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<document id="344E2C2497498436E50B639A864198C2" ID-DOI="10.1093/zoolinnean/zlae112" ID-ISSN="0024-4082" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1732849336941" checkinUser="plazi" docAuthor="Pyrcz, Tomasz, Mahecha-J., Oscar, Boyer, Pierre, Lachowska-Cielik, Dorota, Cerdeña, Jose, Farfán, Jackie, Garlacz, Rafał, Lorenc-Brudecka, Jadwiga, Bálint, Zsolt, Fåhraeus, Christer, Zając-Garlacz, Kamila S. &amp; Espeland, Marianne" docDate="2024" docId="1732B317FFFCFFFFFF76D223B1E9835A" docLanguage="en" docName="zlae112.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlae112" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Nivaliodes puriq Pyrcz, Boyer &amp; Cerdena 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="14" masterDocId="EB0BCB6FFFF4FFF2FFCED575B032854B" masterDocTitle="A snow-dwelling tropical buưerfly? An unprecedented discovery of a new genus of the Pedaliodes clade in an extreme, high-altitude Andean environment (Lepidoptera: Nymphalidae: Satyrinae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="9" updateTime="1735050647935" updateUser="karina">
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<mods:title id="6C34F8527EE7E4862CE8B823197703DB">A snow-dwelling tropical buưerfly? An unprecedented discovery of a new genus of the Pedaliodes clade in an extreme, high-altitude Andean environment (Lepidoptera: Nymphalidae: Satyrinae)</mods:title>
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<mods:namePart id="FD9E64339FB526B5F0CCF1E646EC247B">Pyrcz, Tomasz</mods:namePart>
<mods:affiliation id="8EA17F1ECAAFACEBF5C02CD2264B1C6A">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 380 Kraków, Poland</mods:affiliation>
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<mods:namePart id="64D8A76390A801F9D050AFD789B4A681">Mahecha-J., Oscar</mods:namePart>
<mods:affiliation id="0D346F6B3A0CB22CBF8210E7F8FC7615">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 380 Kraków, Poland &amp; Programa Academico de Biologia, Facultad de Ciencias Matematicas y Naturales, Universidad Distrital Francisco José de Caldas, Carrera 3 No. 26 A - 40, Bogotá, Colombia</mods:affiliation>
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<mods:namePart id="621CC2CD8C7202914061F0954BBB4B9A">Boyer, Pierre</mods:namePart>
<mods:affiliation id="752781AC2FBC3311694AB9A42ABF9EB2">7 Lotissement l’Horizon, Le Puy Sainte Réparade, France</mods:affiliation>
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<mods:affiliation id="FB39981FAB3448AAA6C5397FA3478391">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 380 Kraków, Poland</mods:affiliation>
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<mods:namePart id="FD0B2C9A6D1A2E5AF1C6FB27AA4B8C11">Cerdeña, Jose</mods:namePart>
<mods:affiliation id="5C372595886DE55F5554EC4A56483EFD">Museo de Historia Natural, Universidad Nacional de San Agustín de Arequipa, Arequipa, Peru</mods:affiliation>
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<mods:namePart id="E482B10AF6819967D8FB703DBC2FA067">Farfán, Jackie</mods:namePart>
<mods:affiliation id="65B2C4F5BC40A50D96640DB5E6C96D70">Museo de Historia Natural, Universidad Nacional de San Agustín de Arequipa, Arequipa, Peru</mods:affiliation>
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<mods:namePart id="6A2335D7DA8D7E5B7169F20FADD5506B">Garlacz, Rafał</mods:namePart>
<mods:affiliation id="7309F8EABB9C067B35DF07C0AA2995F5">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 380 Kraków, Poland</mods:affiliation>
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<mods:namePart id="F817810CE9C78A0545FCDA3E687B06A7">Lorenc-Brudecka, Jadwiga</mods:namePart>
<mods:affiliation id="376AC28EC2344FBA73EC12062F2B2E59">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 380 Kraków, Poland</mods:affiliation>
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<mods:namePart id="676D17B2D03F1CDF26FEDFCD0AFB69F7">Bálint, Zsolt</mods:namePart>
<mods:affiliation id="A18CBCFCDB839042C15D6E063F6FC92E">Department of Zoology, Hungarian Natural History Museum, Barros utca 13, 1088 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="015308F045109E91D2DD8B302EB468DB">Fåhraeus, Christer</mods:namePart>
<mods:affiliation id="822CB324673BECE15E7307C74C1908C7">Biodiversity, Department of Biology, Lund University, Lund, Sweden</mods:affiliation>
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<mods:namePart id="279BB14AC11ECF672651832397F8D85C">Zając-Garlacz, Kamila S.</mods:namePart>
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<mods:namePart id="FED36389A95B961448D1B5EB69E17811">Espeland, Marianne</mods:namePart>
<mods:affiliation id="A49F834C5569F85E6E9DDBEFC107040C">Leibniz Institute for the Analysis for Biodiversity Change, Museum Koenig, Adenauerallee 127, 53113 Bonn, Germany</mods:affiliation>
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<mods:title id="407E5BFB1F4031519A7C0DDB5FD2E9E9">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="10FE2CD348D33DCC359D134428CFF72C">2024</mods:date>
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<mods:number id="BF67812BAEE0531035A7118FC8A63B49">2024-10-24</mods:number>
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<mods:number id="BF346914BF460610141871658B03410D">202</mods:number>
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<treatment id="1732B317FFFCFFFFFF76D223B1E9835A" LSID="urn:lsid:plazi:treatment:1732B317FFFCFFFFFF76D223B1E9835A" httpUri="http://treatment.plazi.org/id/1732B317FFFCFFFFFF76D223B1E9835A" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9">
<subSubSection id="D781518AFFFCFFFAFF76D223B2E0823A" box="[184,722,1878,1905]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="9F240201FFFCFFFAFF76D223B2E0823A" blockId="8.[129,778,1878,1972]" box="[184,722,1878,1905]" pageId="8" pageNumber="9">
