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<document id="40118A94D0CC66B6D8C257DEEFD84835" ID-CLB-Dataset="34707" ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="16B6406E056C06275E5C7953416D99DE" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus crenatus Lewis 1888" docType="treatment" docVersion="5" lastPageNumber="73" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="71" updateTime="1732934305285" updateUser="ExternalLinkService">
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<mods:title id="77B9B2724DA8E21C3A2F06C3F24836C4">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart id="851FD12463E9EAB2AE8B4B2ED5517D0B">Caterino, Michael S.</mods:namePart>
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<mods:namePart id="D5F9C4A9989BAEB5242B432DE5FB1008">Tishechkin, Alexey K.</mods:namePart>
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<treatment id="16B6406E056C06275E5C7953416D99DE" ID-GBIF-Taxon="152040986" LSID="urn:lsid:plazi:treatment:16B6406E056C06275E5C7953416D99DE" httpUri="http://treatment.plazi.org/id/16B6406E056C06275E5C7953416D99DE" lastPageId="72" lastPageNumber="73" pageId="70" pageNumber="71" scope_family="Histeridae" scope_order="Coleoptera" scope_tribe="Exosternini">
<subSubSection id="FACDAB7E6DFDEDE28BE2CF3D85E2323A" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="51762F4750CED1565D31E81EFA07B33B" pageId="70" pageNumber="71">
<taxonomicName id="38D7292D88B8C922F7158678A8845FB7" ID-CoL="74NJH" LSID="http://species-id.net/wiki/Operclipygus_crenatus" authority="Lewis, 1888" authorityName="Lewis" authorityYear="1888" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="crenatus">
<pageBreakToken id="9A57DEFE34BD9475BF3713C247E8B51A" pageId="70" pageNumber="71" start="start">Operclipygus</pageBreakToken>
crenatus (Lewis, 1888)
</taxonomicName>
Figs 17
<normalizedToken id="C41C6A63E55EFE1FF56D32E6A009ED9F" originalValue="C–D">C-D</normalizedToken>
18E, GMap 6
</paragraph>
</subSubSection>
<subSubSection id="F3D7C16F4978C530444542DC8EC63434" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="C77B99C790C78F64D932AAD380B8D49D" pageId="70" pageNumber="71">
<taxonomicName id="E3A420673F53B0C99F9A2316887E676D" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister crenatus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="crenatus">Phelister crenatus</taxonomicName>
Lewis, 1888: 192;
<bibRefCitation id="CE13E9D7A759FF7F97CC306219840F90" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 257);
<taxonomicName id="3E36534DBBD60878F0A06ACB75E3CB05" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
:
<bibRefCitation id="67A83E16DD01AA5E603152EEF2B8BA25" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 257).
</paragraph>
<paragraph id="2E6FADC8F5BC6C8068D4106E37EC1189" pageId="70" pageNumber="71">
<taxonomicName id="19E3A6540EF1BEE6FF3A16CF406C8378" genus="Phelisteroides" lsidName="Phelisteroides miladae" pageId="70" pageNumber="71" rank="species" species="miladae">Phelisteroides miladae</taxonomicName>
Wenzel &amp; Dybas, 1941: 450;
<taxonomicName id="1B52E6E9BDE7906646CEED9E1FDA2DF1" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus miladae" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="miladae">Operclipygus miladae</taxonomicName>
:
<bibRefCitation id="7D535EA0128FE7DC594098F3D2B15A4A" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 257). syn. n.
</paragraph>
<paragraph id="90CC780DEAC16D6C4E4A5F419DDA9B59" pageId="70" pageNumber="71">
<taxonomicName id="ECB39899CF6810DFF8A5E97DC2BB173F" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister propygidialis" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="propygidialis">Pseudister propygidialis</taxonomicName>
Hinton, 1935e: 12;
<taxonomicName id="557A70B3C0AD3A0E0D32FA9D8CB309BB" genus="Phelisteroides" lsidName="Phelisteroides propygidialis" pageId="70" pageNumber="71" rank="species" species="propygidialis">Phelisteroides propygidialis</taxonomicName>
:
<bibRefCitation id="A138F9D3960F3065A95C3B61FD0ECC16" author="Wenzel, RL" journalOrPublisher="Fieldiana, Zoology" pageId="377" pageNumber="378" pagination="433 - 472" title="New and little known neotropical Histeridae (Coleoptera)." volume="22" year="1941">Wenzel and Dybas (1941</bibRefCitation>
: 454).
<taxonomicName id="2D7B3F56D76FA6796DA7D7779A6EEFA8" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus propygidialis" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="propygidialis">Operclipygus propygidialis</taxonomicName>
:
<bibRefCitation id="BBDA4F28341196B2A178E8F628A537E4" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel (1976</bibRefCitation>
: 257). syn. n.
