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<document id="53422F3F07D2BBF7989224C6FB17CE6E" ID-DOI="10.1016/j.rbe.2017.08.002" ID-ISSN="1806-9665" ID-Zenodo-Dep="13195627" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722635176695" checkinUser="felipe" docAuthor="González, Guillermo &amp; Almeida, Lúcia M." docDate="2017" docId="170F8796A2135C7BFFFBFC18736DFE0F" docLanguage="en" docName="RevBrasEntomol.61.4.330-338.pdf" docOrigin="Revista Brasileira de Entomologia 61 (4)" docSource="http://dx.doi.org/10.1016/j.rbe.2017.08.002" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Psyllobora lueri González &amp; Almeida, 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="335" masterDocId="EB36FFEEA2175C7EFF8EFF9A722EFFD1" masterDocTitle="Psyllobora picta (Germain) species complex (Coleoptera: Coccinellidae), with descriptions of two new species from Chile" masterLastPageNumber="338" masterPageNumber="330" pageNumber="334" updateTime="1722682715502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BA47A59E939B217B4603B7A89A575866">Psyllobora picta (Germain) species complex (Coleoptera: Coccinellidae), with descriptions of two new species from Chile</mods:title>
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<mods:namePart id="81F15977DFD3FBBFC67A8B3C17DFFE99">González, Guillermo</mods:namePart>
<mods:affiliation id="1D380630CAD5DB93C88D35F8A8C42CBB">La Reina, Santiago, Chile</mods:affiliation>
</mods:name>
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<mods:namePart id="526901FFC4EDA95ADCD1A13584D75D87">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="0C961827DB94859D686F8F547356B392">Universidade Federal do Paraná, Departamento de Zoologia, Laboratório de Sistemática e Bioecologia de Coleoptera, Curitiba, PR, Brazil</mods:affiliation>
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<mods:title id="AC23FFFE3CFF1B41437F4AE8CF439776">Revista Brasileira de Entomologia</mods:title>
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<mods:date id="4566228D1FA87658E61198C2F09954A4">2017</mods:date>
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<mods:number id="A12BBDA54501750970FABE51F6B7C8DC">2017-08-19</mods:number>
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<mods:url id="A76AA475014A33E7F0AF53C60E96CB4A">http://dx.doi.org/10.1016/j.rbe.2017.08.002</mods:url>
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<mods:classification id="09AECE91BFA9AB7FA350F14B36BEFA76">journal article</mods:classification>
<mods:identifier id="2D0F5A2859D706CF6A51A1B368C9BFDA" type="DOI">10.1016/j.rbe.2017.08.002</mods:identifier>
<mods:identifier id="4373DF001DE59E5A7E7E93CE39360109" type="ISSN">1806-9665</mods:identifier>
<mods:identifier id="E9037288DEF42701D147A5D44DC67C36" type="Zenodo-Dep">13195627</mods:identifier>
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<treatment id="170F8796A2135C7BFFFBFC18736DFE0F" LSID="urn:lsid:plazi:treatment:170F8796A2135C7BFFFBFC18736DFE0F" httpUri="http://treatment.plazi.org/id/170F8796A2135C7BFFFBFC18736DFE0F" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334">
<subSubSection id="D7BC650BA2135C7AFFFBFC18735FFC44" box="[117,369,898,917]" pageId="4" pageNumber="334" type="nomenclature">
<paragraph id="9F193680A2135C7AFFFBFC18735FFC44" blockId="4.[117,369,898,917]" box="[117,369,898,917]" pageId="4" pageNumber="334">
<heading id="C45181ECA2135C7AFFFBFC18735FFC44" bold="true" box="[117,369,898,917]" fontSize="8" level="2" pageId="4" pageNumber="334" reason="2">
<emphasis id="ADD2EA92A2135C7AFFFBFC18735FFC44" bold="true" box="[117,369,898,917]" pageId="4" pageNumber="334">