<emphasis id="ADEFDE13FFFCFFFAFF76D223B2E0823A" bold="true" box="[184,722,1878,1905]" pageId="8" pageNumber="9">
<taxonomicName id="589B7982FFFCFFFAFF76D223B244823B" authority="Pyrcz, Boyer &amp; Cerdena" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[184,630,1878,1905]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="puriq" status="sp. nov.">
<emphasis id="ADEFDE13FFFCFFFAFF76D223B140823B" bold="true" box="[184,370,1878,1905]" italics="true" pageId="8" pageNumber="9">Nivaliodes puriq</emphasis>
Pyrcz, Boyer &amp; Cerdeña
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<taxonomicNameLabel id="B6DC6368FFFCFFFAFDB5D222B2E0823A" box="[635,722,1879,1905]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<taxonomicName id="589B7982FFFCFFFAFF4FD208B11282DE" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[129,288,1917,1941]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="puriq">
<emphasis id="ADEFDE13FFFCFFFAFF4FD208B11282DE" box="[129,288,1917,1941]" italics="true" pageId="8" pageNumber="9">Nivaliodes puriq</emphasis>
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P yrcz,Boyer&amp;Cerdeña:LSIDurn:lsid:zoobank. org:act:
<uuid id="EB3D38D4FFFCFFFAFF04D2E9B28682FF" box="[202,692,1948,1972]" pageId="8" pageNumber="9">21C2C99B-BEE4-4B04-A086-9B056763E1FB</uuid>
<emphasis id="ADEFDE13FFFCFFFAFB81D222B49E823B" box="[1103,1196,1879,1904]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
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<paragraph id="9F240201FFFCFFFBFCF7D208B38E82FD" blockId="8.[825,1474,1917,1973]" lastBlockId="9.[810,1459,1857,1975]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
A medium-sized species, externally most similar to its two congeners, from which it differs by the smaller size, being more compact (less elongated) and even darker, with almost black ground colour. Additionally, the FWD whitish costal patch is slightly smaller, with a straight inner edge, produced in
<taxonomicName id="589B7982FFFDFFFBFD2DD2F5B0D282FC" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFDFFFBFD2DD2F5B0D282FC" italics="true" pageId="9" pageNumber="10">N. negrobueno</emphasis>
</taxonomicName>
, and on the HWV the whitish costal patch is very small and narrow, not extending along the postdiscal line as much as in
<taxonomicName id="589B7982FFFDFFFBFC52D214B4158233" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[924,1063,1888,1912]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFDFFFBFC52D214B4158233" box="[924,1063,1888,1912]" italics="true" pageId="9" pageNumber="10">N. negrobueno</emphasis>
</taxonomicName>
; also, there are no traces of any reddish or rufous markings on the HWV as found in some specimens of
<taxonomicName id="589B7982FFFDFFFBFCE4D2D5B38482FC" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[810,950,1951,1975]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFDFFFBFCE4D2D5B38482FC" box="[810,950,1951,1975]" italics="true" pageId="9" pageNumber="10">N. negrobueno</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="CBE45289FFFDFFFBFFBFD398B427826A" pageId="9" pageNumber="10" startId="9.[113,178,1773,1797]" targetBox="[313,1247,161,1710]" targetPageId="9" targetType="figure">
<paragraph id="9F240201FFFDFFFBFFBFD398B427826A" blockId="9.[113,1443,1773,1825]" pageId="9" pageNumber="10">
<emphasis id="ADEFDE13FFFDFFFBFFBFD398B0FB824E" bold="true" box="[113,201,1773,1797]" pageId="9" pageNumber="10">Figure 5.</emphasis>
Male genitalia in lateral view, with aedeagus extracted in lateral and dorsal view. A,
<taxonomicName id="589B7982FFFDFFFBFC1AD398B482824E" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[980,1200,1773,1797]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFDFFFBFC1AD398B482824E" bold="true" box="[980,1200,1773,1797]" italics="true" pageId="9" pageNumber="10">Nivaliodes negrobueno</emphasis>
</taxonomicName>
(prep. genit CEP_7679). B,
<taxonomicName id="589B7982FFFDFFFBFF47D27CB118826A" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[137,298,1801,1825]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="puriq">
<emphasis id="ADEFDE13FFFDFFFBFF47D27CB118826A" bold="true" box="[137,298,1801,1825]" italics="true" pageId="9" pageNumber="10">Nivaliodes puriq</emphasis>
</taxonomicName>
(prep. genit CEP_10558). C,
<taxonomicName id="589B7982FFFDFFFBFD86D27CB323826A" authorityName="Pyrcz &amp; Mahecha-J. &amp; Boyer &amp; Lachowska-Cielik &amp; Cerdeña &amp; Farfán &amp; Garlacz &amp; Lorenc-Brudecka &amp; Bálint &amp; Fåhraeus &amp; Zając-Garlacz &amp; Espeland" authorityYear="2024" box="[584,785,1801,1825]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="viracocha">
<emphasis id="ADEFDE13FFFDFFFBFD86D27CB323826A" bold="true" box="[584,785,1801,1825]" italics="true" pageId="9" pageNumber="10">Nivaliodes viracocha</emphasis>
</taxonomicName>
(prep. genit. CEP_10615).
</paragraph>
</caption>
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<paragraph id="9F240201FFFEFFF8FF4FD186B1D18008" blockId="10.[129,764,1267,1347]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FF4FD186B0EA8040" bold="true" box="[129,216,1267,1291]" pageId="10" pageNumber="11">Figure 6.</emphasis>
Female genitalia in lateral view. A,
<taxonomicName id="589B7982FFFEFFF8FDEED186B2CE8040" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[544,764,1267,1291]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFEFFF8FDEED186B2CE8040" bold="true" box="[544,764,1267,1291]" italics="true" pageId="10" pageNumber="11">Nivaliodes negrobueno</emphasis>
</taxonomicName>
(CEP_7680). B.
<taxonomicName id="589B7982FFFEFFF8FEEFD07AB1F0806C" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[289,450,1295,1319]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="puriq">
<emphasis id="ADEFDE13FFFEFFF8FEEFD07AB1F0806C" bold="true" box="[289,450,1295,1319]" italics="true" pageId="10" pageNumber="11">Nivaliodes puriq</emphasis>
</taxonomicName>
(CEP_10615). C.