</paragraph>
</subSubSection>
<subSubSection id="CA32EF48FC95DE06150EEF244E048106" pageId="70" pageNumber="71" type="type locality">
<paragraph id="BB9DF51844D941D0B6B1CD996B1EE04C" pageId="70" pageNumber="71">Type locality.</paragraph>
<paragraph id="67155B8B92E1CDB3101510AD2A028AC6" pageId="70" pageNumber="71">
PANAMA:
<normalizedToken id="839B97C13DF82B5ACAFEBE10C4964240" originalValue="Chiriquí">Chiriqui</normalizedToken>
:
<normalizedToken id="9DD0083E8ADA0399148E1239337DEA5D" originalValue="Barú">Baru</normalizedToken>
Volcano [
<geoCoordinate id="DC5220CADF9F0E870263A8F1401644AF" direction="north" orientation="latitude" precision="925" value="8.8">8°48'N</geoCoordinate>
,
<geoCoordinate id="80D1FDD2ABB05B48D65495BFFD902066" direction="west" orientation="longitude" precision="92" value="-82.541664">82°32.5'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="9376D6ACEEB52F9E4BC2E9E06C3AB39F" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="type material">
<paragraph id="FD8BD80FC243DE7F49D99B1FE855F8A1" pageId="70" pageNumber="71">Type material.</paragraph>
<paragraph id="A949C79C2F7CDB7F5B7BA570EA64A1C8" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<taxonomicName id="32972D7099536ADD29558C2EE2F23DAC" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister crenatus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="crenatus">Phelister crenatus</taxonomicName>
: Lectotype, here designated, probably male: &quot;Sp. figured
<normalizedToken id="F7481378B968295260D3C98171A736A6" originalValue="”/”">&quot;/&quot;</normalizedToken>
V. de
<normalizedToken id="7E87A8E1DDAE2A7AC0CFFFC1DC8C209C" originalValue="Chiriquí">Chiriqui</normalizedToken>
, 3-4000 ft. Champion&quot; / &quot;LECTOTYPE
<taxonomicName id="70F7BB8D1612DC230C0F9E0D8BD385B5" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister crenatus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="crenatus">Phelister crenatus</taxonomicName>
Lewis 1864, M.S.Caterino &amp; A.K.Tishechkin des. 2010&quot; (BMNH). This species was described from an unspecified number of specimens, and the lectotype designation
<pageBreakToken id="1A17C7D304DC3CA0AF1ECAD607871F01" pageId="71" pageNumber="72" start="start">fixes</pageBreakToken>
primary type status on the only known original specimen.
<taxonomicName id="721E6CC65831C9E443C2A67EE01489CB" genus="Phelisteroides" lsidName="Phelisteroides miladae" pageId="71" pageNumber="72" rank="species" species="miladae">Phelisteroides miladae</taxonomicName>
: Holotype female (FMNH): &quot;Villavicencio, Meta, Colomb. VII:25:38
<normalizedToken id="4B68A07F292FC321ED8788BE11554751" originalValue="”/”">&quot;/&quot;</normalizedToken>
under rotten bark
<normalizedToken id="DBF7332A67DE870AE829FC8C2672A2A9" originalValue="”/”">&quot;/&quot;</normalizedToken>
Coll. by H. Dybas
<normalizedToken id="0FCC5DE6FC5101883D1D041AB417B09F" originalValue="”/”">&quot;/&quot;</normalizedToken>
Type
<taxonomicName id="DEA2AE2DBA5B5165B9439685AA510C76" genus="Phelisteroides" lsidName="Phelisteroides miladae" pageId="71" pageNumber="72" rank="species" species="miladae">Phelisteroides miladae</taxonomicName>
Wenz. &amp; Dybas&quot;; examined 2006.
<taxonomicName id="71613E83F0D8EA23AF5566BA52A1A71F" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister propygidialis" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propygidialis">Pseudister propygidialis</taxonomicName>
: Type repository unknown; published type locality: Morelos, Cuernevaca, Mexico.