<taxonomicName id="58A64D03A2135C7AFFFBFC187334FC44" authorityName="González &amp; Almeida" authorityYear="2017" box="[117,282,898,917]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="lueri" status="sp. nov.">
<emphasis id="ADD2EA92A2135C7AFFFBFC187334FC44" bold="true" box="[117,282,898,917]" italics="true" pageId="4" pageNumber="334">Psyllobora lueri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6E157E9A2135C7AFEAEFC18735FFC44" box="[288,369,898,917]" pageId="4" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D7BC650BA2135C7AFFFBFC047332FC60" box="[117,284,926,945]" pageId="4" pageNumber="334" type="description">
<paragraph id="9F193680A2135C7AFFFBFC047332FC60" blockId="4.[85,756,926,1168]" box="[117,284,926,945]" pageId="4" pageNumber="334">
(
<figureCitation id="079D2A05A2135C7AFFF3FC0472DEFC60" box="[125,240,926,945]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 1525</figureCitation>
,
<figureCitation id="079D2A05A2135C7AFF74FC04733AFC60" box="[250,276,926,945]" captionStart="Fig" captionStartId="2.[890,917,1052,1066]" captionTargetBox="[838,1441,148,1021]" captionTargetId="figure-1196@2.[838,1441,148,1021]" captionTargetPageId="2" captionText="Fig. 53. Distribution map of Psyllobora picta species complex." figureDoi="http://doi.org/10.5281/zenodo.13195631" httpUri="https://zenodo.org/record/13195631/files/figure.png" pageId="4" pageNumber="334">53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D7BC650BA2135C7AFFFBFC2071B4F8CA" pageId="4" pageNumber="334" type="materials_examined">
<paragraph id="9F193680A2135C7AFFFBFC207098FC38" blockId="4.[85,756,926,1168]" pageId="4" pageNumber="334">
<materialsCitation id="2FCE3CDDA2135C7AFFFBFC207098FC38" collectingDate="2005-02-14" collectionCode="MHNS" collectorName="A. Luer" country="Chile" county="Chico" location="Region" municipality="Carrera" pageId="4" pageNumber="334" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="ADD2EA92A2135C7AFFFBFC2072F8FC1C" bold="true" box="[117,214,954,973]" pageId="4" pageNumber="334">
<typeStatus id="401D8822A2135C7AFFFBFC2072F8FC1C" box="[117,214,954,973]" pageId="4" pageNumber="334" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="89A0FD09A2135C7AFF52FC2072C3FC1C" box="[220,237,954,973]" count="1" pageId="4" pageNumber="334" type="male"></specimenCount>
: “
<collectingCountry id="E7B17610A2135C7AFE8DFC207317FC1C" box="[259,313,954,973]" name="Chile" pageId="4" pageNumber="334">Chile</collectingCountry>
, XI-R [
<location id="9A79605BA2135C7AFEF5FC2073EBFC1C" LSID="urn:lsid:plazi:treatment:170F8796A2135C7BFFFBFC18736DFE0F:9A79605BA2135C7AFEF5FC2073EBFC1C" box="[379,453,954,973]" country="Chile" county="Chico" municipality="Carrera" name="Region" pageId="4" pageNumber="334">Región</location>
]/Gral. [
<location id="9A79605BA2135C7AFD80FC20704CFC1C" LSID="urn:lsid:plazi:treatment:170F8796A2135C7BFFFBFC18736DFE0F:9A79605BA2135C7AFD80FC20704CFC1C" box="[526,610,954,973]" country="Chile" county="Chico" municipality="Carrera" name="General" pageId="4" pageNumber="334">General</location>
]
<collectingMunicipality id="7F7DACFAA2135C7AFDE0FC207097FC1C" box="[622,697,954,973]" pageId="4" pageNumber="334">Carrera</collectingMunicipality>
/
<collectingCountry id="E7B17610A2135C7AFD4EFC2070DAFC1C" box="[704,756,954,973]" name="Chile" pageId="4" pageNumber="334">Chile</collectingCountry>
<collectingCounty id="76784E0CA2135C7AFFDBFC4F72BAFC39" box="[85,148,981,1000]" pageId="4" pageNumber="334">Chico</collectingCounty>
/
<date id="EB181040A2135C7AFF12FC4F732FFC39" box="[156,257,981,1000]" pageId="4" pageNumber="334" value="2005-02-14">
<collectingDate id="FB5CE9A8A2135C7AFF12FC4F732FFC39" box="[156,257,981,1000]" pageId="4" pageNumber="334" value="2005-02-14">14-2-2005</collectingDate>
</date>
/coll. [leg.]
