treatments-xml/data/16/04/87/160487E5FA6FFFB8FE95F926FB35F91E.xml
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<document id="6E3113C5873CD77149823E641762B46B" ID-DOI="10.5281/zenodo.157462" ID-GBIF-Dataset="a9ccfc65-1459-414b-8a99-c99649f83d51" ID-ISSN="1175­5326" ID-Zenodo-Dep="157462" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461019330970" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2004" docId="160487E5FA6FFFB8FE95F926FB35F91E" docLanguage="en" docName="zt00538.pdf" docOrigin="Zootaxa 538" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pneuminion natalensis Perkins, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="EA3DFF9DFA66FFB1FF9DF83AFFBAFFCF" masterDocTitle="A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="10" updateTime="1698569824382" updateUser="plazi">
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<mods:title id="3B352AFF6AA5A5A08F760D5430E658ED">A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="B127A4FBDABD38DAAAB7091750857C33">Perkins, Philip D.</mods:namePart>
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<mods:part id="3B61FDC0AA6649EE9F635A694BC8BD24">
<mods:date id="D2AE9D3AE6BF087D03B7E96C93F29AD9">2004</mods:date>
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<mods:identifier id="169C92712D58B4108DCAC39A8B11DEF4" type="DOI">10.5281/zenodo.157462</mods:identifier>
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<treatment id="160487E5FA6FFFB8FE95F926FB35F91E" ID-DOI="http://doi.org/10.5281/zenodo.6271127" ID-GBIF-Taxon="119602182" ID-Zenodo-Dep="6271127" LSID="urn:lsid:plazi:treatment:160487E5FA6FFFB8FE95F926FB35F91E" httpUri="http://treatment.plazi.org/id/160487E5FA6FFFB8FE95F926FB35F91E" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="D6B76578FA6FFFB8FE95F926FE5BFE96" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="9E1236F3FA6FFFB8FE95F926FD0CFEFA" blockId="9.[264,694,283,345]" box="[264,694,283,310]" pageId="9" pageNumber="10">
<heading id="C55A819FFA6FFFB8FE95F926FD0CFEFA" bold="true" box="[264,694,283,310]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="ACD9EAE1FA6FFFB8FE95F926FD0CFEFA" bold="true" box="[264,694,283,310]" pageId="9" pageNumber="10">
<taxonomicName id="59AD4D70FA6FFFB8FE95F926FDA4FEF9" ID-CoL="4KKVC" box="[264,542,284,310]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="natalensis" status="sp. nov.">
<emphasis id="ACD9EAE1FA6FFFB8FE95F926FDA4FEF9" bold="true" box="[264,542,284,310]" italics="true" pageId="9" pageNumber="10">Pneuminion natalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7EA579AFA6FFFB8FDBBF921FD0CFEFA" box="[550,694,283,309]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E1236F3FA6FFFB8FE95F905FE5BFE96" blockId="9.[264,694,283,345]" box="[264,481,319,345]" pageId="9" pageNumber="10">
(
<figureCitation id="06962A76FA6FFFB8FE8DF905FE3BFE96" box="[272,385,319,345]" captionStart="FIGURE 6" captionStartId="19.[264,368,1577,1601]" captionTargetBox="[264,1322,310,1568]" captionTargetId="figure@19.[264,1323,309,1572]" captionTargetPageId="19" captionText="FIGURE 6. Pneuminion natalensis holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169052/files/figure.png" pageId="9" pageNumber="10">Figures 6</figureCitation>
,
<figureCitation id="06962A76FA6FFFB8FE13F905FE14FE96" box="[398,430,319,345]" captionStart="FIGURE 16" captionStartId="29.[264,368,1700,1724]" captionTargetBox="[338,1203,316,1657]" captionTargetId="figure@29.[337,1249,315,1676]" captionTargetPageId="29" captionText="FIGURE 16. Pneuminion natalensis. Aedeagus of holotype." httpUri="https://zenodo.org/record/157476/files/figure.png" pageId="9" pageNumber="10">16</figureCitation>
,