<taxonomicName id="589B7982FFFEFFF8FDB8D07AB0ED8008" authorityName="Pyrcz &amp; Mahecha-J. &amp; Boyer &amp; Lachowska-Cielik &amp; Cerdeña &amp; Farfán &amp; Garlacz &amp; Lorenc-Brudecka &amp; Bálint &amp; Fåhraeus &amp; Zając-Garlacz &amp; Espeland" authorityYear="2024" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="viracocha">
<emphasis id="ADEFDE13FFFEFFF8FDB8D07AB0ED8008" bold="true" italics="true" pageId="10" pageNumber="11">Nivaliodes viracocha</emphasis>
</taxonomicName>
(prep. genit. CEP_10616).
</paragraph>
</caption>
<paragraph id="9F240201FFFEFFF8FEE1D0F4B26980D1" blockId="10.[129,780,1408,2003]" box="[303,603,1408,1435]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FEE1D0F4B1E180D1" box="[303,467,1408,1434]" italics="true" pageId="10" pageNumber="11">Description: male</emphasis>
(
<figureCitation id="07A01E84FFFEFFF8FE2BD0F5B27D80D0" box="[485,591,1408,1435]" captionStart="Figure 4" captionStartId="8.[129,194,1772,1796]" captionTargetBox="[267,1339,157,1705]" captionTargetId="figure-48@8.[225,1377,144,1744]" captionTargetPageId="8" captionText="Figure 4. Adults. A–D, Nivaliodes puriq. A, male holotype, dorsal, Trancapampa. B, idem, ventral (Trancapampa).C, female paratype, dorsal, Trancapampa. D, idem, ventral. E–H, Nivaliodes viracocha. E, male holotype, dorsal, Viracochan.F, idem, ventral. G, female paratype, dorsal, Viracochan. H, idem, ventral." pageId="10" pageNumber="11">Fig. 4A, B</figureCitation>
)
</paragraph>
<paragraph id="9F240201FFFEFFF8FF4FD0D2B49F84AA" blockId="10.[129,780,1408,2003]" lastBlockId="10.[825,1475,144,481]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FF4FD0D2B08F80F4" box="[129,189,1447,1471]" italics="true" pageId="10" pageNumber="11">Head:</emphasis>
Eyes blackish brown, covered with dense and medium long hair, with a milky white collar; palpi two times the length of head, covered with long, dorsally short blackish brown hairy scales, with a lateral row of milky white scales; frons with a tuss of blackish brown hair; antennae reaching two-fisshs the length of costa, slender, dark brown, mostly naked, with a club formed gradually, composed of 11 flagellomeres, flattened dorsoventrally; thorax black, dorsally covered with short and sparse, golden brown hairy scales; mesothoracic legs covered with brown and blackish brown scales; FW length (
<quantity id="5863AFE4FFFEFFF8FE5FD3B4B2358393" box="[401,519,1729,1753]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="10" pageNumber="11" unit="mm" value="22.0" valueMax="23.0" valueMin="21.0">21–23 mm</quantity>
, mean:
<quantity id="5863AFE4FFFEFFF8FD98D3B5B2868393" box="[598,692,1728,1752]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="10" pageNumber="11" unit="mm" value="22.5">22.5 mm</quantity>
,
<emphasis id="ADEFDE13FFFEFFF8FD71D3B4B2E08393" box="[703,722,1729,1752]" italics="true" pageId="10" pageNumber="11">N</emphasis>
= 3), with a subacute apex and slightly convex outer margin, and with long intermittent milky white and black fringes; upperside brown and blackish brown, a shade darker in median half, lustrous, with a small white costal patch one-third of the way from distal end of discal cell and apex, spreading over three spaces, costa, R4 + 5– M1, and M1–M2, widening in the middle, with a straight inner margin, no scent patch; underside dull, blackish brown, with the white costal patch as on the upperside, sprinkled with white scales in the subapical and apical area, and a row of four to five minute white submarginal dots from costa to cell M3–CuA1. HW rounded, with an undulating outer margin and with intermittently grey and milky white fringes; HWD uniform blackish brown, lustrous, hairy in median half; HWV ground colour greyish brown, with a heavy whitish and black ripple pattern covering the entire wing surface without producing any conspicuous pattern except for a very lightly marked, irregular black postdiscal line; the unique well-marked pattern is a short midcostal whitish streak. Abdomen is covered dorsally and laterally with thick black scales and ventrally with dark brown scales.
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D776B53F87BD" blockId="10.[825,1474,514,758]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FCF7D776B3A68750" box="[825,916,515,539]" italics="true" pageId="10" pageNumber="11">Genitalia</emphasis>
(
<figureCitation id="07A01E84FFFEFFF8FC6BD776B3DE8751" box="[933,1004,514,539]" captionStart="Figure 5" captionStartId="9.[113,178,1773,1797]" captionTargetBox="[313,1247,161,1710]" captionTargetId="figure-86@9.[296,1275,145,1745]" captionTargetPageId="9" captionText="Figure 5. Male genitalia in lateral view, with aedeagus extracted in lateral and dorsal view.A, Nivaliodes negrobueno (prep. genit CEP_7679). B, Nivaliodes puriq (prep.genit CEP_10558). C, Nivaliodes viracocha (prep. genit.CEP_10615)." pageId="10" pageNumber="11">Fig. 5B</figureCitation>
): Tegumen with a flat dorsum; uncus as long as the dorsum of tegumen, aligned with tegumen, nearly straight, with a blunt tip; subunci approximately half the length of uncus, stout; pedunculus short, with a sharp extremity curved downwards; saccus wide and deep; valvae as long as tegumen + valva, slender, with a moderately humped, slightly irregular dorsum and a subacute apex; aedeagus short, wide, and tubular, about the same length throughout, with an acute tip.