<collectorName id="32535356A2135C7AFEFFFC4C7396FC38" box="[369,440,982,1001]" pageId="4" pageNumber="334">A. Lüer</collectorName>
” “
<specimenCount id="89A0FD09A2135C7AFE5EFC4F73CFFC39" box="[464,481,981,1000]" count="1" pageId="4" pageNumber="334" type="male"></specimenCount>
1657” (mv) [
<collectionCode id="F9B7AE45A2135C7AFDEAFC4C7082FC38" box="[612,684,982,1001]" pageId="4" pageNumber="334">MHNS</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="9F193680A2135C7AFFFBFC6B73D8FB41" blockId="4.[85,756,926,1168]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFFFBFC6B72F0FBD5" bold="true" box="[117,222,1009,1028]" pageId="4" pageNumber="334">
<typeStatus id="401D8822A2135C7AFFFBFC6B72F0FBD5" box="[117,222,1009,1028]" pageId="4" pageNumber="334" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="89A0FD09A2135C7AFF62FC6B7350FBD5" box="[236,382,1009,1028]" count="20" pageId="4" pageNumber="334" type="generic">20 specimens</specimenCount>
):
<materialsCitation id="2FCE3CDDA2135C7AFE1EFC6B730DFBF1" collectingDate="2005-02-14" collectionCode="CPAL" collectorName="A. Luer" country="Chile" county="Chico" location="Region" municipality="Carrera" pageId="4" pageNumber="334" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFE1EFC6B739AFBD5" box="[400,436,1009,1028]" count="1" pageId="4" pageNumber="334" type="male">1 ♂</specimenCount>
: same data as holotype except “
<specimenCount id="89A0FD09A2135C7AFFD1FB97725EFBF1" box="[95,112,1037,1056]" count="1" pageId="4" pageNumber="334" type="male"></specimenCount>
148” (mv) [
<collectionCode id="F9B7AE45A2135C7AFF6DFB977331FBF1" box="[227,287,1037,1056]" pageId="4" pageNumber="334">CPAL</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="2FCE3CDDA2135C7AFEA5FB9772F1FBED" collectingDate="2005-02-16" collectionCode="MHNS" collectorName="A. Luer" country="Chile" county="Chico" location="Region" municipality="Carrera" pageId="4" pageNumber="334" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFEA5FB977363FBF1" box="[299,333,1037,1056]" count="1" pageId="4" pageNumber="334" type="male">1 ♂</specimenCount>
and
<specimenCount id="89A0FD09A2135C7AFEF2FB9773B6FBF1" box="[380,408,1037,1056]" count="2" pageId="4" pageNumber="334" type="female">2 ♀</specimenCount>
: same data except date
<date id="EB181040A2135C7AFD06FB9770DDFBF1" box="[648,755,1037,1056]" pageId="4" pageNumber="334" value="2005-02-16">
<collectingDate id="FB5CE9A8A2135C7AFD06FB9770DDFBF1" box="[648,755,1037,1056]" pageId="4" pageNumber="334" value="2005-02-16">16-2-2005</collectingDate>
</date>
(mv) [
<collectionCode id="F9B7AE45A2135C7AFF1FFBB372F7FBED" box="[145,217,1065,1084]" pageId="4" pageNumber="334">MHNS</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="2FCE3CDDA2135C7AFF66FBB37366FB89" collectingDate="2005-02-12" collectingDateMax="2005-02-17" collectingDateMin="2005-02-12" collectionCode="CPAL" collectorName="A. Luer" country="Chile" county="Chico" location="Region" municipality="Carrera" pageId="4" pageNumber="334" specimenCount="12" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFF66FBB37356FBED" box="[232,376,1065,1084]" count="12" pageId="4" pageNumber="334" type="generic">12 specimens</specimenCount>
: same data except several dates from
<date id="EB181040A2135C7AFFDBFBDF72D3FB89" box="[85,253,1093,1112]" pageId="4" pageNumber="334" value="2005-02-12" valueMax="2005-02-17" valueMin="2005-02-12">
<collectingDate id="FB5CE9A8A2135C7AFFDBFBDF72D3FB89" box="[85,253,1093,1112]" pageId="4" pageNumber="334" value="2005-02-12" valueMax="2005-02-17" valueMin="2005-02-12">12 to 172-2005</collectingDate>
