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<treatment id="15830D1A8129D96656AF3E65BAAF168E" ID-GBIF-Taxon="231525340" LSID="urn:lsid:plazi:treatment:15830D1A8129D96656AF3E65BAAF168E" httpUri="http://treatment.plazi.org/id/15830D1A8129D96656AF3E65BAAF168E" lastPageNumber="930" pageNumber="930" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13000065">
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<paragraph id="D35E5C69EFB85FEDCBDA481B07BE9BCB" pageNumber="930">
<taxonomicName id="3CA828C5D6188D062A3598334FC79694" ID-CoL="JDZ" authority="Roberts 1951" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae">Cricetomyinae Roberts 1951</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="E4674885D79297E85AC394589A5DFA16" pageNumber="930">
<treatmentCitation id="0B25D6C87A4257F203E03EE2360B07EB" author="Roberts" authority="Roberts 1951" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" page="434" pageNumber="930" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae" title="Cricetomyinae" volumeTitle="Mammals of South Africa" year="1951">
<taxonomicName id="EE2AB34D006D32ADCBC71077E7C7A694" ID-CoL="JDZ" authority="Roberts 1951" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae">
Cricetomyinae
<bibRefCitation id="E7CD3B13B04A10FC2297D3DB52AA823E" author="Roberts, A." refId="ref306581" refString="Roberts, A. 1951. The mammals of South Africa. Trustees of &quot; The mammals of South Africa &quot; book fund, Johannesburg, 700 pp." year="1951">Roberts 1951</bibRefCitation>
</taxonomicName>
,
<bibCitation id="93F3B3E49D98921321386FE99E5AB68C" author="Roberts" pageNumber="930" pagination="434" title="Cricetomyinae" volumeTitle="Mammals of South Africa" year="1951">
Mammals of
<collectingCountry id="FDF16DB40FB2732477DB1DCFDD36F8AA" name="South Africa">South Africa</collectingCountry>
: 434
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FCFE4047203C1766C93A9803C36B2225" pageNumber="930" type="synonym_list">
<paragraph id="48A299F12BB40D1ED7945FD8FA4B891F" pageNumber="930">
<emphasis id="52569F472A4E13BBA021A0E284824214" inLineHeading="true" pageNumber="930">Synonyms:</emphasis>
<taxonomicName id="38A9B5E0B3394D5F25A5F4D216F50809" authority="Chaline, Mein, and F. Petter 1977" authorityName="Chaline, Mein, and F. Petter" authorityYear="1977" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="subFamily" subFamily="Cricetomyidae">
Cricetomyidae Chaline, Mein, and F.
<bibRefCitation id="77FDB812FCEC8D248219F63F260AFB7E" author="Petter, J. - J. &amp; R. Albignac &amp; Y. Rumpler" refId="ref285142" refString="Petter, J. - J., R. Albignac, and Y. Rumpler. 1977. Faune de Madagascar. 44. Mammiferes lemuriens. O. R. S. T. O. M., Paris, 513 pp." year="1977">Petter 1977</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E12AE533EFBB2084805D19570BA899D8" authority="Roberts 1951" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="subFamily" subFamily="Saccostomurinae">
Saccostomurinae
<bibRefCitation id="302E4ECF986AF91A2A3B182CB36C3485" author="Roberts, A." refId="ref306581" refString="Roberts, A. 1951. The mammals of South Africa. Trustees of &quot; The mammals of South Africa &quot; book fund, Johannesburg, 700 pp." year="1951">Roberts 1951</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C4CE541524BD8B09FE831EA1D2369B72" pageNumber="930" type="child_taxa">
<paragraph id="BB527FC390D812A1EACD6C3C8E31A6E4" pageNumber="930">
<emphasis id="288B92A6014308B97F52C467A4FD42FE" inLineHeading="true" pageNumber="930">Genera:</emphasis>
3 genera with 8 species:
</paragraph>
<paragraph id="4004318665EC6D57CFB48EC6F62C1876" pageNumber="930">
Genus
<taxonomicName id="1CE90CC3099A44081F3F4495448E6415" authority="Thomas 1909" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Nesomyidae" genus="Beamys" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="genus">