</paragraph>
<paragraph id="9F240201FFFEFFF8FC10D663B52F867B" blockId="10.[825,1474,789,883]" box="[990,1309,789,816]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FC10D663B4A18664" box="[990,1171,789,815]" italics="true" pageId="10" pageNumber="11">Description: female</emphasis>
(
<figureCitation id="07A01E84FFFEFFF8FB6CD660B523867B" box="[1186,1297,789,816]" captionStart="Figure 4" captionStartId="8.[129,194,1772,1796]" captionTargetBox="[267,1339,157,1705]" captionTargetId="figure-48@8.[225,1377,144,1744]" captionTargetPageId="8" captionText="Figure 4. Adults. A–D, Nivaliodes puriq. A, male holotype, dorsal, Trancapampa. B, idem, ventral (Trancapampa).C, female paratype, dorsal, Trancapampa. D, idem, ventral. E–H, Nivaliodes viracocha. E, male holotype, dorsal, Viracochan.F, idem, ventral. G, female paratype, dorsal, Viracochan. H, idem, ventral." pageId="10" pageNumber="11">Fig. 4C, D</figureCitation>
)
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D649B4B98638" blockId="10.[825,1474,789,883]" pageId="10" pageNumber="11">
FW length
<quantity id="5863AFE4FFFEFFF8FC63D649B3C48618" box="[941,1014,828,852]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="25.0">25 mm</quantity>
(
<emphasis id="ADEFDE13FFFEFFF8FBC8D648B42B861F" box="[1030,1049,829,852]" italics="true" pageId="10" pageNumber="11">N</emphasis>
= 2), lighter coloured on both the upperand underside, otherwise similar.
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D6E0B4988102" blockId="10.[825,1474,917,1097]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FCF7D6E0B3A686E6" box="[825,916,917,941]" italics="true" pageId="10" pageNumber="11">Genitalia</emphasis>
(
<figureCitation id="07A01E84FFFEFFF8FC69D6E0B3C386E6" box="[935,1009,917,941]" captionStart="Figure 6" captionStartId="10.[129,194,1267,1291]" captionTargetBox="[184,755,146,1228]" captionTargetId="figure-683@10.[149,757,144,1238]" captionTargetPageId="10" captionText="Figure 6. Female genitalia in lateral view.A, Nivaliodes negrobueno (CEP_7680). B. Nivaliodes puriq (CEP_10615). C. Nivaliodes viracocha (prep. genit.CEP_10616)." pageId="10" pageNumber="11">Fig. 6B</figureCitation>
): Similar to
<taxonomicName id="589B7982FFFEFFF8FB4CD6E0B52386E6" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[1154,1297,917,941]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFEFFF8FB4CD6E0B52386E6" box="[1154,1297,917,941]" italics="true" pageId="10" pageNumber="11">N. negrobueno</emphasis>
</taxonomicName>
, with prominent papillae anales, covered with long but rather sparse hair; strongly sclerotized postvaginal lamella with a smooth surface, wide, enclosing a spacious antrum; short, mildly sclerotized, wide ductus burse, opening into a large, pear-like corpus bursae, with a smooth surface, no apparent signa.
</paragraph>
<paragraph id="9F240201FFFEFFF8FB9FD11FB49881C8" blockId="10.[825,1474,1130,1254]" box="[1105,1194,1130,1155]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FB9FD11FB49881C8" box="[1105,1194,1130,1155]" italics="true" pageId="10" pageNumber="11">Variation</emphasis>
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D1FAB4D781AD" blockId="10.[825,1474,1130,1254]" pageId="10" pageNumber="11">Individual variation is negligible and affects mostly the size of the FWD postdiscal white patch; individuals from Puente Carrizales are characterized by a slightly larger patch.</paragraph>
<paragraph id="9F240201FFFEFFF8FBF3D077B48D8050" blockId="10.[825,1474,1281,1437]" box="[1085,1215,1281,1307]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FBF3D077B48D8050" box="[1085,1215,1281,1307]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="4020BCA3FFFEFFF8FBF3D077B4598050" box="[1085,1131,1282,1307]" pageId="10" pageNumber="11">Type</typeStatus>
material
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="D781518AFFFEFFF8FCF7D052B5F1827B" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="9F240201FFFEFFF8FCF7D052B39A80D6" blockId="10.[825,1474,1281,1437]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FCF7D052B3A08074" box="[825,914,1319,1343]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="4020BCA3FFFEFFF8FCF7D052B3A08074" box="[825,914,1319,1343]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
<materialsCitation id="2FF3085CFFFEFFF8FC53D05FB39680D6" collectingDate="2021-09-29" collectionCode="CEPUJ" collectorName="P. Boyer" country="Peru" elevation="3220" latitude="-12.046667" location="Est de Huari" longLatPrecision="21" longitude="-74.91805" municipality="Est de Huari" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Huancavelica" typeStatus="holotype">
<specimenCount id="899DC988FFFEFFF8FC53D05FB3858075" box="[925,951,1322,1342]" count="1" pageId="10" pageNumber="11" type="male">♂</specimenCount>
:
<collectingCountry id="E78C4291FFFEFFF8FC07D052B3C98074" box="[969,1019,1319,1343]" name="Peru" pageId="10" pageNumber="11">Perú</collectingCountry>
,
<collectingRegion id="5D5FCCE3FFFEFFF8FBC4D052B4A78074" box="[1034,1173,1319,1343]" country="Peru" name="Huancavelica" pageId="10" pageNumber="11">Huancavelica</collectingRegion>
,
<collectingMunicipality id="7F40987BFFFEFFF8FB6DD05DB5028074" box="[1187,1328,1319,1343]" pageId="10" pageNumber="11">Est de Huari</collectingMunicipality>
,
<locationDeviation id="8E961B23FFFEFFF8FAF0D052B3F48015" pageId="10" pageNumber="11">
<quantity id="5863AFE4FFFEFFF8FAF0D052B5438074" box="[1342,1393,1319,1343]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="km" value="4.0">4 km</quantity>
east of Trancapampa
</locationDeviation>
, HV109, PK26,
<geoCoordinate id="FAAF64C6FFFEFFF8FBB1D032B4C38015" box="[1151,1265,1349,1375]" degrees="12" direction="south" minutes="02" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="48" value="-12.046667">
12°02
<emphasis id="ADEFDE13FFFEFFF8FB75D030B4F38015" box="[1211,1217,1349,1374]" italics="true" pageId="10" pageNumber="11">ʹ</emphasis>
48″S
</geoCoordinate>
,
<geoCoordinate id="FAAF64C6FFFEFFF8FACFD033B54D8015" box="[1281,1407,1349,1374]" degrees="74" direction="west" minutes="55" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="05" value="-74.91805">
74°55
<emphasis id="ADEFDE13FFFEFFF8FAF3D030B5718015" box="[1341,1347,1349,1374]" italics="true" pageId="10" pageNumber="11">ʹ</emphasis>
05″W
</geoCoordinate>
,
<quantity id="5863AFE4FFFEFFF8FA40D032B37D8036" metricMagnitude="3" metricUnit="m" metricValue="3.22" pageId="10" pageNumber="11" unit="m" value="3220.0">
<elevation id="14B6E532FFFEFFF8FA40D032B37D8036" metricMagnitude="3" metricUnit="m" metricValue="3.22" pageId="10" pageNumber="11" unit="m" value="3220.0">3220 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FC92D013B4198035" box="[860,1067,1382,1406]" pageId="10" pageNumber="11" value="2021-09-29">
<collectingDate id="FB61DD29FFFEFFF8FC92D013B4198035" box="[860,1067,1382,1406]" pageId="10" pageNumber="11" value="2021-09-29">29 September 2021</collectingDate>
</date>
,
<collectorName id="326E67D7FFFEFFF8FBF9D013B4BF8035" box="[1079,1165,1382,1406]" pageId="10" pageNumber="11">P. Boyer</collectorName>
leg.,
<collectionCode id="F98A9AC4FFFEFFF8FB08D013B5218035" box="[1222,1299,1382,1406]" pageId="10" pageNumber="11">CEPUJ</collectionCode>
, to be deposited in MUSM
</materialsCitation>
.
