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<document id="958CCA4649DB64864F38B2E4B1A16305" ID-DOI="10.5281/zenodo.171981" ID-GBIF-Dataset="8ee587b3-3852-49d2-9297-995428a3531a" ID-ISSN="1175­5326" ID-Zenodo-Dep="171981" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761366646" checkinUser="plazi" docAuthor="Rafael, José Albertino &amp; Cumming, Jeffrey M." docDate="2006" docId="163B5B4D346CFFEFFEF7CA3DFB5747A3" docLanguage="en" docName="zt01136p064.pdf" docOrigin="Zootaxa 1136" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrostomus tarsalis Rafael &amp; Cumming, 2006, n. sp." docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="EA0223353467FFE0FFFFC949FFC1465D" masterDocTitle="New species of Macrostomus Wiedemann (Diptera, Empididae) from Colombia, Ecuador and Peru" masterLastPageNumber="64" masterPageNumber="49" pageNumber="60" updateTime="1698215234110" updateUser="plazi">
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<mods:title id="2DD93B3A259D49D7AAD5512485CF1319">New species of Macrostomus Wiedemann (Diptera, Empididae) from Colombia, Ecuador and Peru</mods:title>
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<mods:namePart id="46096B4B2B56055748E1E7C9C07C7E22">Rafael, José Albertino</mods:namePart>
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<mods:namePart id="849A2F14BB18AFA4BF0904FC2B50D1C3">Cumming, Jeffrey M.</mods:namePart>
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<mods:date id="523F0E57825B6DEE5A9F94A24FF09716">2006</mods:date>
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<treatment id="163B5B4D346CFFEFFEF7CA3DFB5747A3" ID-DOI="http://doi.org/10.5281/zenodo.6493802" ID-GBIF-Taxon="119344964" ID-Zenodo-Dep="6493802" LSID="urn:lsid:plazi:treatment:163B5B4D346CFFEFFEF7CA3DFB5747A3" httpUri="http://treatment.plazi.org/id/163B5B4D346CFFEFFEF7CA3DFB5747A3" lastPageId="15" lastPageNumber="64" pageId="11" pageNumber="60">
<subSubSection id="D688B9D0346CFFEBFEF7CA3DFE1345EC" pageId="11" pageNumber="60" type="nomenclature">
<paragraph id="9E2DEA5B346CFFEBFEF7CA3DFD9C45D3" blockId="11.[264,605,884,945]" box="[264,605,884,910]" pageId="11" pageNumber="60">
<heading id="C5655D37346CFFEBFEF7CA3DFD9C45D3" bold="true" box="[264,605,884,910]" fontSize="11" level="1" pageId="11" pageNumber="60" reason="1">
<emphasis id="ACE63649346CFFEBFEF7CA3DFD9C45D3" bold="true" box="[264,605,884,910]" pageId="11" pageNumber="60">
<taxonomicName id="599291D8346CFFEBFEF7CA3DFDCB45D3" ID-CoL="3XBBH" box="[264,522,884,910]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="tarsalis" status="sp. nov.">
<emphasis id="ACE63649346CFFEBFEF7CA3DFDCB45D3" bold="true" box="[264,522,884,910]" italics="true" pageId="11" pageNumber="60">Macrostomus tarsalis</emphasis>
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,
<taxonomicNameLabel id="B7D58B32346CFFEBFDE5CA3DFD9C45D3" box="[538,605,884,910]" pageId="11" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="9E2DEA5B346CFFEBFEF7CADEFE1345EC" blockId="11.[264,605,884,945]" box="[264,466,919,945]" pageId="11" pageNumber="60">
(
<figureCitation id="06A9F6DE346CFFEBFEEECADEFE5E45EC" box="[273,415,919,945]" captionStart-0="FIGURES 29 ­ 33" captionStart-1="FIGURES 34 ­ 39" captionStartId-0="12.[264,382,1762,1786]" captionStartId-1="13.[264,382,1324,1348]" captionTargetBox-0="[307,1280,473,1738]" captionTargetBox-1="[290,1297,327,1298]" captionTargetId-0="figure@12.[305,1282,446,1738]" captionTargetId-1="figure@13.[289,1297,326,1298]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." captionText-1="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri-0="https://zenodo.org/record/171987/files/figure.png" httpUri-1="https://zenodo.org/record/171988/files/figure.png" pageId="11" pageNumber="60">Figs. 2939</figureCitation>
,
<figureCitation id="06A9F6DE346CFFEBFE54CADEFE0B45EC" box="[427,458,919,945]" captionStart="FIGURE 40" captionStartId="14.[264,368,1741,1765]" captionTargetBox="[264,1323,846,1717]" captionTargetId="figure@14.[264,1323,846,1717]" captionTargetPageId="14" captionText="FIGURE 40. Distributions of new species of Macrostomus. Symbols represent the known collection localities as follows: square = M. alpinus; diamond = M. ciliaticosta; circle = M. lineatus; star = M. occidentalis; triangle = M. penai; cross = M. tarsalis." httpUri="https://zenodo.org/record/171989/files/figure.png" pageId="11" pageNumber="60">40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D688B9D0346CFFEFFEF7CAAEFB5747A3" lastPageId="15" lastPageNumber="64" pageId="11" pageNumber="60" type="diagnosis">