<emphasis id="1FCF66922084DA041F6AAB29A075B0E5" italics="true" pageNumber="930">Beamys</emphasis>
Thomas 1909
</taxonomicName>
(2 species)
</paragraph>
<paragraph id="9ECEB5C5B3AA540AF139B915B600878F" pageNumber="930">
Genus
<taxonomicName id="106641F3253194A79FA1E4ACC664836F" authority="Waterhouse 1840" authorityName="Waterhouse" authorityYear="1840" class="Mammalia" family="Nesomyidae" genus="Cricetomys" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="genus">
<emphasis id="15549B43E9564053AA434DED475095F2" italics="true" pageNumber="930">Cricetomys</emphasis>
Waterhouse 1840
</taxonomicName>
(4 species)
</paragraph>
<paragraph id="C9B8242CA98E329DD715962DD4DD3272" pageNumber="930">
Genus
<taxonomicName id="3F6401EA8B55FFCDD662076EEDDB29CF" authority="Peters 1846" authorityName="Peters" authorityYear="1846" class="Mammalia" family="Nesomyidae" genus="Saccostomus" kingdom="Animalia" order="Rodentia" pageNumber="930" phylum="Chordata" rank="genus">
<emphasis id="485CE744FD4CF9007D4A2B977D738D87" italics="true" pageNumber="930">Saccostomus</emphasis>
Peters 1846
</taxonomicName>
(2 species)
</paragraph>
</subSubSection>
<subSubSection id="6C13CCDCB97275E809C5344379E3D391" pageNumber="930" type="discussion">
<paragraph id="761B67D332F092171BF13CD935904892" pageNumber="930">
<emphasis id="2CBA28F6B42EE426C99901297EA25296" inLineHeading="true" pageNumber="930">Discussion:</emphasis>
<paragraph id="D396154246DA89DEDD3E3C525B17D1AC" pageNumber="930">
Although initially classified within Murinae (e.g.,
<bibRefCitation id="D40E8C2FD9FB84DE5CA6477712B65724" author="Ellerman, J. R." refId="ref92857" refString="Ellerman, J. R. 1941. The families and genera of living rodents. Vol. II. Family Muridae. British Museum (Natural History), London, 690 pp." year="1941">Ellerman, 1941</bibRefCitation>
;
<bibRefCitation id="EBC6A3CA943A0E8A22A051CE1F82C2C2" author="Simpson, G. G." refId="ref328876" refString="Simpson, G. G. 1945. The principles of classification and a classification of mammals. Bulletin of the American Museum of Natural History, 85: 1 - 350." year="1945">Simpson, 1945</bibRefCitation>
;
<bibRefCitation id="E63D680A5E4B29BFAED7B41E198B6191" author="Thomas, O." refId="ref352089" refString="Thomas, O. 1896 [1897]. On the genera of rodents: An attempt to bring up to date the current arrangement of the order. Proceedings of the Zoological Society of London, 1896: 1012 - 1028." year="1896">Thomas, 1896</bibRefCitation>
), systematists have acknowledged a closer affinity of African pouched rats to one another than to other murines, a distinction later formalized by Roberts (1951) diagnosis of
<taxonomicName id="CAB230B5847BEEBA6E5FFE6C1EFF04B3" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae">Cricetomyinae</taxonomicName>
. Most subsequent systematic arrangements have followed F.
<bibRefCitation id="4692DACEC6CBF0212908131648FEF29D" author="Petter, F." refId="ref283596" refString="Petter, F. 1966 a. Affinites des genres Beamys, Saccostomus et Cricetomys (Rongeurs, Cricetomyinae). Annales Musee Royal de l'Afrique Centrale, ser. 8 (Sciences Zoologiques), 144: 13 - 25." year="1966">
Petter (1966
<emphasis id="F36E8EE58601AB1E8D4925A80EEA3AEB" italics="true" pageNumber="930">a</emphasis>
)
</bibRefCitation>
in allying cricetomyines with cricetids (Reig, 1980, 1981;
<bibRefCitation id="432593889C4ABAC11CEC1E3EAA662E50" author="Rosevear, D. R." refId="ref310288" refString="Rosevear, D. R. 1969. The rodents of West Africa. Trustees of the British Museum (Natural History), London, 677: 1 - 604." year="1969">Rosevear, 1969</bibRefCitation>
) based on his interpretation of molar cusp homologies.