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D0B0B53283D2" blockId="10.[825,1474,1477,1689]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FCF7D0B0B3AA8097" box="[825,920,1477,1500]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="4020BCA3FFFEFFF8FCF7D0B0B3AA8097" box="[825,920,1477,1500]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="899DC988FFFEFFF8FC66D0B0B3D48097" box="[936,998,1477,1501]" count="10" pageId="10" pageNumber="11" type="male">10 ♂</specimenCount>
and
<specimenCount id="899DC988FFFEFFF8FBD5D0B0B4518097" box="[1051,1123,1477,1500]" count="2" pageId="10" pageNumber="11" type="female">two ♀</specimenCount>
):
<materialsCitation id="2FF3085CFFFEFFF8FBB3D0B0B38080B0" collectingDate="2021-09-29" collectionCode="PBF" collectorName="de Huari &amp; Trancapampa, HV &amp; P. Boyer" country="Peru" elevation="3220" latitude="-12.046667" location="Trancapampa" longLatPrecision="21" longitude="-74.91805" municipality="Est de Huari" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" stateProvince="Huancavelica" typeStatus="paratype">
<specimenCount id="899DC988FFFEFFF8FBB3D0B0B4FF8097" box="[1149,1229,1477,1500]" count="2" pageId="10" pageNumber="11" type="male">Two ♂</specimenCount>
, same data as the holotype [
<collectionCode id="F98A9AC4FFFEFFF8FCBAD091B39980B0" box="[884,939,1508,1531]" pageId="10" pageNumber="11">PBF</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="2FF3085CFFFEFFF8FC72D091B58C8350" collectingDate="1929-09-30" collectingDateMax="1929-10-01" collectingDateMin="1929-09-30" collectionCode="CEPUJ" collectorName="T. Pyrcz" country="Peru" elevation="3100" location="Trancapampa" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3" stateProvince="Huancavelica" typeStatus="paratype">
<specimenCount id="899DC988FFFEFFF8FC72D091B42080B0" box="[956,1042,1508,1532]" count="3" pageId="10" pageNumber="11" type="male">three ♂</specimenCount>
:
<collectingRegion id="5D5FCCE3FFFEFFF8FBEED091B49980B7" box="[1056,1195,1508,1532]" country="Peru" name="Huancavelica" pageId="10" pageNumber="11">Huancavelica</collectingRegion>
,
<location id="9A4454DAFFFEFFF8FB7BD091B57080B0" LSID="urn:lsid:plazi:treatment:1732B317FFFCFFFFFF76D223B1E9835A:9A4454DAFFFEFFF8FB7BD091B57080B0" box="[1205,1346,1508,1531]" country="Peru" name="Trancapampa" pageId="10" pageNumber="11" stateProvince="Huancavelica">Trancapampa</location>
,
<quantity id="5863AFE4FFFEFFF8FA82D091B37D8350" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="10" pageNumber="11" unit="m" value="3100.0" valueMax="3200.0" valueMin="3000.0">
<elevation id="14B6E532FFFEFFF8FA82D091B37D8350" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="10" pageNumber="11" unit="m" value="3100.0" valueMax="3200.0" valueMin="3000.0">3000–3200 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FC93D376B4A28350" box="[861,1168,1539,1563]" pageId="10" pageNumber="11" value="1929-09-30" valueMax="1929-10-01" valueMin="1929-09-30">
<collectingDate id="FB61DD29FFFEFFF8FC93D376B4A28350" box="[861,1168,1539,1563]" pageId="10" pageNumber="11" value="1929-09-30" valueMax="1929-10-01" valueMin="1929-09-30">29–30 September–1 October</collectingDate>
</date>
2021,
<collectorName id="326E67D7FFFEFFF8FB12D371B5018357" box="[1244,1331,1540,1564]" pageId="10" pageNumber="11">T. Pyrcz</collectorName>
leg.,
<collectionCode id="F98A9AC4FFFEFFF8FAA0D376B58C8350" box="[1390,1470,1539,1563]" pageId="10" pageNumber="11">CEPUJ</collectionCode>
</materialsCitation>
;
<materialsCitation id="2FF3085CFFFEFFF8FCF7D356B5078311" collectingDate="1929-09-30" collectingDateMax="1929-10-01" collectingDateMin="1929-09-30" collectionCode="CEPUJ" collectorName="T. Pyrcz" country="Peru" elevation="4200" location="Trancapampa" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" stateProvince="Huancavelica" typeStatus="paratype">
<specimenCount id="899DC988FFFEFFF8FCF7D356B3B48371" box="[825,902,1571,1594]" count="1" pageId="10" pageNumber="11" type="female">one ♀</specimenCount>
:
<collectingRegion id="5D5FCCE3FFFEFFF8FC54D356B4178370" box="[922,1061,1571,1595]" country="Peru" name="Huancavelica" pageId="10" pageNumber="11">Huancavelica</collectingRegion>
, above
<location id="9A4454DAFFFEFFF8FBB3D356B5388371" LSID="urn:lsid:plazi:treatment:1732B317FFFCFFFFFF76D223B1E9835A:9A4454DAFFFEFFF8FBB3D356B5388371" box="[1149,1290,1571,1594]" country="Peru" name="Trancapampa" pageId="10" pageNumber="11" stateProvince="Huancavelica">Trancapampa</location>
,
<quantity id="5863AFE4FFFEFFF8FAD4D356B5428371" box="[1306,1392,1571,1595]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="11" unit="m" value="4200.0">
<elevation id="14B6E532FFFEFFF8FAD4D356B5428371" box="[1306,1392,1571,1595]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="11" unit="m" value="4200.0">4200 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FA4ED356B42F8311" pageId="10" pageNumber="11" value="1929-09-30" valueMax="1929-10-01" valueMin="1929-09-30">