<paragraph id="9E2DEA5B346CFFEBFEF7CAAEFDCD42FB" blockId="11.[264,1324,999,1881]" pageId="11" pageNumber="60">Diagnosis. Prosternum yellow; anterolateral part of scutum, including postpronotal lobe without pruinosity; female hind leg with ventral pinnate scales at distal quarter of femur and some dorsal near middle of femur, dorsal along dorsal face and ventral at basal half of hind tibia; male hind tarsomere 3 widened in dorsal view; cell dm rather acute; M1, M2 and A1 rather distinct.</paragraph>
<paragraph id="9E2DEA5B346CFFECFEC7CDF3FE6C47DB" blockId="11.[264,1324,999,1881]" lastBlockId="12.[264,1323,284,390]" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
Description. Male. Head lost. Thorax black with prosternum yellow; gray­blue pruinose with anterolateral part of scutum, including postpronotal lobe without pruinosity. 3 dorsocentral setae, the mid one weaker; 3 notopleural setae; 45 antepronotal setae; 2 proepisternal setae being one strong; 78 katatergital setae. Legs, predominantly yellow with apex of mid femur and apex of mid tibia, distal half of hind femur, entire hind tibia and all tarsi, black; tarsomere 2 inconspicuously yellow medially. Slightly stronger setae: mid tibia with 1 anterodorsal near base, 23 ventral very short along distal three­quarter; hind femur with 2 anterior at basal three­quarter and at distal three­quarter, no anteroventral setae. Slender setae: hind tibia with 4 dorsal; hind tarsomere 1 with 1 dorsal.
<collectingCountry id="E685AACB346CFFEBFEF7CF6BFE824061" box="[264,323,1570,1596]" name="India" pageId="11" pageNumber="60">Hind</collectingCountry>
tarsomere 3 modified, widened in dorsal view. Wing (
<figureCitation id="06A9F6DE346CFFEBFC28CF6BFBEE4061" box="[983,1071,1570,1596]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="11" pageNumber="60">Fig. 29</figureCitation>
) brown­infuscated at costal cell, pterostigma and apex of cell r2+3; cell dm rather acute; M1, M2 and A1 rather distinct. Halter light yellow to white. Terminalia (
<figureCitation id="06A9F6DE346CFFEBFC9CCF3EFC3640CC" box="[867,1015,1655,1681]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="11" pageNumber="60">Figs. 3033</figureCitation>
). Tergite 8 (
<figureCitation id="06A9F6DE346CFFEBFB74CF3EFB2540CC" box="[1163,1252,1655,1681]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="11" pageNumber="60">Fig. 30</figureCitation>
) with large broad projection and wide basal median cleft; sternite 8 (
<figureCitation id="06A9F6DE346CFFEBFC16CFD6FBFF40E4" box="[1001,1086,1695,1721]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="11" pageNumber="60">Fig. 31</figureCitation>
) with posterolateral projection. Epandrium (
<figureCitation id="06A9F6DE346CFFEBFDD6CF8EFD4540BC" box="[553,644,1735,1761]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="11" pageNumber="60">Fig. 32</figureCitation>
) with anterodorsal and posterodorsal lobes of subequal height. Anterior cercus elongate; posterior cercus complex, slightly bifid distally with long slender setae at margin; with a wide ventral protuberance with elongate digitiform yellow setae distally, internally with elongate protuberance with short distal yellow setae and a smaller and thinner simple protuberance near base of the latter. Hypandrium (
<figureCitation id="06A9F6DE346BFFECFB6DC855FB27476B" box="[1170,1254,284,310]" captionStart="FIGURES 29 ­ 33" captionStartId="12.[264,382,1762,1786]" captionTargetBox="[307,1280,473,1738]" captionTargetId="figure@12.[305,1282,446,1738]" captionTargetPageId="12" captionText="FIGURES 29 ­ 33. Macrostomus tarsalis, n. sp., holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view." httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="12" pageNumber="61">Fig. 33</figureCitation>
) with large sinus at apex and a unique protuberance inside sinus, many long setae on distal posterior half.