</date>
[
<collectionCode id="F9B7AE45A2135C7AFE89FBDF736DFB89" box="[263,323,1093,1112]" pageId="4" pageNumber="334">CPAL</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="2FCE3CDDA2135C7AFEDEFBDF73BAFBA5" collectingDate="2016-02-10" collectorName="R. Honour" country="Chile" location="Chile" pageId="4" pageNumber="334" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFEDEFBDF735DFB89" box="[336,371,1093,1112]" count="2" pageId="4" pageNumber="334" type="male">2 ♂</specimenCount>
: “
<date id="EB181040A2135C7AFE07FBDF702FFB89" box="[393,513,1093,1112]" pageId="4" pageNumber="334" value="2016-02-10">
<collectingDate id="FB5CE9A8A2135C7AFE07FBDF702FFB89" box="[393,513,1093,1112]" pageId="4" pageNumber="334" value="2016-02-10">10/02/2016</collectingDate>
</date>
/
<quantity id="585E9B65A2135C7AFD87FBDF706AFB89" box="[521,580,1093,1112]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="334" unit="km" value="13.0">13 km</quantity>
W
<collectingCountry id="E7B17610A2135C7AFDEBFBDF70B7FB89" box="[613,665,1093,1112]" name="Chile" pageId="4" pageNumber="334">Chile</collectingCountry>
Chico/XI Región/
<collectingCountry id="E7B17610A2135C7AFF2CFBFB72F8FBA5" box="[162,214,1121,1140]" name="Chile" pageId="4" pageNumber="334">Chile</collectingCountry>
leg.
<collectorName id="32535356A2135C7AFE84FBFB735CFBA5" box="[266,370,1121,1140]" pageId="4" pageNumber="334">R. Honour</collectorName>
S.”
</materialsCitation>
<materialsCitation id="2FCE3CDDA2135C7AFE28FBFB70EAFBA5" box="[422,708,1117,1140]" collectionCode="CPRH" country="Chile" pageId="4" pageNumber="334" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFE28FBFB73E6FBA5" box="[422,456,1121,1140]" count="1" pageId="4" pageNumber="334" type="male">♂♂</specimenCount>
1733
<emphasis id="ADD2EA92A2135C7AFD8DFBC77022FBBA" bold="true" box="[515,524,1117,1131]" pageId="4" pageNumber="334">
<superScript id="68D39BC8A2135C7AFD8DFBC77022FBBA" attach="left" box="[515,524,1117,1131]" fontSize="6" pageId="4" pageNumber="334">ƖƖ</superScript>
</emphasis>
(mv) [
<collectionCode id="F9B7AE45A2135C7AFDDDFBFB70BFFBA5" box="[595,657,1121,1140]" pageId="4" pageNumber="334">CPRH</collectionCode>
] and
</materialsCitation>
<materialsCitation id="2FCE3CDDA2135C7AFD42FBFB73DCFB41" collectionCode="CPMD" country="Chile" pageId="4" pageNumber="334" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="89A0FD09A2135C7AFD42FBFB70C2FBA5" box="[716,748,1121,1140]" count="2" pageId="4" pageNumber="334" type="female">2 ♀</specimenCount>
: same data except “
<specimenCount id="89A0FD09A2135C7AFE9AFBE77305FB41" box="[276,299,1149,1168]" count="1" pageId="4" pageNumber="334" type="female">♀♀</specimenCount>
1733” (ab) [
<collectionCode id="F9B7AE45A2135C7AFE29FBE773C3FB41" box="[423,493,1149,1168]" pageId="4" pageNumber="334">CPMD</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="9F193680A2135C7AFFDBFB21729DFB1E" blockId="4.[85,179,1211,1231]" box="[85,179,1211,1231]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFFDBFB21729DFB1E" box="[85,179,1211,1231]" italics="true" pageId="4" pageNumber="334">Diagnosis</emphasis>
</paragraph>
<paragraph id="9F193680A2135C7AFFFBFB6E735BF9EB" blockId="4.[85,757,1268,1985]" pageId="4" pageNumber="334">
The dorsal color yellowish brown with ten light-brown spots with dark border, ringed, and the very thick tube of the male penis (its width 1/6 of the maximum length), separate