<collectingDate id="FB61DD29FFFEFFF8FA4ED356B42F8311" pageId="10" pageNumber="11" value="1929-09-30" valueMax="1929-10-01" valueMin="1929-09-30">29–30 September–1 October</collectingDate>
</date>
2021,
<collectorName id="326E67D7FFFEFFF8FBAED337B4818311" box="[1120,1203,1602,1626]" pageId="10" pageNumber="11">T. Pyrcz</collectorName>
leg.,
<collectionCode id="F98A9AC4FFFEFFF8FB2BD337B5078311" box="[1253,1333,1602,1626]" pageId="10" pageNumber="11">CEPUJ</collectionCode>
</materialsCitation>
;
<materialsCitation id="2FF3085CFFFEFFF8FAF3D337B4C983D2" collectingDate="2021-09-30" collectionCode="MUSM" collectorName="J. Cerdena" country="Peru" elevation="3200" location="Trancapampa" municipality="Trancapampa" pageId="10" pageNumber="11" specimenCount="3" specimenCount-male="3" stateProvince="Huancavelica" typeStatus="paratype">
<specimenCount id="899DC988FFFEFFF8FAF3D337B5A28312" box="[1341,1424,1602,1626]" count="3" pageId="10" pageNumber="11" type="male">three ♂</specimenCount>
: HV [uancavelica],
<collectingMunicipality id="7F40987BFFFEFFF8FC00D317B4698332" box="[974,1115,1634,1657]" pageId="10" pageNumber="11">Trancapampa</collectingMunicipality>
,
<locationDeviation id="8E961B23FFFEFFF8FBA8D314B5508331" box="[1126,1378,1633,1658]" pageId="10" pageNumber="11">
<quantity id="5863AFE4FFFEFFF8FBA8D314B49B8332" box="[1126,1193,1633,1657]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="30.0">30 km</quantity>
east of Huancayo
</locationDeviation>
,
<quantity id="5863AFE4FFFEFFF8FAA3D314B58C8332" box="[1389,1470,1633,1657]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="11" unit="m" value="3200.0">
<elevation id="14B6E532FFFEFFF8FAA3D314B58C8332" box="[1389,1470,1633,1657]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="11" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FCF5D3F4B43783D2" box="[827,1029,1665,1689]" pageId="10" pageNumber="11" value="2021-09-30">
<collectingDate id="FB61DD29FFFEFFF8FCF5D3F4B43783D2" box="[827,1029,1665,1689]" pageId="10" pageNumber="11" value="2021-09-30">30 September 2021</collectingDate>
</date>
,
<collectorName id="326E67D7FFFEFFF8FBC3D3F4B44583D2" box="[1037,1143,1665,1689]" pageId="10" pageNumber="11">J. Cerdeña</collectorName>
leg.,
<collectionCode id="F98A9AC4FFFEFFF8FB64D3F4B4C983D2" box="[1194,1275,1665,1689]" collectionName="Peru, Lima, Universidad Nacional Mayor de San Marcos, Museo de Historia Natural" pageId="10" pageNumber="11">MUSM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D3CFB5F1827B" blockId="10.[825,1475,1722,1840]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FCF7D3CFB3C58399" box="[825,1015,1722,1746]" italics="true" pageId="10" pageNumber="11">Additional material:</emphasis>
<materialsCitation id="2FF3085CFFFEFFF8FC33D3CFB506825B" collectingDate="2009-10-15" collectionCode="PBF" collectorName="P. Boyer" country="Peru" county="Puente Carrizales" elevation="3375" latitude="-11.4867" location="Puente Carrizales" longLatPrecision="7" longitude="-74.8859" municipality="Route Satipo-Mariposa" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Junin">
<specimenCount id="899DC988FFFEFFF8FC33D3CFB47B839A" box="[1021,1097,1722,1746]" count="1" pageId="10" pageNumber="11" type="male">One ♂</specimenCount>
:
<collectingCountry id="E78C4291FFFEFFF8FB9BD3CFB4B78399" box="[1109,1157,1722,1746]" name="Peru" pageId="10" pageNumber="11">Perú</collectingCountry>
,
<collectingRegion id="5D5FCCE3FFFEFFF8FB40D3CFB4F68399" box="[1166,1220,1722,1746]" country="Peru" name="Junin" pageId="10" pageNumber="11">Junín</collectingRegion>
,
<collectingCounty id="76457A8DFFFEFFF8FB00D3CFB54E8399" box="[1230,1404,1722,1746]" pageId="10" pageNumber="11">Puente Carrizales</collectingCounty>
,
<locationDeviation id="8E961B23FFFEFFF8FA4BD3CFB4ED83BA" pageId="10" pageNumber="11">Route Satipo-Mariposa vers Concepción km 72</locationDeviation>
,
<geoCoordinate id="FAAF64C6FFFEFFF8FB20D3ACB57D83BA" box="[1262,1359,1753,1777]" degrees="11.4867" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="5" value="-11.4867">−11.4867</geoCoordinate>
,
<geoCoordinate id="FAAF64C6FFFEFFF8FA90D3ACB58C83BA" box="[1374,1470,1753,1777]" degrees="74.8859" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="5" value="-74.8859">−74.8859</geoCoordinate>
,
<quantity id="5863AFE4FFFEFFF8FCF7D38DB3F6825B" box="[825,964,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="3.375" metricValueMax="3.45" metricValueMin="3.3" pageId="10" pageNumber="11" unit="m" value="3375.0" valueMax="3450.0" valueMin="3300.0">
<elevation id="14B6E532FFFEFFF8FCF7D38DB3F6825B" box="[825,964,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="3.375" metricValueMax="3.45" metricValueMin="3.3" pageId="10" pageNumber="11" unit="m" value="3375.0" valueMax="3450.0" valueMin="3300.0">3300–3450 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FC04D38DB442825A" box="[970,1136,1784,1809]" pageId="10" pageNumber="11" value="2009-10-15">