</paragraph>
</subSubSection>
<subSubSection id="BF9AC23723AEB57017B9C244582D5CCD" pageId="71" pageNumber="72" type="other material">
<paragraph id="9B30ADCBEDBB93C3BC114E067B855F2D" pageId="71" pageNumber="72">Other material.</paragraph>
<paragraph id="82701BBE5AF5FAA31CE26156A5C1E9EC" pageId="71" pageNumber="72">
ARGENTINA: Jujuy: 1: Parque Nac. Calilegua, Aguas Negras, 440m, 18-28.xii.1987, malaise/FIT, S. &amp; J. Peck (CHSM); 2: Parque Nac. Calilegua, El Cortaderal, km6, 800m, 18-28.xii.1987, malaise/FIT, forest, S. &amp; J. Peck (CHSM). COSTA RICA: Puntarenas: 1: Parque Nac. Corcovado, Sirena Stn. Corcovado Trail,
<geoCoordinate id="E3E9950B14CA1348868FD48B04F1DA4D" direction="north" orientation="latitude" precision="15" value="8.485278">8°29'7&quot;N</geoCoordinate>
,
<geoCoordinate id="003671CED726E6A669434C631F8A44B9" direction="west" orientation="longitude" precision="15" value="-83.5775">83°34'39&quot;W</geoCoordinate>
, 150m, 28.
<normalizedToken id="3FFD793614B77FF363B0420F8681658C" originalValue="vi–">vi-</normalizedToken>
1.vii.2000, FIT, Z. Falin (SEMC); 1: Rancho Quemado, Peninsula de Osa, 200m, v.1992, F. Quesada y G. Varela, (INBIO), 1: xii.1991, F. Quesada, (INBIO); 1: Est. Agujas, Sendero Ajo, 300m, 24-26.ix.1996, in fruit, A. Azofeifa, (INBIO). Cartago: 1: Ref. Nac. Fauna Silv. Tapanti, Quebrada Segunda, 1250m, iii.1992, G. Mora, (INBIO); same data except as noted: 1: ix.1992, (INBIO); 5: v.1992 (INBIO), 1: v.1993, (INBIO), 1: 1250m, vi.1992, (INBIO), 20: 1250m, vii.1992, (INBIO), 2: 1250m, x.1992, (INBIO), 5: 1300m, vi.1992, (INBIO), 1: Grano de Oro, Chirripo, Turrialba, 1120m, iii.1993, P. Campos, (INBIO). FRENCH GUIANA: 1: Montagne des Chevaux,
<geoCoordinate id="832224C77E4ABD720BBB4ADF1E9CFA01" direction="north" orientation="latitude" precision="925" value="4.7166667">4°43'N</geoCoordinate>
,
<geoCoordinate id="20CFEB00696C50B2598511FD6C903EED" direction="west" orientation="longitude" precision="925" value="-52.4">52°24'W</geoCoordinate>
, 11.x.2009, FIT, SEAG (CHND). PERU: Cusco: 1: Consuelo, Manu rd. km 165, 11.x.1982, palm litter, L. Watrous &amp; G. Mazurek (FMNH); same data except as noted: 5: 3.x.1982 (FMNH), 1: 5.x.1982 (FMNH), 2: 8.x.1982 (FMNH). Huanuco: 1:Tingo Maria region, 6-14.vi.1987, F. Woytkowski (SEMC).
<normalizedToken id="A0924334C241303B2980979E27986D40" originalValue="Junín">Junin</normalizedToken>
: 3: La Merced, 9.5km NE La Merced-Villa Rica Rd.,
<geoCoordinate id="FBB871BFEAC1598119C9C949A3A9A9C5" direction="south" orientation="latitude" precision="15" value="-10.9783325">10°58'42&quot;S</geoCoordinate>
,
<geoCoordinate id="7503944026CC0344B7DB4F5A6F23DAE8" direction="west" orientation="longitude" precision="15" value="-75.305">75°18'18&quot;W</geoCoordinate>
, 880m, 15-21.x.1999, FIT, R. Brooks (SEMC); 1: 7.5km NE Puerto Ocopa, Cananeden,
<geoCoordinate id="FA8E303B98998DEB8BFBA8AB3BD3D3FC" direction="south" orientation="latitude" precision="92" value="-11.081667">11°4.9'S</geoCoordinate>
,
<geoCoordinate id="A7A76E3E5FF04D51194E08BB7D29BAB2" direction="west" orientation="longitude" precision="92" value="-74.26833">74°16.1'W</geoCoordinate>
, 1180m, 5-9.i.2007, A.V. Petrov (AKTC). Madre de Dios: 1:Manu National Park, Rio Alto Madre de Dios, Pantiacolla Lodge,
<geoCoordinate id="CD7E1AE5BE7772A1F605B28A8BB41422" direction="south" orientation="latitude" precision="92" value="-12.655">12°39.3'S</geoCoordinate>
,
<geoCoordinate id="B849EDD868CF4CE152BD2D6C6215C5AF" direction="west" orientation="longitude" precision="92" value="-71.23167">71°13.9'W</geoCoordinate>
, 14-19.xi.2007, FIT, D. Brzoska (SEMC). VENEZUELA: Monagas: 3: Caripe, Cueva
<normalizedToken id="AF4DE028B61DC7B23CC4D15CE2A88875" originalValue="Guácharo">Guacharo</normalizedToken>
, 700m, 20-30.vii.1987, malaise/FIT, forest over coffee, S. &amp; J. Peck (FMNH).