<figureCitation id="06962A76FA6FFFB8FE27F905FE63FE96" box="[442,473,319,345]" captionStart="FIGURES 29 30" captionStartId="37.[264,382,1650,1674]" captionTargetBox="[277,1310,326,1614]" captionTargetId="figure@37.[277,1311,326,1626]" captionTargetPageId="37" captionText="FIGURES 29 30. Pneuminion distribution maps. — 29. P. punctatum. — 30. P. natalensis." httpUri="https://zenodo.org/record/169061/files/figure.png" pageId="9" pageNumber="10">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6FFFB8FE95F9B4FED1FDEE" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="9E1236F3FA6FFFB8FE95F9B4FED1FDEE" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">
<emphasis id="ACD9EAE1FA6FFFB8FE95F9B4FE7AFE67" bold="true" box="[264,448,398,424]" pageId="9" pageNumber="10">
<typeStatus id="41168851FA6FFFB8FE95F9B4FEF9FE67" box="[264,323,398,424]" pageId="9" pageNumber="10">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="41168851FA6FFFB8FE48F9B5FDF8FE66" box="[469,578,399,425]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="E6BA7663FA6FFFB8FD2CF9B5FCF4FE66" box="[689,846,399,425]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
:
<collectingCountry id="E6BA7663FA6FFFB8FCC7F9B5FC26FE66" box="[858,924,399,425]" name="South Africa" pageId="9" pageNumber="10">Natal</collectingCountry>
,
<quantity id="59559B16FA6FFFB8FC37F9B5FC57FE66" box="[938,1005,399,425]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="10" unit="km" value="75.0">75km</quantity>
WSW Estcourt, Cathedral Peaks For. Sta.,
<date id="EA131033FA6FFFB8FE56F98DFDE8FE1E" box="[459,594,439,465]" pageId="9" pageNumber="10" value="1979-12-07" valueMax="1979-12-31" valueMin="1979-12-07">731.xii.79</date>
, S. &amp; J. Peck, Meteorology Sta., LittleBerg,
<quantity id="59559B16FA6FFFB8FBF2F98DFB79FE1E" box="[1135,1219,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="9" pageNumber="10" unit="m" value="1860.0">1860m</quantity>
, malaise trough,
<date id="EA131033FA6FFFB8FEF4F9E5FE45FE36" box="[361,511,479,505]" pageId="9" pageNumber="10" value="1979-12-21" valueMax="1979-12-31" valueMin="1979-12-21">2131.xii.79</date>
; deposited in the
<collectionCode id="F8BCAE36FA6FFFB8FD7BF9E5FC81FE36" box="[742,827,479,505]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="9" pageNumber="10">TMSA</collectionCode>
.
<typeStatus id="41168851FA6FFFB8FCC7F9E5FC7FFE36" box="[858,965,479,505]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
: Same data as
<typeStatus id="41168851FA6FFFB8FB0AF9E5FABAFE36" box="[1175,1280,479,505]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
, (1
<collectionCode id="F8BCAE36FA6FFFB8FE95FA3DFEE4FDEE" box="[264,350,519,545]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="9" pageNumber="10">TMSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6FFFB8FEA5FA14FC7AFCF6" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="9E1236F3FA6FFFB8FEA5FA14FC7AFCF6" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">
<emphasis id="ACD9EAE1FA6FFFB8FEA5FA14FE0DFD87" bold="true" box="[312,439,558,584]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Similar to
<taxonomicName id="59AD4D70FA6FFFB8FDC1FA15FD41FD87" box="[604,763,559,584]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="ACD9EAE1FA6FFFB8FDC1FA15FD41FD87" box="[604,763,559,584]" italics="true" pageId="9" pageNumber="10">P. punctatum</emphasis>
</taxonomicName>
in dorsal sculpture, recognized by the more strongly sculptured, subrugulose pronotal foveae, the less rugulose frons foveae, the smaller elytral granules, the smaller elytral punctures, the more nearly equally developed rows of setae on the elytral series and intervals, and the shallower pronotal postocular emarginations (
<figureCitation id="06962A76FA6FFFB8FE20FAF5FDBFFD26" box="[445,517,719,745]" captionStart="FIGURE 6" captionStartId="19.[264,368,1577,1601]" captionTargetBox="[264,1322,310,1568]" captionTargetId="figure@19.[264,1323,309,1572]" captionTargetPageId="19" captionText="FIGURE 6. Pneuminion natalensis holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169052/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Reliable determinations of the two species will be based on examination of the quite dissimilar male genitalia. The aedeagus of