<collectingDate id="FB61DD29FFFEFFF8FC04D38DB442825A" box="[970,1136,1784,1809]" pageId="10" pageNumber="11" value="2009-10-15">15 October2009</collectingDate>
</date>
,
<collectorName id="326E67D7FFFEFFF8FBB9D38CB4F4825A" box="[1143,1222,1785,1809]" pageId="10" pageNumber="11">P.Boyer</collectorName>
leg.,[
<collectionCode id="F98A9AC4FFFEFFF8FB37D38CB51C825B" box="[1273,1326,1785,1808]" pageId="10" pageNumber="11">PBF</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="2FF3085CFFFEFFF8FAF5D38CB58D827B" collectingDate="2019-06-05" collectionCode="CEPUJ" collectorName="T. Pyrcz" country="Peru" elevation="3425" location="Puente Carrizales" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Junin">
<specimenCount id="899DC988FFFEFFF8FAF5D38CB54C825B" box="[1339,1406,1785,1808]" count="1" pageId="10" pageNumber="11" type="male">one ♂</specimenCount>
:
<collectingRegion id="5D5FCCE3FFFEFFF8FA46D38CB58D825A" box="[1416,1471,1785,1809]" country="Peru" name="Junin" pageId="10" pageNumber="11">Junín</collectingRegion>
,
<location id="9A4454DAFFFEFFF8FCF7D26DB3D7827B" LSID="urn:lsid:plazi:treatment:1732B317FFFCFFFFFF76D223B1E9835A:9A4454DAFFFEFFF8FCF7D26DB3D7827B" box="[825,997,1816,1840]" country="Peru" name="Puente Carrizales" pageId="10" pageNumber="11" stateProvince="Junin">Puente Carrizales</location>
,
<quantity id="5863AFE4FFFEFFF8FC22D26DB4448264" box="[1004,1142,1816,1840]" metricMagnitude="3" metricUnit="m" metricValue="3.425" metricValueMax="3.45" metricValueMin="3.4" pageId="10" pageNumber="11" unit="m" value="3425.0" valueMax="3450.0" valueMin="3400.0">
<elevation id="14B6E532FFFEFFF8FC22D26DB4448264" box="[1004,1142,1816,1840]" metricMagnitude="3" metricUnit="m" metricValue="3.425" metricValueMax="3.45" metricValueMin="3.4" pageId="10" pageNumber="11" unit="m" value="3425.0" valueMax="3450.0" valueMin="3400.0">3400–3450 m</elevation>
</quantity>
,
<date id="EB2524C1FFFEFFF8FBB3D26DB4C3827B" box="[1149,1265,1816,1840]" pageId="10" pageNumber="11" value="2019-06-05">
<collectingDate id="FB61DD29FFFEFFF8FBB3D26DB4C3827B" box="[1149,1265,1816,1840]" pageId="10" pageNumber="11" value="2019-06-05">5 June 2019</collectingDate>
</date>
,
<collectorName id="326E67D7FFFEFFF8FB36D26DB575827B" box="[1272,1351,1816,1840]" pageId="10" pageNumber="11">T.Pyrcz</collectorName>
leg.,
<collectionCode id="F98A9AC4FFFEFFF8FABAD26DB58D827B" box="[1396,1471,1816,1840]" pageId="10" pageNumber="11">CEPUJ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D781518AFFFEFFF8FB85D220B3FF8298" pageId="10" pageNumber="11" type="etymology">
<paragraph id="9F240201FFFEFFF8FB85D220B4838224" blockId="10.[1099,1201,1877,1903]" box="[1099,1201,1877,1903]" pageId="10" pageNumber="11">
<emphasis id="ADEFDE13FFFEFFF8FB85D220B4838224" box="[1099,1201,1877,1903]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
</paragraph>
<paragraph id="9F240201FFFEFFF8FCF7D209B3FF8298" blockId="10.[825,1474,1916,2003]" pageId="10" pageNumber="11">
The specific epithet of this taxon,
<taxonomicName id="589B7982FFFEFFF8FB7AD209B4DC82DF" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[1204,1262,1916,1940]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="puriq">
‘
<emphasis id="ADEFDE13FFFEFFF8FB77D208B4DC82DF" box="[1209,1262,1917,1940]" italics="true" pageId="10" pageNumber="11">puriq</emphasis>
</taxonomicName>
’, means traveller in Quechua and is an allusion to the apparent dispersalist tendency of this species.
</paragraph>
</subSubSection>
<caption id="CBE45289FFFFFFF9FFBFD399B141823F" pageId="11" pageNumber="12" startId="11.[113,178,1772,1796]" targetBox="[260,1313,149,1737]" targetPageId="11" targetType="figure">
<paragraph id="9F240201FFFFFFF9FFBFD399B141823F" blockId="11.[113,1422,1772,1909]" pageId="11" pageNumber="12">
<emphasis id="ADEFDE13FFFFFFF9FFBFD399B0FB824E" bold="true" box="[113,201,1772,1797]" pageId="11" pageNumber="12">Figure 7.</emphasis>
Scanning electron microscopy images of
<taxonomicName id="589B7982FFFFFFF9FD83D399B31B824F" authorityName="Pyrcz, Fahraeus &amp; Boyer" authorityYear="2024" box="[589,809,1772,1796]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="negrobueno">
<emphasis id="ADEFDE13FFFFFFF9FD83D399B31B824F" bold="true" box="[589,809,1772,1796]" italics="true" pageId="11" pageNumber="12">Nivaliodes negrobueno</emphasis>
</taxonomicName>
scales. A, scale-covered wing membrane, with visible discal veins also covered by ground scales. B, elongated cover scales with hairy scales. C, two morphotypes of scales: elongated cover scales and wide, shovel-shaped ground scales. D, elongated cover scales and hairy scales. E, elongated cover scale with pointed apical termini. F, shovellike cover scale with undulating terminus. G, detail of cover scale micro- and structures, median area. H, detail of cover scale micro- and nanostructures, apical area.