</paragraph>
<caption id="CAEDBAD3346BFFECFEF7CFABFC8C4162" httpUri="https://zenodo.org/record/171987/files/figure.png" pageId="12" pageNumber="61" targetBox="[307,1280,473,1738]" targetPageId="12">
<paragraph id="9E2DEA5B346BFFECFEF7CFABFC8C4162" blockId="12.[264,1322,1762,1855]" pageId="12" pageNumber="61">
<emphasis id="ACE63649346BFFECFEF7CFABFE0740A7" bold="true" box="[264,454,1762,1786]" pageId="12" pageNumber="61">FIGURES 29­33</emphasis>
.
<taxonomicName id="599291D8346BFFECFE29CFABFD7E40A4" box="[470,703,1762,1785]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="tarsalis" status="sp. nov.">
<emphasis id="ACE63649346BFFECFE29CFABFD7E40A4" box="[470,703,1762,1785]" italics="true" pageId="12" pageNumber="61">Macrostomus tarsalis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B7D58B32346BFFECFD30CFABFCCD40A4" box="[719,780,1762,1785]" pageId="12" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, holotype ♂. 29, wing; 30, tergite 8, dorsal view, right setae not represented; 31, sternite 8, ventral view, left setae not represented; 32, terminalia, lateral view; 33, hypandrium and phallus, lateral view.
</paragraph>
</caption>
<caption id="CAEDBAD3346AFFEDFEF7CC65FD3143F1" httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="13" pageNumber="62" targetBox="[290,1297,327,1298]" targetPageId="13">
<paragraph id="9E2DEA5B346AFFEDFEF7CC65FD3143F1" blockId="13.[264,1322,1322,1452]" pageId="13" pageNumber="62">
<emphasis id="ACE63649346AFFEDFEF7CC65FE044319" bold="true" box="[264,453,1324,1348]" pageId="13" pageNumber="62">FIGURES 34­39</emphasis>
.
<taxonomicName id="599291D8346AFFEDFE23CC64FD054319" box="[476,708,1325,1348]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="tarsalis" status="sp. nov.">
<emphasis id="ACE63649346AFFEDFE23CC64FD054319" box="[476,708,1325,1348]" italics="true" pageId="13" pageNumber="62">Macrostomus tarsalis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B7D58B32346AFFEDFD2CCC65FCCE431E" box="[723,783,1324,1347]" pageId="13" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork.
