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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1039.64740" ID-GBIF-Dataset="8fe0041e-40a6-4fd0-a674-948f3f919703" ID-PMC="PMC8159918" ID-Pensoft-Pub="1313-2970-1039-1" ID-Pensoft-UUID="FA2622A854585C59A7E8182A4E66C6DF" ID-PubMed="34084064" ID-ZooBank="3487C3570FAC4907A5C0C0A0C801C6E3" ModsDocID="1313-2970-1039-1" checkinTime="1621541808324" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Bezdek, Jan" docDate="2021" docId="1681EBF9D68C50A3B656E12F2FD44BAA" docLanguage="en" docName="ZooKeys 1039: 1-108" docOrigin="ZooKeys 1039" docPubDate="2021-05-20" docSource="http://dx.doi.org/10.3897/zookeys.1039.64740" docTitle="Pyrrhalta tsoui Bezdek &amp; Lee 2019" docType="treatment" docVersion="4" id="FA2622A854585C59A7E8182A4E66C6DF" lastPageNumber="1" masterDocId="FA2622A854585C59A7E8182A4E66C6DF" masterDocTitle="Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei" masterLastPageNumber="108" masterPageNumber="1" pageNumber="1" updateTime="1668152740594" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the genera Xanthogaleruca Laboissiere, 1932 and Pyrrhalta Joannis, 1865 (Coleoptera, Chrysomelidae, Galerucinae) of Taiwan, with type designation of Galerucella lineatipes Takei</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bezdek, Jan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4358-7211</mods:nameIdentifier>
<mods:affiliation>Mendel University in Brno, Department of Zoology, Fisheries, Hydrobiology and Apiculture, Zemĕdĕlska 1, 613 00, Brno, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1039</mods:number>
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<mods:start>1</mods:start>
<mods:end>108</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1039.64740</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1039.64740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1039-1</mods:identifier>
<mods:identifier type="ZooBank">3487C3570FAC4907A5C0C0A0C801C6E3</mods:identifier>
<mods:identifier type="Pensoft-UUID">FA2622A854585C59A7E8182A4E66C6DF</mods:identifier>
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<treatment ID-GBIF-Taxon="182248294" LSID="urn:lsid:plazi:treatment:1681EBF9D68C50A3B656E12F2FD44BAA" httpUri="http://treatment.plazi.org/id/1681EBF9D68C50A3B656E12F2FD44BAA" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="1681EBF9-D68C-50A3-B656-E12F2FD44BAA" authority="Bezdek &amp; Lee, 2019" authorityName="Bezdek &amp; Lee" authorityYear="2019" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta tsoui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tsoui">
Pyrrhalta tsoui
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
&amp; Lee, 2019
</taxonomicName>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Habitus of Pyrrhalta maculata Gressitt &amp; Kimoto and P. tsoui Bezdek &amp; Lee A P. maculata, female, dorsal view B ditto, ventral view C ditto, lateral view D P. tsoui female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure23" httpUri="https://binary.pensoft.net/fig/545553" pageId="0" pageNumber="1">Figs 23D-F</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">, 25</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Field photographs of adults of Pyrrhalta species on host plants A P. tsoui Bezdek &amp; Lee B P. formosanensis sp. nov. C P. semifulva (Jacoby) D P. ishiharai Kimoto." figureDoi="10.3897/zookeys.1039.64740.figure26" httpUri="https://binary.pensoft.net/fig/545556" pageId="0" pageNumber="1">, 26A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bezdek &amp; Lee" authorityYear="2019" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrrhalta tsoui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tsoui">Pyrrhalta tsoui</taxonomicName>
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
&amp; Lee, 2019: 531.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material</paragraph>
<paragraph pageId="0" pageNumber="1">
(specimens examined by
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4664.4.4" author="Bezdek, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="518 - 534" refId="B7" refString="Bezdek, J, Lee, C-F, 2019. Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra. Zootaxa 4664: 518 - 534, DOI: https://doi.org/10.11646/zootaxa.4664.4.4" title="Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra." url="https://doi.org/10.11646/zootaxa.4664.4.4" volume="4664" year="2019">
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
and Lee (2019)
</bibRefCitation>
excluded).
<materialsCitation collectingDate="1976-06-06" collectorName="H. Makihara" country="Taiwan" location="Nantou" specimenCount="2" specimenCount-female="2" stateProvince="Nantou">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
.
<collectingRegion country="Taiwan" name="Nantou">Nantou</collectingRegion>
:
<specimenCount type="female">2♀</specimenCount>
(KMNH), Lushan Wenchuan (廬山溫泉),
<collectingDate value="1976-06-06">6.VI.1976</collectingDate>
, leg.