</paragraph>
</caption>
<caption id="CBE45289FFF8FFFEFF4FD39EB1D18238" pageId="12" pageNumber="13" startId="12.[129,194,1771,1795]" targetBox="[275,1328,148,1736]" targetPageId="12" targetType="figure">
<paragraph id="9F240201FFF8FFFEFF4FD39EB1D18238" blockId="12.[129,1468,1771,1907]" pageId="12" pageNumber="13">
<emphasis id="ADEFDE13FFF8FFFEFF4FD39EB0EA8248" bold="true" box="[129,216,1771,1795]" pageId="12" pageNumber="13">Figure 8.</emphasis>
Scanning electron microscopy images of
<taxonomicName id="589B7982FFF8FFFEFD92D39EB2CF8248" authorityName="Pyrcz, Boyer &amp; Cerdena" authorityYear="2024" box="[604,765,1771,1795]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puriq">
<emphasis id="ADEFDE13FFF8FFFEFD92D39EB2CF8248" bold="true" box="[604,765,1771,1795]" italics="true" pageId="12" pageNumber="13">Nivaliodes puriq</emphasis>
</taxonomicName>
scales. A, scale-covered wing membrane with visible veins free of scales. B, transformed cover scales with sparse hair-like androconial scales. C, transformed cover scales with elongated shape on top, shovel-like ground scales below. D, transformed cover scales and sparse androconial scales. E, transformed cover scale with pointed termini. F, fragments of a transformed cover scale and an androconial scale. G, detail of terminal part of a hair-like androconial scale. H, micro- and nanostructures in the transformed cover scale, median area.
</paragraph>
</caption>
<caption id="CBE45289FFF9FFFFFFBDD058B532800E" box="[115,1280,1325,1350]" pageId="13" pageNumber="14" startId="13.[115,180,1325,1349]" targetBox="[149,1422,147,1293]" targetPageId="13" targetType="figure">
<paragraph id="9F240201FFF9FFFFFFBDD058B532800E" blockId="13.[115,1280,1325,1350]" box="[115,1280,1325,1350]" pageId="13" pageNumber="14">
<emphasis id="ADEFDE13FFF9FFFFFFBDD058B0F8800E" bold="true" box="[115,202,1325,1349]" pageId="13" pageNumber="14">Figure 9.</emphasis>
Temperature range of three species of
<taxonomicName id="589B7982FFF9FFFFFDFBD058B2A9800E" authorityName="Pyrcz &amp; Mahecha-J. &amp; Boyer &amp; Lachowska-Cielik &amp; Cerdeña &amp; Farfán &amp; Garlacz &amp; Lorenc-Brudecka &amp; Bálint &amp; Fåhraeus &amp; Zając-Garlacz &amp; Espeland" authorityYear="2024" box="[565,667,1325,1349]" class="Insecta" family="Boreidae" genus="Nivaliodes" kingdom="Animalia" order="Mecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="ADEFDE13FFF9FFFFFDFBD058B2A9800E" bold="true" box="[565,667,1325,1349]" italics="true" pageId="13" pageNumber="14">Nivaliodes</emphasis>
</taxonomicName>
and
<taxonomicName id="589B7982FFF9FFFFFD06D058B3AA800E" baseAuthorityName="Nieme" baseAuthorityYear="1905" box="[712,920,1325,1349]" class="Insecta" family="Nymphalidae" genus="Pherepedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naevia">
<emphasis id="ADEFDE13FFF9FFFFFD06D058B3AA800E" box="[712,920,1325,1349]" italics="true" pageId="13" pageNumber="14">Pherepedaliodes naevia</emphasis>
</taxonomicName>
(box plot with median and quartiles).
</paragraph>
</caption>
<subSubSection id="D781518AFFF9FFFFFEB2D000B1E9835A" pageId="13" pageNumber="14" type="distribution">
<paragraph id="9F240201FFF9FFFFFEB2D000B1DD80C4" blockId="13.[113,763,1397,1553]" box="[380,495,1397,1423]" pageId="13" pageNumber="14">
<emphasis id="ADEFDE13FFF9FFFFFEB2D000B1DD80C4" box="[380,495,1397,1423]" italics="true" pageId="13" pageNumber="14">Distribution</emphasis>
</paragraph>
<paragraph id="9F240201FFF9FFFFFFBFD0E9B1E9835A" blockId="13.[113,763,1397,1553]" pageId="13" pageNumber="14">
So far, this species is known from three localities, all of which are within a
<quantity id="5863AFE4FFF9FFFFFF31D0CEB1778098" box="[255,325,1467,1491]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="km" value="60.0">60 km</quantity>
radius in the departments of
<collectingRegion id="5D5FCCE3FFF9FFFFFD5CD0CEB2FB8098" box="[658,713,1467,1491]" country="Peru" name="Junin" pageId="13" pageNumber="14">Junín</collectingRegion>
and
<collectingRegion id="5D5FCCE3FFF9FFFFFFBFD0AFB0C880B9" box="[113,250,1498,1522]" country="Peru" name="Huancavelica" pageId="13" pageNumber="14">Huancavelica</collectingRegion>
(
<figureCitation id="07A01E84FFF9FFFFFEC5D0AFB16380B9" box="[267,337,1498,1522]" captionStart="Figure 10" captionStartId="14.[129,194,1694,1718]" captionTargetBox="[129,1473,143,1666]" captionTargetId="figure-145@14.[129,1473,143,1666]" captionTargetPageId="14" captionText="Figure 10. Map of central Peruvian Andes with known localities of three species of Nivaliodes." pageId="13" pageNumber="14">Fig. 10</figureCitation>
). It occurs along the upper limit of cloud forest and in lower puna grasslands.
</paragraph>
</subSubSection>
</treatment>
</document>
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