</paragraph>
</caption>
<paragraph id="9E2DEA5B346AFFEDFEC7CCB4FAEB410A" blockId="13.[264,1323,1533,1886]" pageId="13" pageNumber="62">
Female. Frons wider than anterior ocellus width; face approximately anterior ocellus width, without pruinosity; 2 pairs of ocellar setae; postgena setae rather long. Prosternum yellow and propleuron partially yellow. 34 dorsocentral setae; 34 antepronotal setae (dorsocentral and antepronotal setae variable in the same specimen). Slightly stronger setae: mid tibia with 1 anterodorsal near base, 1 posterodorsal near middle, 2 posteroventral at base and apex of median third and 1 anteroventral at apex of median third. Pinnate scales on hindleg (
<figureCitation id="06A9F6DE346AFFEDFD75CFA4FD22415A" box="[650,739,1773,1799]" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="13" pageNumber="62">Fig. 34</figureCitation>
): ventral at distal quarter and some dorsal thinner near middle of femur; dorsal along dorsal face and ventral at basal half of tibia. Wing (
<figureCitation id="06A9F6DE346AFFEDFB04CE5CFEE7410A" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="13" pageNumber="62">Fig. 35</figureCitation>
) wider than in male, brown­ infuscated at costal and r1 cells and apex of r2+3 cell.
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEF7C855FE6647F3" blockId="14.[264,1323,284,790]" pageId="14" pageNumber="63">
Terminalia (
<figureCitation id="06A9F6DE3469FFEEFE67C855FDEA476B" box="[408,555,284,310]" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="14" pageNumber="63">Figs. 3639</figureCitation>
). Tergite 8 and sternite 8 as depicted in
<figureCitation id="06A9F6DE3469FFEEFBF1C855FBB3476B" box="[1038,1138,284,310]" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="14" pageNumber="63">Figs. 36</figureCitation>
. Tergite 8 (
<figureCitation id="06A9F6DE3469FFEEFB04C855FEE74703" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="14" pageNumber="63">Fig. 38</figureCitation>
) with large distal cleft. sternite 8 (
<figureCitation id="06A9F6DE3469FFEEFD36C80DFCE04703" box="[713,801,324,350]" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="14" pageNumber="63">Fig. 37</figureCitation>
) rather membranous ventrally. Genital fork (
<figureCitation id="06A9F6DE3469FFEEFEEEC825FEB147DB" box="[273,368,364,390]" captionStart="FIGURES 34 ­ 39" captionStartId="13.[264,382,1324,1348]" captionTargetBox="[290,1297,327,1298]" captionTargetId="figure@13.[289,1297,326,1298]" captionTargetPageId="13" captionText="FIGURES 34 ­ 39. Macrostomus tarsalis, n. sp., paratype Ψ. 34 right anterior leg, posterior view, larger and pinnate setae represented; 35, wing; 36, tergite 8 and sternite 8, lateral view, setae not represented; 37, sternite 8, ventral view, left setae not represented; 38, female tergite 8, dorsal view, setae not represented; 39, female genital fork." httpUri="https://zenodo.org/record/171988/files/figure.png" pageId="14" pageNumber="63">Fig. 39</figureCitation>
) elongate with short lateral arms not projected laterally. Genital chamber membranous.
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEC7C8F5FCA0478B" blockId="14.[264,1323,284,790]" box="[312,865,444,470]" pageId="14" pageNumber="63">
Size. Body length not measured; wing
<quantity id="596A47BE3469FFEEFD00C8F5FC9F478B" box="[767,862,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="63" unit="mm" value="4.8">4.8 mm</quantity>
.
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEC7C8ADFC8B47A3" blockId="14.[264,1323,284,790]" box="[312,842,484,510]" pageId="14" pageNumber="63">
Geographical distribution.
<collectingCountry id="E685AACB3469FFEEFD8CC8ADFD1447A3" box="[627,725,484,510]" name="Ecuador" pageId="14" pageNumber="63">Ecuador</collectingCountry>
(
<figureCitation id="06A9F6DE3469FFEEFD1AC8ADFCFC47A3" box="[741,829,484,510]" captionStart="FIGURE 40" captionStartId="14.[264,368,1741,1765]" captionTargetBox="[264,1323,846,1717]" captionTargetId="figure@14.[264,1323,846,1717]" captionTargetPageId="14" captionText="FIGURE 40. Distributions of new species of Macrostomus. Symbols represent the known collection localities as follows: square = M. alpinus; diamond = M. ciliaticosta; circle = M. lineatus; star = M. occidentalis; triangle = M. penai; cross = M. tarsalis." httpUri="https://zenodo.org/record/171989/files/figure.png" pageId="14" pageNumber="63">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEC7CB45FD6E442B" blockId="14.[264,1323,284,790]" pageId="14" pageNumber="63">
Material.