<taxonomicName id="58A64D03A2135C7AFDBBFAB07059FAEF" authorityName="González &amp; Almeida" authorityYear="2017" box="[565,631,1322,1342]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="lueri">
<emphasis id="ADD2EA92A2135C7AFDBBFAB07059FAEF" box="[565,631,1322,1342]" italics="true" pageId="4" pageNumber="334">P. lueri</emphasis>
</taxonomicName>
from the big majority of the genera species. From the similar looking
<taxonomicName id="58A64D03A2135C7AFD1AFADC70DAFA8B" authorityName="González &amp; Almeida" authorityYear="2017" box="[660,756,1350,1370]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pauline">
<emphasis id="ADD2EA92A2135C7AFD1AFADC70DAFA8B" box="[660,756,1350,1370]" italics="true" pageId="4" pageNumber="334">P. pauline</emphasis>
</taxonomicName>
is distinguished by the penis guide nearly straight with slightly converging sides and the elytral spots 214567 almost always linked together, forming an “S” shape, while from the other similar looking species
<taxonomicName id="58A64D03A2135C7AFF79FA2C73A2FA1B" authorityName="Boheman Penis" authorityYear="1858" box="[247,396,1462,1482]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="bicongregata">
<emphasis id="ADD2EA92A2135C7AFF79FA2C73A2FA1B" box="[247,396,1462,1482]" italics="true" pageId="4" pageNumber="334">P. bicongregata</emphasis>
</taxonomicName>
is distinguished by the penis guide elongate more than 4.0 times longer than wide.Otherwise
<taxonomicName id="58A64D03A2135C7AFD19FA4870DAFA37" authorityName="González &amp; Almeida" authorityYear="2017" box="[663,756,1490,1510]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="pauline">
<emphasis id="ADD2EA92A2135C7AFD19FA4870DAFA37" box="[663,756,1490,1510]" italics="true" pageId="4" pageNumber="334">P.pauline</emphasis>
</taxonomicName>
is restricted to Copiapó to
<taxonomicName id="58A64D03A2135C7AFEEFFA7573FDF9D3" authorityName="Arias" authorityYear="2008" box="[353,467,1519,1538]" class="Insecta" family="Elateridae" genus="Llanquihue" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="genus">Llanquihue</taxonomicName>
provinces from
<collectingCountry id="E7B17610A2135C7AFDF7FA757080F9D3" box="[633,686,1519,1538]" name="Chile" pageId="4" pageNumber="334">Chile</collectingCountry>
, and
<taxonomicName id="58A64D03A2135C7AFD6DFA7472FDF9CF" authorityName="Boheman Penis" authorityYear="1858" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="bicongregata">
<emphasis id="ADD2EA92A2135C7AFD6DFA7472FDF9CF" italics="true" pageId="4" pageNumber="334">P. bicongregata</emphasis>
</taxonomicName>
to
<collectingCountry id="E7B17610A2135C7AFF77F9917371F9CF" box="[249,351,1547,1566]" name="Argentina" pageId="4" pageNumber="334">Argentina</collectingCountry>
,
<collectingCountry id="E7B17610A2135C7AFEE4F991738BF9CF" box="[362,421,1547,1566]" name="Brazil" pageId="4" pageNumber="334">Brasil</collectingCountry>
,
<collectingCountry id="E7B17610A2135C7AFE3FF9917020F9CF" box="[433,526,1547,1566]" name="Paraguay" pageId="4" pageNumber="334">Paraguay</collectingCountry>
and
<collectingCountry id="E7B17610A2135C7AFDCAF99170B3F9CF" box="[580,669,1547,1566]" name="Uruguay" pageId="4" pageNumber="334">Uruguay</collectingCountry>
, regions where
<taxonomicName id="58A64D03A2135C7AFF15F9BF72F1F9E8" authorityName="González &amp; Almeida" authorityYear="2017" box="[155,223,1573,1593]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="lueri">
<emphasis id="ADD2EA92A2135C7AFF15F9BF72F1F9E8" box="[155,223,1573,1593]" italics="true" pageId="4" pageNumber="334">P. lueri</emphasis>
</taxonomicName>
is not present.