<bibRefCitation id="F4C32AED891C29C5C230A67B9C4C043A" author="Lavocat, R." refId="ref205495" refString="Lavocat, R. 1973. Les rongeurs du Miocene d'Afrique Orientale. Memoires et Travaux de l'Ecole Pratique des Hautes Etudes, Institut de Montpellier, 1: 1 - 284." year="1973">Lavocat (1973</bibRefCitation>
,
<bibRefCitation id="B4BDF7A65AA78CC29757400006B47242" author="Lavocat, R." refId="ref205553" refString="Lavocat, R. 1978. Rodentia and Lagomorpha. Pp. 69 - 89, in Evolution of African mammals (V. J. Maglio and H. B. S. Cooke, eds.). Harvard University Press, Cambridge, MA, 641 pp." year="1978">1978</bibRefCitation>
) first suggested phylogenetic association with other archaic African groups in a broadly defined
<taxonomicName id="E2B750505EE57AF984A9EEA46CD43DFD" authorityName="Major" authorityYear="1897" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="family">Nesomyidae</taxonomicName>
, an interpretation generally endorsed by
<bibRefCitation id="9105AAB76E99D37855C3FF007B099E99" author="Carleton, M. D. &amp; G. G. Musser" refId="ref52182" refString="Carleton, M. D., and G. G. Musser. 1984. Muroid rodents. Pp. 289 - 379, in Orders and families of Recent mammals of the world (S. Anderson and J. K. Jones, Jr., eds.). John Wiley and Sons, New York, 686 pp." year="1984">Carleton and Musser (1984)</bibRefCitation>
. Configuration of molar cusps (F.
<bibRefCitation id="42F7764B956D5884B252E0D91CFAD14D" author="Petter, F." refId="ref283596" refString="Petter, F. 1966 a. Affinites des genres Beamys, Saccostomus et Cricetomys (Rongeurs, Cricetomyinae). Annales Musee Royal de l'Afrique Centrale, ser. 8 (Sciences Zoologiques), 144: 13 - 25." year="1966">
Petter, 1966
<emphasis id="947D61782707A8050B911C5AD20B7D90" italics="true" pageNumber="930">a</emphasis>
</bibRefCitation>
), anatomy of internal cheek pouches (
<bibRefCitation id="08FD2498629014277C86C1DC9483CA23" author="Ryan, J. M." refId="ref312988" refString="Ryan, J. M. 1989 b. Evolution of cheek pouches in African pouched rats (Rodentia: Cricetomyinae). Journal of Mammalogy, 70: 267 - 274." year="1989">
Ryan, 1989
<emphasis id="CFDCF45612F22839849D6967C7F00DAD" italics="true" pageNumber="930">b</emphasis>
</bibRefCitation>
), and mitochondrial and nuclear DNA sequence data (
<bibRefCitation id="46F5D77403C993AEC2D3B741E835287B" author="Jansa, S. A. &amp; S. M. Goodman &amp; P. K. Tucker" refId="ref172368" refString="Jansa, S. A., S. M. Goodman, and P. K. Tucker. 1999. Molecular phylogeny and biogeography of the native rodents of Madagascar (Muridae: Nesomyinae): A test of the single-origin hypothesis. Cladistics, 15: 253 - 270." year="1999">Jansa et al., 1999</bibRefCitation>
;
<bibRefCitation id="3079AE2B1E404BF9AA943A9908FA6D43" author="Michaux, J. &amp; A. Reyes &amp; F. Catzeflis" refId="ref240856" refString="Michaux, J., A. Reyes, and F. Catzeflis. 2001 b. Evolutionary history of the most speciose mammals: Molecular phylogeny of muroid rodents. Molecular Biology and Evolution, 18 (11): 2017 - 2031." year="2001">
Michaux et al., 2001
<emphasis id="14703942BC17469FE257A6AA80B4895C" italics="true" pageNumber="930">b</emphasis>
</bibRefCitation>
;
<bibRefCitation id="301ED4A41304AD8ADEA3CC84515D103B" author="Michaux, J. &amp; F. Catzeflis" refId="ref240457" refString="Michaux, J., and F. Catzeflis. 2000. The bushlike radiation of muroid rodents is exemplified by the molecular phylogeny of the LCAT nuclear gene. Molecular Phylogenetics and Evolution, 17: 280 - 293." year="2000">Michaux and Catzeflis, 2000</bibRefCitation>
) support the monophyly of the subfamily, which forms an unresolved polytomy with species of
<taxonomicName id="D76D095708F2707FD458F61927A91DDD" authorityName="G. M. Allen" authorityYear="1939" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subFamily" subFamily="Dendromurinae">Dendromurinae</taxonomicName>
and
<taxonomicName id="96B955286E38C830595C920E5EEF0F24" authorityName="Wagner" authorityYear="1841" class="Mammalia" family="Nesomyidae" genus="Mystromys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="BC3FC8B8362D81F0AA1EA9068A1C6E85" italics="true" pageNumber="930">Mystromys</emphasis>
</taxonomicName>
according to phyletic evaluation of nuclear protein-coding genes (
<bibRefCitation id="00A2B922A51E7182B5CD0996E395CBED" author="Michaux, J. &amp; A. Reyes &amp; F. Catzeflis" refId="ref240856" refString="Michaux, J., A. Reyes, and F. Catzeflis. 2001 b. Evolutionary history of the most speciose mammals: Molecular phylogeny of muroid rodents. Molecular Biology and Evolution, 18 (11): 2017 - 2031." year="2001">
Michaux et al., 2001
<emphasis id="9730A0AA7932EFE39A71C2FE5CE64096" italics="true" pageNumber="930">b</emphasis>
</bibRefCitation>
;
<bibRefCitation id="FDA2CBF4299F69DC5D0BD93E950E0447" author="Michaux, J. &amp; F. Catzeflis" refId="ref240457" refString="Michaux, J., and F. Catzeflis. 2000. The bushlike radiation of muroid rodents is exemplified by the molecular phylogeny of the LCAT nuclear gene. Molecular Phylogenetics and Evolution, 17: 280 - 293." year="2000">Michaux and Catzeflis, 2000</bibRefCitation>
).