<collectorName>H. Makihara</collectorName>
(one identified as
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
by Kimoto, 1983)
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
(TARI), Tsuifeng (翠峰),
<date value="1984-09-12" valueMax="1984-09-14" valueMin="1984-09-12">12-14.IX.1984</date>
, leg. K. S. Lin &amp; S. C. Lin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length 4.6-5.3 mm, width 2.3-2.8 mm. Body color (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Habitus of Pyrrhalta maculata Gressitt &amp; Kimoto and P. tsoui Bezdek &amp; Lee A P. maculata, female, dorsal view B ditto, ventral view C ditto, lateral view D P. tsoui female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure23" httpUri="https://binary.pensoft.net/fig/545553" pageId="0" pageNumber="1">23D-F</figureCitation>
) reddish brown; vertex with one black spot at center; pronotum with three large black spots, one poorly defined elongate spot at center, from basal 1/3 to middle, one pair laterally; scutellum black; five pairs of large black spots on elytra, one pair near base at middle, two pairs in line with middle, one pair at apical 2/5 laterally, one pair at apical 1/5 near suture; metathoracic ventrites darker. Eyes small, interocular space 2.37-2.42
<normalizedToken originalValue="×">x</normalizedToken>
diameter of eye. Antennae filiform in males (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25A</figureCitation>
), antennomere I swollen, IV with a large tubercle on outer sides at middle, length ratios of antennomeres I-XI 1.0: 0.5: 0.8: 0.9: 0.7: 0.7: 0.6: 0.6: 0.6: 0.6: 1.0, length to width ratios of antennomeres I-XI 2.1: 1.9: 2.7: 2.6: 2.1: 1.9: 1.6: 1.5: 1.5: 1.4: 2.4; filiform in females (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25B</figureCitation>
), antennomere I and IV not modified, length ratios of antennomeres I-XI 1.0: 0.6: 0.9: 0.8: 0.7: 0.6: 0.6: 0.6: 0.6: 0.6: 1.0, length to width ratios of antennomeres I-XI 2.4: 2.1: 3.4: 2.7: 2.2: 1.9: 1.7: 1.5: 1.6: 1.6: 2.5. Pronotum and elytra convex. Pronotum 1.9-2.0
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, disc with reticulate microsculpture; dense, extremely coarse punctures and extremely short pubescence; with median longitudinal and lateral depressions; lateral margins moderately rounded, apical and basal margins straight; anterior and posterior setiferous punctures erect. Elytra elongate and broad, parallel-sided, 1.4-1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; disc with reticulate microsculpture, and with dense extremely coarse punctures and short pubescence. Apical spur of tibia of middle leg small (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25E</figureCitation>
), tarsomere I not modified in males. Aedeagus (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25C, D</figureCitation>
) extremely slender in dorsal view, 8.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, sides asymmetric, curved near apex, apex truncate; strongly curved at apical 1/3 and near base in lateral view, apex truncate; ostium not covered by membrane, ventrally located, along lateral margin; two endophallic sclerites elongate, apex of primary endophallic sclerite with several teeth, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus, secondary sclerite much shorter, 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as primary endophallic sclerite, apex acute, with one tooth near apex. Only apices of gonocoxae (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25J</figureCitation>
) sclerotized and transverse, with two long setae at apex of each gonocoxa. Ventrite VIII (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25F</figureCitation>
) transverse; disc with several long setae and dense short setae along apical margin; spiculum long. Receptacle of spermatheca (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25G</figureCitation>
) slightly swollen; pump short and strongly curved; sclerotized proximal spermathecal duct wide and short. Apical margin of abdominal ventrite V truncate, with deeply rounded depression at middle in males (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25I</figureCitation>
); slightly concave in females (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adults of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
<normalizedToken originalValue="Bezděk">Bezdek</normalizedToken>
&amp; Lee and
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
Gressitt &amp; Kimoto may be separated from others within the species group by the five pairs of large black spots on the elytra (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Habitus of Pyrrhalta maculata Gressitt &amp; Kimoto and P. tsoui Bezdek &amp; Lee A P. maculata, female, dorsal view B ditto, ventral view C ditto, lateral view D P. tsoui female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1039.64740.figure23" httpUri="https://binary.pensoft.net/fig/545553" pageId="0" pageNumber="1">23</figureCitation>
), the strongly curved aedeagus in lateral view (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24C</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25C</figureCitation>
), and gonocoxa with only two setae (Figs
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24H</figureCitation>
,
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25J</figureCitation>
). adults of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
differ from those of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
by the normal antennomere III and antennomere IV with a large tubercle in males (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25A</figureCitation>
) (antennomere III apically broadened in
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24A</figureCitation>
)), and slender antennomere III, &lt;4.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in females (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25B</figureCitation>
) (extremely slender antennomere III,&gt; 4.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; in those of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24B</figureCitation>
)). In males of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. tsoui" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="tsoui">
<emphasis italics="true" pageId="0" pageNumber="1">P. tsoui</emphasis>
</taxonomicName>
, the apex of the primary endophallic sclerite has several teeth and one additional tooth on the secondary sclerite (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Diagnostic characters of Pyrrhalta tsoui Bezdek &amp; Lee A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H abdominal ventrite V, female I abdominal ventrite V, male J gonocoxae." figureDoi="10.3897/zookeys.1039.64740.figure25" httpUri="https://binary.pensoft.net/fig/545555" pageId="0" pageNumber="1">25C, D</figureCitation>
). In
<taxonomicName class="Dicotyledoneae" family="Rosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="P. maculata" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. maculata</emphasis>
</taxonomicName>
the primary endophallic sclerite is acute apically, and the secondary sclerite lacks additional teeth (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Diagnostic characters of Pyrrhalta maculata Gressitt &amp; Kimoto A antenna, male B antenna, female C aedeagus, dorsal view D ditto, lateral view E apex of tibia of middle leg, male F abdominal ventrite VIII G spermatheca H gonocoxae I abdominal ventrite V, female J abdominal ventrite V, male K tarsi of middle leg, male." figureDoi="10.3897/zookeys.1039.64740.figure24" httpUri="https://binary.pensoft.net/fig/545554" pageId="0" pageNumber="1">24C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="food plant">
<paragraph pageId="0" pageNumber="1">Food plant.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adults feed on flowers of
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Meliosma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Meliosma rhoifolia" order="Rosales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="rhoifolia">
<emphasis italics="true" pageId="0" pageNumber="1">Meliosma rhoifolia</emphasis>
</taxonomicName>
Maxim. (
<taxonomicName class="Dicotyledoneae" family="Sabiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Sabiaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">This species is widespread in lowlands of Taiwan.</paragraph>
</subSubSection>
</treatment>
</document>