</paragraph>
<paragraph id="9F193680A2135C7AFFFBF9D972F3F8BC" blockId="4.[85,757,1268,1985]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFFFBF9D97376F984" bold="true" box="[117,344,1602,1622]" pageId="4" pageNumber="334">
<typeStatus id="401D8822A2135C7AFFFBF9D972F8F987" box="[117,214,1603,1622]" pageId="4" pageNumber="334" type="holotype">Holotype</typeStatus>
description
</emphasis>
: Male. Length: 2.4 mm, width: 1.9 mm. Oval with maximum width near middle of elytra. Back yellowish brown, with five dark-brown spots on pronotum and two small supernumerary spots near border, elytra with ten brown spots with darker, irregular borders, with spot ten widely scattered and a clearer supernumerary spot between spots 5 and 7. Spots 3214578 joined together, forming an “S” pattern. Legs, antennae and mouthparts yellowish brown. Ventral surface light brown with dark-brown metaventrite. Dorsal punctation inconspicuous, irregular, surface opaque, without reticulation (
<figureCitation id="079D2A05A2135C7AFFD3F8C372FEF8BD" box="[93,208,1881,1900]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 1520</figureCitation>
).
</paragraph>
<paragraph id="9F193680A2135C7AFFFBF8EF761CFF34" blockId="4.[85,757,1268,1985]" lastBlockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFFFBF8EF7282F859" bold="true" box="[117,172,1909,1928]" pageId="4" pageNumber="334">Head</emphasis>
: Color light brown, with two brown spots between eyes at insertion of antennas and large dark-brown transverse spot above eyes, extending laterally on vertex. Antennas, labrum, maxillary palps and labium light brown. Eyes rounded frons three times width of eye. Punctation irregular, denser laterally, with points separated by two or more diameters.
</paragraph>
<paragraph id="9F193680A2135C7AFCCAFF747785FE31" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAFF7471A1FED0" bold="true" box="[836,911,238,257]" pageId="4" pageNumber="334">Thorax</emphasis>
: Pronotum light brown, with anterior and lateral borders yellowish and translucent. Small central brown spot near base, surrounded with semicircle of four brown spots, two posterior spots touching base and two anterior spots half length of pronotum, all equidistant from each other, and two small, pale, somewhat irregular spots on each side. Scutellum small, triangular, dark brown (
<figureCitation id="079D2A05A2135C7AFC29FE0C7639FE78" box="[935,1047,405,425]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 15, 17</figureCitation>
). Hypomeres and mesosternum yellowish brown, metaventrite dark brown. Legs yellowish brown. Pronotum with almost imperceptible punctation, points widely separated.
</paragraph>
<paragraph id="9F193680A2135C7AFCCAFE737140FBDF" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAFE7371ADFE2D" bold="true" box="[836,899,489,508]" pageId="4" pageNumber="334">Elytra</emphasis>
: Slightly yellowish brown, with ten brown spots, with somewhat darker borders (3:1:2:1:2:1) and clearer supernumerary spot in space between spots 5, 7 and suture. Spot 1 very large, almost ¼ length of elytra, spot 2 slightly smaller and spot 3 even smaller, adjacent and in contact with base, connected together at middle; spot 4 longitudinal-oval, occupying 1/5 to 2/5 of length of the elytra, close to suture but not touching, near spots 1 and 5; spots 5 and 6 nearly half length of the elytra, spot 5 large and connected to spots 4 and 7, and spot 6 separate and half divided longitudinally, homogeneous dark brown; spot 7 immediately behind large spot 5, semicircular, connected at outer border, both forming large “C” shaped spot, in space between these and suture, with supernumerary spot much clearer, small and irregular; spot 8 near suture, half of diameter of spot 4, spot 9 on same transverse line but still smaller, near lateral border, dark brown; spot 10 disjointed in three small irregular inconspicuous spots, arranged in triangle (
<figureCitation id="079D2A05A2135C7AFACEFC167798FC4E" box="[1344,1462,908,927]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 1518</figureCitation>
). Punctation with large distinct points, irregularly distributed, separated by about two diameters on average. Pale yellow epipleuron strongly descendant, with small dark spot in middle of outer border.
</paragraph>
<paragraph id="9F193680A2135C7AFCCAFB8D7663FB4B" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAFB8D7185FBFB" bold="true" box="[836,939,1047,1066]" pageId="4" pageNumber="334">Abdomen</emphasis>
: Brown, with middle of first two ventrites dark brown. Postcoxal lines descending diagonally at 45
<emphasis id="ADD2EA92A2135C7AFAA9FBB57701FBEC" bold="true" box="[1319,1327,1071,1085]" pageId="4" pageNumber="334">
<superScript id="68D39BC8A2135C7AFAA9FBB57701FBEC" attach="none" box="[1319,1327,1071,1085]" fontSize="6" pageId="4" pageNumber="334"></superScript>
</emphasis>
, bent to touch back border attached to it. Ventrite V emarginate around posterior margin, ventrite VI as short as half of 5 ventrite, gently rounded with shallow notch in center.