</paragraph>
<paragraph id="E096257121C46DF7409CD43C41434EBF" pageNumber="930">
General and diagnostic morphological traits, taxonomic history, and phylogenetic interpretations summarized by
<bibRefCitation id="4546BD20583203E2A1D2BF4B06332C53" author="Carleton, M. D. &amp; G. G. Musser" refId="ref52182" refString="Carleton, M. D., and G. G. Musser. 1984. Muroid rodents. Pp. 289 - 379, in Orders and families of Recent mammals of the world (S. Anderson and J. K. Jones, Jr., eds.). John Wiley and Sons, New York, 686 pp." year="1984">Carleton and Musser (1984)</bibRefCitation>
. Certain early to middle Miocene taxa of East Africa (e.g.,
<taxonomicName id="562E892194D58BF155F9F06BC82240D6" authorityName="Lavocat" authorityYear="1964" class="Mammalia" family="Muridae" genus="Leakeymys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="AD0E5846F3D32FFF9B1EE1E39B6A0498" italics="true" pageNumber="930">Leakeymys</emphasis>
</taxonomicName>
and
<taxonomicName id="69425DFA4708C11A0BC381E809D0FF8C" authorityName="Lavocat" authorityYear="1973" class="Mammalia" family="Muridae" genus="Notocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="3E8EB1845EC41A0001BAE29C6F87503C" italics="true" pageNumber="930">Notocricetodon</emphasis>
</taxonomicName>
) have been speculatively linked to the autochthonous African origin of
<taxonomicName id="06C87E175BE627C1FAB60D8B3D1766F0" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae">Cricetomyinae</taxonomicName>
, but their phyletic association so far lacks persuasive demonstration (see
<bibRefCitation id="E68950BE8859F7B3E53EB450EA9DCFDA" author="Chaline, J. &amp; P. Mein &amp; F. Petter" refId="ref57068" refString="Chaline, J., P. Mein, and F. Petter. 1977. Les grandes lignes d'une classification evolutive des Muroidea. Mammalia, 41: 245 - 252." year="1977">Chaline et al., 1977</bibRefCitation>
;
<bibRefCitation id="AD713513154962A4B41F88F096C22AA5" author="Tong, H. &amp; J. - J. Jaeger" refId="ref357989" refString="Tong, H., and J. - J. Jaeger. 1993. Muroid rodents from the Middle Miocene Fort Ternan locality (Kenya) and their contribution to the phylogeny of muroids. Palaeontographica. Abteilung A, Palaozoologie-Stratigraphie, 229 (1 - 3): 51 - 73." year="1993">Tong and Jaeger, 1993</bibRefCitation>
). Indisputable fossil representatives extend from the early Pliocene to the Recent of eastern and southern Africa (
<bibRefCitation id="9FB68841D5E3E2F9ED5A96798125187C" author="McKenna, M. C. &amp; S. K. Bell" refId="ref233772" refString="McKenna, M. C., and S. K. Bell. 1997. Classification of mammals above the species level. Columbia University Press, New York, 631 pp." year="1997">McKenna and Bell, 1997</bibRefCitation>
).