</paragraph>
</subSubSection>
<subSubSection id="DED02D4B8415E6D53806EAFFBA6B0CF6" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="B5FAF1A55EB6654B0D1F3BFB60BCD36D" pageId="71" pageNumber="72">Diagnosis.</paragraph>
<paragraph id="AE2EDA3986E83015DB224D639BB2C316" pageId="71" pageNumber="72">
Length: 2.25-2.62 mm, width: 1.93-2.28 mm; body piceous; median portion of frontal stria weaker, occasionally absent across middle; supraorbital stria complete across vertex; anterior submarginal pronotal stria generally attached to lateral submarginal stria (angulate behind median pronotal gland opening); anterior emargination of pronotum barely or not outwardly angulate, with a complete marginal stria; elytra with a few fragments of inner subhumeral stria generally present, sutural stria from two-thirds to three-quarters complete; pygidium with fine, dense ground punctation and with coarser secondary punctures uniformly interspersed, apical sulcus complete, crenulate, shallower than in
<taxonomicName id="5971C0B15D40A9433E4733DF6EA24C96" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
. Male genitalia (Figs 18E, G): T8 lacking unique secondary sclerotization along edge of apical emargination; S9 slightly broader, abruptly expanded near apex, with midline sclerotization vague; tegmen evenly rounded, narrowing toward apex, narrowed but more or less straight to near apex in lateral aspect.
</paragraph>
</subSubSection>
<subSubSection id="60DB6A814D68AA64B6161DBC396420A0" pageId="72" pageNumber="73" type="remarks">
<paragraph id="38217EF00FD8E04903A8933D5669F0A6" pageId="72" pageNumber="73">
<pageBreakToken id="03CD50F1158AE3E0DD22B140AD48FB1B" pageId="72" pageNumber="73" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="C73119D9829D7D24AB5A9B3D3EBC704D" pageId="72" pageNumber="73">
See above for characters distinguishing
<taxonomicName id="2F0E81E25CF1BD1B46BAA191EF16FCCD" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
from
<taxonomicName id="8DE59382D3AFDBEFE84ED92314270673" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
. While their distributions overlap, and
<taxonomicName id="DF0D769AC4D10815C40E215A8B9B68EF" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
is very widespread, it is curious that, in Panama,
<taxonomicName id="E5699075E595938E0157D5685EC6BEF2" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
is only known from the type locality.
</paragraph>
<paragraph id="BAE5E59F32E32C380698A5E938E984EA" pageId="72" pageNumber="73">
<bibRefCitation id="6599DA8E8EAE0C2F4D3457D534EAA250" author="Wenzel, RL" journalOrPublisher="Fieldiana, Zoology" pageId="377" pageNumber="378" pagination="433 - 472" title="New and little known neotropical Histeridae (Coleoptera)." volume="22" year="1941">Wenzel and Dybas's (1941)</bibRefCitation>
<taxonomicName id="47A8B019AF464F50329E65C6C77D2690" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus miladae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="miladae">Operclipygus miladae</taxonomicName>
was not compared with other species in the original description, and at the time
<taxonomicName id="7ECE58D2C598DABD7B9AC098CAC6467A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
was still included in the genus
<taxonomicName id="88A784247F6D83867EC42C51F06AE756" class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">Phelister</taxonomicName>
. However, in
<bibRefCitation id="C0A5FC4E4A97615D2B3DF6681F18447D" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel's (1976)</bibRefCitation>
key, the characters presented do not allow their consistent separation, and the variation seen in
<taxonomicName id="8CBFB430FE0C7A5C6094025980847C90" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
fully encompasses the states found in
<taxonomicName id="4DCD21EA4E0334E84CDD699444D51340" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus miladae" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="miladae">Operclipygus miladae</taxonomicName>
. The proposed synonymy of
<taxonomicName id="68A0C355AFFA3CA1C6032BB76089B4A8" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister propygidialis" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="propygidialis">Pseudister propygidialis</taxonomicName>
here is admittedly more tentative, in the absence of a type specimen. However, Hinton's description and figure suggest a close match, and specimens identified as
<taxonomicName id="FCCBE94893B2C766D34BAC25C9CDC73E" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus propygidialis" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="propygidialis">Operclipygus propygidialis</taxonomicName>
in various collections are clearly
<taxonomicName id="1E1A26B516226771DC04F9E9FA4A9958" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus crenatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="crenatus">Operclipygus crenatus</taxonomicName>
. Hopefully the type specimen will eventually surface and this can be confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>
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