</paragraph>
<paragraph id="9F193680A2135C7AFCCAFB397155F9A5" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAFB3971F5FB64" bold="true" box="[836,987,1186,1206]" pageId="4" pageNumber="334">Male genitalia</emphasis>
: Tegmen 4.5 times longer than wide, phallobase subtrapezoidal, 1.1 times longer than wide and with posterior bor- der slightly rounded. Terminal strut 1/3 length of phallobase. Penis guide in dorsal view with slightly converging sides, slightly sinuous, narrowing to 3/4 of width at 5/6 of length, apical 1/6 ending in triangle, with concave sides and slight apical projection; in lateral view triangular, sinuous, bent toward parameres in distal 1/6 of apex. Parameres in dorsal view narrow and slightly curved toward basal lobe, 1/5 shorter than this, with pubescence in distal half, exceeding paramere in almost half of length (
<figureCitation id="079D2A05A2135C7AFA8CFA047759FA60" box="[1282,1399,1438,1457]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 2224</figureCitation>
). Penis width about 1/6 of maximum length, in a semicircle in basal half, slightly curved to 2/3 length, distal 1/3 straight. Apex 1/6 of length and slightly wider than penis tube, separated from it by a sclerosed union at widely oblique angle, with ejaculatory duct “S” shaped and sclerose spine on apex; penis capsule with internal branch hook-shaped, outer branch twice as wide as internal, basal margin convex (
<figureCitation id="079D2A05A2135C7AFCA2F9FB7141F9A5" box="[812,879,1633,1652]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="9F193680A2135C7AFCCAF9E771B4F8CA" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAF9E771BFF941" bold="true" box="[836,913,1661,1680]" pageId="4" pageNumber="334">Female</emphasis>
: Similar to male. Spermatheca “C” shaped, about ten times longer than wide, narrowing regularly toward apex, nodulus lateral and short, ramus 1/10 of total length, seminal duct very short. Infundibulum “Y” shaped, half size of spermathecal (
<figureCitation id="079D2A05A2135C7AFAFAF94B7798F932" box="[1396,1462,1744,1764]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Fig. 25</figureCitation>
). Coxites paddle-shaped, with very wide, pubescent apex and evident stylus.
</paragraph>
</subSubSection>
<subSubSection id="D7BC650BA2135C7BFCCAF8BE72E4FE5B" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334" type="description">
<paragraph id="9F193680A2135C7AFCCAF8BE761CF812" blockId="4.[804,1475,155,1987]" pageId="4" pageNumber="334">
<emphasis id="ADD2EA92A2135C7AFCCAF8BE7186F8E6" bold="true" box="[836,936,1828,1847]" pageId="4" pageNumber="334">Variation</emphasis>
: Length 2.12.6 mm. Color nearly homogenous, spots varying from light brown dark-ringed to homogeneous dark brown. The spots, especially on the pronotum and supernumerary of the elytra tend to disappear, while the elytral spots may be partially disjointed to rarely losing the “S” shape, the posterior spot sometimes complete (
<figureCitation id="079D2A05A2135C7AFC1EF82A760BF812" box="[912,1061,1968,1987]" captionStart="Figs" captionStartId="1.[833,868,1053,1067]" captionTargetBox="[851,1485,153,1022]" captionTargetId="figure-523@1.[848,1489,150,1023]" captionTargetPageId="1" captionText="Figs. 125. 114: Psyllobora pauline sp. nov.: 15: male holotype habitus (dorsal, lateral,frontal,posterior and ventral detail);69:variations (dorsal view); 10: male sipho; 1113: male tegmen (dorsal, lateral and basal lobe detail); 14: female genitalia (spermatheca and infundibulum). 1525: Psyllobora lueri n. sp.: 1518: male holotype habitus (dorsal, lateral, frontal and posterior); 1920: variations (dorsal view); 21: male penis; 2224: male tegmen (dorsal, lateral and basal lobe detail). 25: female genitalia (spermatheca and infundibulum)." figureDoi="http://doi.org/10.5281/zenodo.13195629" httpUri="https://zenodo.org/record/13195629/files/figure.png" pageId="4" pageNumber="334">Figs. 15, 19, 20</figureCitation>
).