<bibRefCitation id="ABD5D767A7F53E0F0FE376B42E770045" author="Roberts, A." refId="ref306581" refString="Roberts, A. 1951. The mammals of South Africa. Trustees of &quot; The mammals of South Africa &quot; book fund, Johannesburg, 700 pp." year="1951">Roberts (1951)</bibRefCitation>
taxonomically segregated
<taxonomicName id="0DBD9A5CDBF2A436847730E6A3FBE656" authority="(Saccostomurinae)" baseAuthorityName="Saccostomurinae" class="Mammalia" family="Nesomyidae" genus="Saccostomus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="D4A0B07B4477C808F86600C0D2943243" italics="true" pageNumber="930">Saccostomus</emphasis>
(Saccostomurinae)
</taxonomicName>
from other African pouched rats (
<taxonomicName id="5CE731341581770821643BA3E646832B" authorityName="Roberts" authorityYear="1951" class="Mammalia" family="Nesomyidae" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subFamily" subFamily="Cricetomyinae">Cricetomyinae</taxonomicName>
), a division not recognized by later systematists (F.
<bibRefCitation id="58DBB9D0FA974DA92CCBBEAF3BEE07FF" author="Petter, F." refId="ref283596" refString="Petter, F. 1966 a. Affinites des genres Beamys, Saccostomus et Cricetomys (Rongeurs, Cricetomyinae). Annales Musee Royal de l'Afrique Centrale, ser. 8 (Sciences Zoologiques), 144: 13 - 25." year="1966">
Petter, 1966
<emphasis id="7F3EFDAE4EBC040C467FD41B6BD4FD2A" italics="true" pageNumber="930">a</emphasis>
</bibRefCitation>
;
<bibRefCitation id="C83B2E84E987EE28E416426E46E66E76" author="Ryan, J. M." refId="ref312988" refString="Ryan, J. M. 1989 b. Evolution of cheek pouches in African pouched rats (Rodentia: Cricetomyinae). Journal of Mammalogy, 70: 267 - 274." year="1989">
Ryan, 1989
<emphasis id="32BAEFAE3132AEF3A80DFC1220F12308" italics="true" pageNumber="930">b</emphasis>
</bibRefCitation>
). Many morphological traits, however, depict
<taxonomicName id="AEE3A18964F94157366AC64C3200AFAD" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Nesomyidae" genus="Beamys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="DBA35DF44F9DCF55B2E81242850BB1A2" italics="true" pageNumber="930">Beamys</emphasis>
</taxonomicName>
and
<taxonomicName id="61AE6226187AF479098C4B4914ECF762" authorityName="Waterhouse" authorityYear="1840" class="Mammalia" family="Nesomyidae" genus="Cricetomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="79A8F07063238DA7160B24D4A926C64A" italics="true" pageNumber="930">Cricetomys</emphasis>
</taxonomicName>
as sister species apart from
<taxonomicName id="C179C5ACDA9F3371ACDA52597300391E" class="Mammalia" family="Nesomyidae" genus="Saccostomus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="58D785A7E30862C72D7FC7177ACD540B" italics="true" pageNumber="930">Saccostomus</emphasis>
</taxonomicName>
, as enumerated by
<bibRefCitation id="5C29EC125F2373C77E4FE476B7D435F6" author="Carleton, M. D. &amp; G. G. Musser" refId="ref52182" refString="Carleton, M. D., and G. G. Musser. 1984. Muroid rodents. Pp. 289 - 379, in Orders and families of Recent mammals of the world (S. Anderson and J. K. Jones, Jr., eds.). John Wiley and Sons, New York, 686 pp." year="1984">Carleton and Musser (1984)</bibRefCitation>
, and we employ those family-group taxa as tribes. The closer affinity of
<taxonomicName id="5D828B8DA89F570170072439EAAC0E59" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Nesomyidae" genus="Beamys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="58B4DC9B27D2A32601824E29F3E49011" italics="true" pageNumber="930">Beamys</emphasis>
</taxonomicName>
and
<taxonomicName id="C3E836C5CFF7F20DEF1B47E62AF79AF8" authorityName="Waterhouse" authorityYear="1840" class="Mammalia" family="Nesomyidae" genus="Cricetomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="D11B6ADB79F7DE112BF9FEB9FCE33958" italics="true" pageNumber="930">Cricetomys</emphasis>
</taxonomicName>
is also reflected in gene-sequence analyses (
<bibRefCitation id="E01874481CE00614E4740427B3FB99A7" author="Corti, M. &amp; R. Castiglia &amp; F. Annesi &amp; W. Verheyen" refId="ref68783" refString="Corti, M., R. Castiglia, F. Annesi, and W. Verheyen. 2004. Mitochondrial sequences and karyotypes reveal hidden diversity in African pouched mice (Subfamily Cricetomyinae, Genus Saccostomus). Journal of Zoology, London, 262: 1 - 12." year="2004">Corti et al., 2004</bibRefCitation>
)
</paragraph>
.
</paragraph>
</subSubSection>
</treatment>
</document>