99 lines
14 KiB
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99 lines
14 KiB
XML
<document id="D4C91E32BA073B330956C97FA0FE7736" ID-DOI="10.5281/zenodo.157462" ID-GBIF-Dataset="a9ccfc65-1459-414b-8a99-c99649f83d51" ID-ISSN="11755326" ID-Zenodo-Dep="157462" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461019330970" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2004" docId="160487E5FA65FFB5FE95F9E3FB80FC11" docLanguage="en" docName="zt00538.pdf" docOrigin="Zootaxa 538" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pneuminion Perkins 1997" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="EA3DFF9DFA66FFB1FF9DF83AFFBAFFCF" masterDocTitle="A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="4" updateTime="1698569824382" updateUser="plazi">
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<mods:title id="3C63F8EB0738D58E577443AAAAF578C6">A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="D65790903854C9D2E02CB5A47B7CF608">Perkins, Philip D.</mods:namePart>
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<mods:date id="5E8DDB7F602D01E5AE60620C68749AD3">2004</mods:date>
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<treatment id="160487E5FA65FFB5FE95F9E3FB80FC11" ID-DOI="http://doi.org/10.5281/zenodo.6271115" ID-GBIF-Taxon="119602179" ID-Zenodo-Dep="6271115" LSID="urn:lsid:plazi:treatment:160487E5FA65FFB5FE95F9E3FB80FC11" httpUri="http://treatment.plazi.org/id/160487E5FA65FFB5FE95F9E3FB80FC11" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="D6B76578FA65FFB2FE95F9E3FDF5FE3C" box="[264,591,473,499]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="9E1236F3FA65FFB2FE95F9E3FDF5FE3C" blockId="3.[264,591,473,499]" box="[264,591,473,499]" pageId="3" pageNumber="4">
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<heading id="C55A819FFA65FFB2FE95F9E3FDF5FE3C" bold="true" box="[264,591,473,499]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<taxonomicName id="59AD4D70FA65FFB2FE95F9E3FDF5FE3C" ID-CoL="8W2K4" authority="Perkins, 1997" authorityName="Perkins" authorityYear="1997" box="[264,591,473,499]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="ACD9EAE1FA65FFB2FE95F9E3FDF5FE3C" bold="true" box="[264,591,473,499]" pageId="3" pageNumber="4">
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<emphasis id="ACD9EAE1FA65FFB2FE95F9E3FE24FE3C" bold="true" box="[264,414,473,499]" italics="true" pageId="3" pageNumber="4">Pneuminion</emphasis>
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<bibRefCitation id="FA3C4B02FA65FFB2FE38F9E3FDF5FE3C" author="Perkins" box="[421,591,473,499]" pageId="3" pageNumber="14" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="D6B76578FA65FFB2FE95FA2DFBC9FDE2" box="[264,1139,534,558]" pageId="3" pageNumber="4" type="reference_group">
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<paragraph id="9E1236F3FA65FFB2FE95FA2DFBC9FDE2" blockId="3.[264,1139,534,558]" box="[264,1139,534,558]" pageId="3" pageNumber="4">
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<treatmentCitationGroup id="BEBD11DDFA65FFB2FE95FA2DFBC9FDE2" box="[264,1139,534,558]" pageId="3" pageNumber="4">
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<taxonomicName id="59AD4D70FA65FFB2FE95FA2DFDE2FDE2" ID-CoL="8W2K4" authority="Perkins, 1997: 171" authorityName="Perkins" authorityPageNumber="171" authorityYear="1997" box="[264,600,534,558]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="ACD9EAE1FA65FFB2FE95FA2DFE32FDE1" box="[264,392,535,558]" italics="true" pageId="3" pageNumber="4">Pneuminion</emphasis>
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<treatmentCitation id="1F0C10E2FA65FFB2FE0DFA2CFDE2FDE2" author="Perkins" box="[400,600,534,557]" page="171" pageId="3" pageNumber="4" year="1997">
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<bibRefCitation id="FA3C4B02FA65FFB2FE0DFA2CFD9CFDE2" author="Perkins" box="[400,550,534,557]" pageId="3" pageNumber="14" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