</paragraph>
<paragraph id="9F193680A2125C7BFFFCFF0D72F6FF7A" blockId="5.[114,216,151,171]" box="[114,216,151,171]" pageId="5" pageNumber="335">
<emphasis id="ADD2EA92A2125C7BFFFCFF0D72F6FF7A" box="[114,216,151,171]" italics="true" pageId="5" pageNumber="335">Discussion</emphasis>
</paragraph>
<paragraph id="9F193680A2125C7BFF1CFF55736DFE83" blockId="5.[114,785,207,478]" pageId="5" pageNumber="335">
<taxonomicName id="58A64D03A2125C7BFF1CFF557303FF32" authorityName="González &amp; Almeida" authorityYear="2017" box="[146,301,207,227]" class="Insecta" family="Coccinellidae" genus="Psyllobora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="lueri" status="sp. nov.">
<emphasis id="ADD2EA92A2125C7BFF1CFF557303FF32" box="[146,301,207,227]" italics="true" pageId="5" pageNumber="335">Psyllobora lueri</emphasis>
</taxonomicName>
<emphasis id="ADD2EA92A2125C7BFEB8FF4A73A4FF32" bold="true" box="[310,394,208,227]" pageId="5" pageNumber="335">
<taxonomicNameLabel id="B6E157E9A2125C7BFEB8FF4A73A4FF32" box="[310,394,208,227]" pageId="5" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs in the coloration pattern and in the male genitalia (see species key). Also, it is geographically isolated from other species of the genus, probably occurring in
<collectingCountry id="E7B17610A2125C7BFFFCFEB972F8FEE7" box="[114,214,291,310]" name="Argentina" pageId="5" pageNumber="335">Argentina</collectingCountry>
to
<collectingRegion id="5D62F862A2125C7BFF7CFEB973B4FEE7" box="[242,410,291,310]" country="Argentina" name="Chubut" pageId="5" pageNumber="335">Chubut Province</collectingRegion>
, where no
<taxonomicName id="58A64D03A2125C7BFD8DFEB8709BFEE7" box="[515,693,290,310]" pageId="5" pageNumber="335">
<emphasis id="ADD2EA92A2125C7BFD8DFEB87046FEE7" box="[515,616,290,310]" italics="true" pageId="5" pageNumber="335">Psyllobora</emphasis>
species
</taxonomicName>
has been recorded previously.
</paragraph>
<paragraph id="9F193680A2125C7BFF1CFEC172E4FE5B" blockId="5.[114,785,207,478]" pageId="5" pageNumber="335">
<emphasis id="ADD2EA92A2125C7BFF1CFEC1738BFEBF" bold="true" box="[146,421,347,366]" pageId="5" pageNumber="335">Geographical distribution</emphasis>
: Austral
<collectingCountry id="E7B17610A2125C7BFD8CFEC17019FEBF" box="[514,567,347,366]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
,
<collectingCountry id="E7B17610A2125C7BFDCEFEC1705AFEBF" box="[576,628,347,366]" name="Chile" pageId="5" pageNumber="335">Chile</collectingCountry>
Chico Province (
<figureCitation id="079D2A05A2125C7BFFF4FEED7293FE5B" box="[122,189,375,394]" captionStart="Fig" captionStartId="2.[890,917,1052,1066]" captionTargetBox="[838,1441,148,1021]" captionTargetId="figure-1196@2.[838,1441,148,1021]" captionTargetPageId="2" captionText="Fig. 53. Distribution map of Psyllobora picta species complex." figureDoi="http://doi.org/10.5281/zenodo.13195631" httpUri="https://zenodo.org/record/13195631/files/figure.png" pageId="5" pageNumber="335">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D7BC650BA2125C7BFF1CFE09736DFE0F" pageId="5" pageNumber="335" type="etymology">
<paragraph id="9F193680A2125C7BFF1CFE09736DFE0F" blockId="5.[114,785,207,478]" pageId="5" pageNumber="335">
<emphasis id="ADD2EA92A2125C7BFF1CFE09732AFE77" bold="true" box="[146,260,403,422]" pageId="5" pageNumber="335">Etymology</emphasis>
: This species is named for Alfredo Lüer, dedicated Chilean entomologist and collector of many of the known specimens of this species.
</paragraph>
</subSubSection>
</treatment>
</document>