<taxonomicName id="59AD4D70FA6FFFB8FC78FACDFBC2FCDF" box="[997,1144,759,784]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="ACD9EAE1FA6FFFB8FC78FACDFBC2FCDF" box="[997,1144,759,784]" italics="true" pageId="9" pageNumber="10">P. natalensis</emphasis>
</taxonomicName>
is most similar to that of
<emphasis id="ACD9EAE1FA6FFFB8FEE4FB25FE4EFCF7" box="[377,500,799,824]" italics="true" pageId="9" pageNumber="10">P. v e l a m e n</emphasis>
(refer to the diagnosis of that species).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6FFFB8FEA5FB7CFE10F966" pageId="9" pageNumber="10" type="description">
<paragraph id="9E1236F3FA6FFFB8FEA5FB7CFD2DFB9E" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">
<emphasis id="ACD9EAE1FA6FFFB8FEA5FB7CFE6AFCAF" bold="true" box="[312,464,838,864]" pageId="9" pageNumber="10">Description.</emphasis>
Size (mm x 100; length/width): body (length to elytral apices) 163/74, head 26/44, pronotum 37/61, elytra 104/74. Dorsum piceus, legs and maxillary palpi dark brown; dorsal setae conspicuous, especially on head and pronotum, whitish. Labrum and clypeus dull, micropunctulate, except shining on anterior margin of clypeus. Frons disc moderately coarsely punctate, most punctures heart­shaped, interpunctal surface smooth, shining, punctures separated by ca. 1.5xpd; fovea in front of ocellus micropunctulate, medial margin moderately abrupt.
</paragraph>
<paragraph id="9E1236F3FA6FFFB8FEA5FC65FDCCFAA6" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">Pronotum rugulose in foveae and laterally, weakly shining on disc, dull laterally; punctures on reliefs small, ca. 0.25x size of heart­shaped punctures in U­shaped fovea, separated by ca. 13xpd, each with short whitish, granule­based seta; interpunctal surfaces on relief smooth and shining; oblique anterior fovea on each side shallowly connecting to midlongitudinal fovea; posterior U­shaped fovea well­developed; setae at lateral margin flat on cuticle in dry specimens. Pronotal postocular emargination deeper and less symmetrical in females than males.</paragraph>
<paragraph id="9E1236F3FA6FFFB8FEA5FD4DFEDAF9FE" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">Elytra with setae of series and intervals very nearly equally developed, resulting in discrete, narrowly separated, unilinear rows of setae, about 18 on each elytron, each seta granule­based; serial punctures moderately large, slightly wider than intervals, separated serially by less than puncture diameter; explanate margin narrow in male, slightly wider in female.</paragraph>
<paragraph id="9E1236F3FA6FFFB8FEA5FE05FE10F966" blockId="9.[264,1324,398,1745]" pageId="9" pageNumber="10">
Aedeagus: main­piece length 44, total length 61; main­piece distal portion with ridge/ groove to support duct and apex weakly hook­shaped; duct very slender over distal onehalf (
<figureCitation id="06962A76FA6FFFB8FEDBFEB5FE26F966" box="[326,412,1679,1705]" captionStart="FIGURE 16" captionStartId="29.[264,368,1700,1724]" captionTargetBox="[338,1203,316,1657]" captionTargetId="figure@29.[337,1249,315,1676]" captionTargetPageId="29" captionText="FIGURE 16. Pneuminion natalensis. Aedeagus of holotype." httpUri="https://zenodo.org/record/157476/files/figure.png" pageId="9" pageNumber="10">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6FFFB8FEA5FE8CFB35F91E" box="[312,1167,1718,1745]" pageId="9" pageNumber="10" type="etymology">
<paragraph id="9E1236F3FA6FFFB8FEA5FE8CFB35F91E" blockId="9.[264,1324,398,1745]" box="[312,1167,1718,1745]" pageId="9" pageNumber="10">
<emphasis id="ACD9EAE1FA6FFFB8FEA5FE8CFE7EF91F" bold="true" box="[312,452,1718,1744]" pageId="9" pageNumber="10">Etymology.</emphasis>
Named in reference to the known geographical distribution.
</paragraph>
</subSubSection>
</treatment>
</document>