<typeStatus id="412954F93469FFEEFE53CB45FDD8447B" box="[428,537,524,550]" pageId="14" pageNumber="63" type="holotype">Holotype</typeStatus>
ɗ:
<collectingCountry id="E685AACB3469FFEEFDB6CB45FD1B447B" box="[585,730,524,550]" name="Ecuador" pageId="14" pageNumber="63">ECUADOR</collectingCountry>
, Napo­Pastaza,
<quantity id="596A47BE3469FFEEFC65CB45FC33447B" box="[922,1010,524,550]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" metricValueMax="12.874752" metricValueMin="3.218688" pageId="14" pageNumber="63" unit="mi" value="5.0" valueMax="8.0" valueMin="2.0">28 mi</quantity>
[les] N[orth] of Puyo,
<quantity id="596A47BE3469FFEEFB00CB45FEE34413" metricMagnitude="2" metricUnit="m" metricValue="9.53" pageId="14" pageNumber="63" unit="m" value="953.0">953 m</quantity>
, II.9.1955 (sic), E.I. Schlinger &amp; E.S. Ross collectors (CAS).
<typeStatus id="412954F93469FFEEFBF9CB7DFBB14413" box="[1030,1136,564,590]" pageId="14" pageNumber="63" type="paratype">Paratype</typeStatus>
. Same data and same pin as the
<typeStatus id="412954F93469FFEEFE3CCB15FDEB442B" box="[451,554,604,630]" pageId="14" pageNumber="63" type="holotype">holotype</typeStatus>
(1Ψ CAS).
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEC7CBCDFC40449B" blockId="14.[264,1323,284,790]" pageId="14" pageNumber="63">
<typeStatus id="412954F93469FFEEFEC7CBCDFE6444C3" box="[312,421,644,670]" pageId="14" pageNumber="63" type="holotype">Holotype</typeStatus>
condition. Head lost. Right wing mounted in microslide; terminalia in glycerine together with female terminalia of
<typeStatus id="412954F93469FFEEFCEBCBE5FCBC449B" box="[788,893,684,710]" pageId="14" pageNumber="63" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="9E2DEA5B3469FFEEFEC7CB9DFE58454B" blockId="14.[264,1323,284,790]" pageId="14" pageNumber="63">
Etymology. From Greek
<emphasis id="ACE636493469FFEEFD89CB9DFD7C44B0" box="[630,701,724,749]" italics="true" pageId="14" pageNumber="63">tarsos</emphasis>
= tarsus, in reference to the shape of the hind tarsomere 3.
</paragraph>
<caption id="CAEDBAD33469FFEEFEF7CF84FC674177" httpUri="https://zenodo.org/record/171989/files/figure.png" pageId="14" pageNumber="63" targetBox="[264,1323,846,1717]" targetPageId="14">
<paragraph id="9E2DEA5B3469FFEEFEF7CF84FC674177" blockId="14.[264,1323,1741,1834]" pageId="14" pageNumber="63">
<emphasis id="ACE636493469FFEEFEF7CF84FE5840B8" bold="true" box="[264,409,1741,1765]" pageId="14" pageNumber="63">FIGURE 40</emphasis>
. Distributions of new species of
<taxonomicName id="599291D83469FFEEFCD1CF87FC7F40B8" box="[814,958,1742,1765]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="ACE636493469FFEEFCD1CF87FC7F40B8" box="[814,958,1742,1765]" italics="true" pageId="14" pageNumber="63">Macrostomus</emphasis>
</taxonomicName>
. Symbols represent the known collection localities as follows: square =
<taxonomicName id="599291D83469FFEEFD47CFB8FCE94155" box="[696,808,1777,1800]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="ACE636493469FFEEFD47CFB8FCE94155" box="[696,808,1777,1800]" italics="true" pageId="14" pageNumber="63">M. alpinus</emphasis>
</taxonomicName>
; diamond =
<taxonomicName id="599291D83469FFEEFC50CFB8FB894155" box="[943,1096,1777,1800]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="ciliaticosta">
<emphasis id="ACE636493469FFEEFC50CFB8FB894155" box="[943,1096,1777,1800]" italics="true" pageId="14" pageNumber="63">M. ciliaticosta</emphasis>