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: 171
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus id="41168851FA65FFB2FDFBFA2CFD2FFDE2" box="[614,661,534,557]" pageId="3" pageNumber="4">type</typeStatus>
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species
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<taxonomicName id="59AD4D70FA65FFB2FD6DFA2DFBD1FDE2" ID-CoL="4KKVG" authority="Perkins, 1997" authorityName="Perkins" authorityYear="1997" box="[752,1131,534,558]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="velamen">
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<emphasis id="ACD9EAE1FA65FFB2FD6DFA2DFC6AFDE1" box="[752,976,535,558]" italics="true" pageId="3" pageNumber="4">Pneuminion velamen</emphasis>
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<bibRefCitation id="FA3C4B02FA65FFB2FC4AFA2CFBD1FDE2" author="Perkins" box="[983,1131,534,557]" pageId="3" pageNumber="14" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
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</taxonomicName>
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)
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</paragraph>
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<subSubSection id="D6B76578FA65FFB2FE95FA69FEDBFC49" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="9E1236F3FA65FFB2FE95FA69FEDBFC49" blockId="3.[264,1323,595,1902]" pageId="3" pageNumber="4">
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<emphasis id="ACD9EAE1FA65FFB2FE95FA69FE32FDA2" bold="true" box="[264,392,595,621]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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Recognized by the combination of the transverse, noncordiform pronotum lacking hyaline margins and with welldeveloped postocular emarginations, the elytra with ten very regular series of punctures and all interseries equally developed (except
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<emphasis id="ACD9EAE1FA65FFB2FB2BFA9EFE8DFD2A" italics="true" pageId="3" pageNumber="4">P. i m p re s sum</emphasis>
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), the 11 articles of the antenna (six + club), the ocelli located midway between the midline and the eyes, the moderately short maxillary palpi with length ratios of last three articles as five:four:eight, the restricted abdominal hydrofuge pubescence, the very large wethypomeron, and the features of the postocular/hypomeral antennal pocket (see below).
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</paragraph>
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<subSubSection id="D6B76578FA65FFB5FEA5FBAEFB80FC11" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
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<paragraph id="9E1236F3FA65FFB2FEA5FBAEFEDEFB09" blockId="3.[264,1323,595,1902]" pageId="3" pageNumber="4">
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<emphasis id="ACD9EAE1FA65FFB2FEA5FBAEFE75FC61" bold="true" box="[312,463,916,942]" pageId="3" pageNumber="4">Description.</emphasis>
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Body size ca.
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<quantity id="59559B16FA65FFB2FD1DFBAEFD52FC61" box="[640,744,916,942]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="4" unit="mm" value="1.3">1.30 mm</quantity>
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to 2.00 mm, form elongate oval, moderately convex dorsally, head not deflexed. Antennomeres 11 (six + club), third slightly longer than combined lengths of fourth to sixth. Maxillary palpi moderately short, length ratios of articles two to four (last) as five: four: eight. Eyes welldeveloped, not elevated. Labrum large, set at angle to clypeus, apicomedially deeply incised; anterior margin fringed with long setae. Clypeus anterior margin straight. Ocelli located midway between midline and eyes, frons very weakly convex between ocelli; area behind and between ocelli lacking carinae.
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</paragraph>
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<paragraph id="9E1236F3FA65FFB2FEA5FCEEFCAFF9B1" blockId="3.[264,1323,595,1902]" pageId="3" pageNumber="4">
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Pronotum with anterior midlongitudinal fovea, behind which is Ushaped fovea; lateral depressions not demarked by foveae; lateral margin arcuate. Hyaline borders absent; setal fringe on base in front of scutellum. Elytra usually with ten rows of punctures. Prosternum in front of procoxae shorter than procoxal cavities, latter open behind (
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<figureCitation id="06962A76FA65FFB2FB4DFD76FAA1FAA9" box="[1232,1307,1356,1382]" captionStart="FIGURE 8" captionStartId="21.[264,368,1652,1676]" captionTargetBox="[264,1322,310,1644]" captionTargetId="figure@21.[264,1323,308,1648]" captionTargetPageId="21" captionText="FIGURE 8. Pneuminion semisulcatum holotype. Ventral habitus." httpUri="https://zenodo.org/record/169054/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
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). Metasternum with shallow midlongitudinal impression. Intercoxal sternite small, triangular. Legs moderately elongate, tarsi fivefivefive. Abdominal sterna onefour about equal in length, fifth slightly longer than fourth; in male, seventh concealed beneath sixth; in female seventh visible beyond posterior margin of sixth; first and basal part of second hydrofuge pubescent, remainder with sparse, fine, long setae. Hydrofuge pubescence also present on postocular area, bubblehypomeron (except antennal pockets), notal postcoxal projections, mesosternum, and metasternum.
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</paragraph>
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<paragraph id="9E1236F3FA65FFB2FEA5FEB6FD4AF8A1" blockId="3.[264,1323,595,1902]" pageId="3" pageNumber="4">
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Antennal pocket (terminology as in
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<bibRefCitation id="FA3C4B02FA65FFB2FD78FEB6FC33F969" author="Perkins" box="[741,905,1676,1702]" pageId="3" pageNumber="14" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
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) located in anterior face of prothorax, moderately deep, opened ventromedially (between hapsetae and the prosternum) and dorsally (at postocular emarginations); middle portion of antennal pocket convex, partially dividing pocket into dorsal and ventral concavity. About nine exocrine pores located along the medial margin of pocket, next to notosternal suture; otherwise pocket has very few, randomly distributed exocrine pores.
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</paragraph>
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<paragraph id="9E1236F3FA62FFB5FEA5F926FC7CFD51" blockId="4.[264,1323,284,990]" pageId="4" pageNumber="5">Hypomeral carina well defined, angulate posteriorly and becoming obsolete anteriorly at the antennal pocket. Bubblehypomeron narrow and very sparsely pubescent opposite notosternal suture. Wethypomeron very wide, slightly wider anteriorly than posteriorly, reticulate, nonsetose (except hapsetae) and with very sparse randomly distributed exocrine pores. Row of 12–15 welldeveloped hypomeral antennal pocket setae located along the ventral margin of antennal pocket; setae closely spaced, each widest in middle, tapering apically, and flattened; setae becoming progressively longer from most lateral to most medial, latter arising from anterior extreme of hydrofuge hypomeron, adjacent to lateral extreme of procoxal cavity. There are no organized clusters of exocrine pores or endapparatus, and no secretion sulcus exists on the hypomeron.</paragraph>
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<paragraph id="9E1236F3FA62FFB5FEA5FA96FE51FCD9" blockId="4.[264,1323,284,990]" pageId="4" pageNumber="5">A dense cluster of sharp spines, the prosternal antennal cleaner, is located on anterolateral margin of prosternum. The antennal cleaner is surrounded by sparsely pubescent asperite hydrofuge.</paragraph>
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<paragraph id="9E1236F3FA62FFB5FEA5FB1EFB80FC11" blockId="4.[264,1323,284,990]" pageId="4" pageNumber="5">Periocular exocrine pores are present along subocular antennal groove and at posterior margin of eye; medial margin of antennal groove raised slightly, not cariniform; few exocrine pores located near posteroventral angle of eye; welldeveloped postocular ridge delimits postocular and subocular areas; postocular area clothed in hydrofuge pubescence, but lacks any specialized exocrine secretion delivery setae or sensilla.</paragraph>
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</subSubSection>
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</treatment>
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