</taxonomicName>
; circle =
<taxonomicName id="599291D83469FFEEFB53CFB8FAE24155" box="[1196,1315,1777,1800]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ACE636493469FFEEFB53CFB8FAE24155" box="[1196,1315,1777,1800]" italics="true" pageId="14" pageNumber="63">M. lineatus</emphasis>
</taxonomicName>
; star =
<taxonomicName id="599291D83469FFEEFEB4CE5AFE2E4177" box="[331,495,1811,1834]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="ACE636493469FFEEFEB4CE5AFE2E4177" box="[331,495,1811,1834]" italics="true" pageId="14" pageNumber="63">M. occidentalis</emphasis>
</taxonomicName>
; triangle =
<taxonomicName id="599291D83469FFEEFD94CE5AFD0B4177" box="[619,714,1811,1834]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="ACE636493469FFEEFD94CE5AFD0B4177" box="[619,714,1811,1834]" italics="true" pageId="14" pageNumber="63">M. penai</emphasis>
</taxonomicName>
; cross =
<taxonomicName id="599291D83469FFEEFCD4CE5AFC5E4177" box="[811,927,1811,1834]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="ACE636493469FFEEFCD4CE5AFC5E4177" box="[811,927,1811,1834]" italics="true" pageId="14" pageNumber="63">M. tarsalis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="9E2DEA5B3468FFEFFEC7C855FB5747A3" blockId="15.[264,1323,284,510]" pageId="15" pageNumber="64">
Discussion. As mentioned previously,
<taxonomicName id="599291D83468FFEFFCEBC855FC584768" box="[788,921,284,310]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="ACE636493468FFEFFCEBC855FCED4768" box="[788,812,284,309]" italics="true" pageId="15" pageNumber="64">M</emphasis>
.
<emphasis id="ACE636493468FFEFFCBEC855FC584768" box="[833,921,284,309]" italics="true" pageId="15" pageNumber="64">tarsalis</emphasis>
</taxonomicName>
apparently belongs to the same group as
<taxonomicName id="599291D83468FFEFFE7CC80DFDC94700" box="[387,520,324,349]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="ACE636493468FFEFFE7CC80DFDC94700" box="[387,520,324,349]" italics="true" pageId="15" pageNumber="64">M. alpinus</emphasis>
</taxonomicName>
,
<taxonomicName id="599291D83468FFEFFDE1C80DFD0E4700" box="[542,719,324,349]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="ciliaticosta">
<emphasis id="ACE636493468FFEFFDE1C80DFD0E4700" box="[542,719,324,349]" italics="true" pageId="15" pageNumber="64">M. ciliaticosta</emphasis>
</taxonomicName>
and
<taxonomicName id="599291D83468FFEFFCE7C80DFC624700" box="[792,931,324,349]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="ACE636493468FFEFFCE7C80DFC624700" box="[792,931,324,349]" italics="true" pageId="15" pageNumber="64">M. lineatus</emphasis>
</taxonomicName>
, on the basis of these species possessing a rather membranous area medially on the hypandrium.
<taxonomicName id="599291D83468FFEFFBD2C825FB6F47D8" box="[1069,1198,364,390]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="ACE636493468FFEFFBD2C825FB8447D8" box="[1069,1093,364,389]" italics="true" pageId="15" pageNumber="64">M</emphasis>
.
<emphasis id="ACE636493468FFEFFBA9C825FB6F47D8" box="[1110,1198,364,389]" italics="true" pageId="15" pageNumber="64">tarsalis</emphasis>
</taxonomicName>
is distinct from the remaining species in having tergite 8 with a large broad distal projection and a wide basal median cleft, sternite 8 with a posterolateral projection, and the hypandrium with a large sinus at the apex and many long setae on the distal posterior half.
</paragraph>
</subSubSection>
</treatment>
</document>