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<document ID-CLB-Dataset="24755" ID-DOI="http://dx.doi.org/10.3897/zookeys.893.46923" ID-GBIF-Dataset="c941edd5-de40-4cbf-aa06-1baac5d36f1c" ID-PMC="PMC6901611" ID-Pensoft-Pub="1313-2970-893-91" ID-Pensoft-UUID="E8A0151ADE0F532CA16E0710F6F2BC50" ID-PubMed="31844401" ID-ZooBank="4E267EC6A94F4227ADF5700DC17F0D87" ModsDocID="1313-2970-893-91" checkinTime="1575325048369" checkinUser="pensoft" docAuthor="Ohtsuka, Susumu &amp; Boxshall, Geoffrey A." docDate="2019" docId="1490FC06A592512B8B14F71B4E47B420" docLanguage="en" docName="ZooKeys 893: 91-113" docOrigin="ZooKeys 893" docSource="http://dx.doi.org/10.3897/zookeys.893.46923" docTitle="Caligus kajii Ohtsuka &amp; Boxshall, 2019, sp. nov." docType="treatment" docUuid="9A3A7B46-6140-4865-AAFA-99E718FBA8D6" docUuidSource="ZooBank" docVersion="7" id="E8A0151ADE0F532CA16E0710F6F2BC50" lastPageNumber="91" masterDocId="E8A0151ADE0F532CA16E0710F6F2BC50" masterDocTitle="Two new species of the genus Caligus (Crustacea, Copepoda, Siphonostomatoida) from the Sea of Japan, with a note on the establishment of a new species group" masterLastPageNumber="113" masterPageNumber="91" pageNumber="91" updateTime="1732848802802" updateUser="ExternalLinkService">
<mods:mods id="A5127F0359AC3148EEDAD726356CA247" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="91E87C651CABC451F7D575469FD3EB40">
<mods:title id="C56F7841DE722C69D2C29B4F2D7577FA">Two new species of the genus Caligus (Crustacea, Copepoda, Siphonostomatoida) from the Sea of Japan, with a note on the establishment of a new species group</mods:title>
</mods:titleInfo>
<mods:name id="4634EDF1BCED7E77722B73D23469444A" type="personal">
<mods:role id="841A42E624ACDEDE8F798A6E2AE4FF13">
<mods:roleTerm id="42DF27C7EACA3D3761D28592F48A8083">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="76F7A0D9FF15D854A058B9C3F9F30591">Ohtsuka, Susumu</mods:namePart>
</mods:name>
<mods:name id="A6E4E102A1082FDC131CEC5FDB2791C0" type="personal">
<mods:role id="B78B81F410509A274E5BAF023FCA3D65">
<mods:roleTerm id="A62CAB4347DB2E2553D5F7F1024303DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96F5697970BC127DBE48FF45B51DDA8A">Boxshall, Geoffrey A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CB67CD6960638C2ACB71F3AA4C617650">text</mods:typeOfResource>
<mods:relatedItem id="87EF6B4CA3E3583D1115CCF286B02B32" type="host">
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<mods:title id="C7D015CF0E8E84CBAF95C9921CBBD70B">ZooKeys</mods:title>
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<mods:part id="BA464BCE9EBACD43A691AD598C95FDC5">
<mods:date id="8F97DD7D5E83C8A81A6AA0A1CA3AB291">2019</mods:date>
<mods:detail id="15435D4532E6CDFF87432AE296AB3F2B" type="volume">
<mods:number id="D5F3DF3994E117DBD73BF04D2519537A">893</mods:number>
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<mods:start id="EB530E4CA4230B355428352D0847D334">91</mods:start>
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<mods:location id="78FD2316D2C4D361DA894DE880AD6C76">
<mods:url id="70C6AD0548B3672D6F3EBDAF524106AC">http://dx.doi.org/10.3897/zookeys.893.46923</mods:url>
</mods:location>
<mods:classification id="AC21DF7E538C5AD3E9EF7BD6260BE0E4">journal article</mods:classification>
<mods:identifier id="759048BB28CE8678E4D6C309C761B561" type="DOI">http://dx.doi.org/10.3897/zookeys.893.46923</mods:identifier>
<mods:identifier id="95F87E08C47BF37E9469058EA0B5D3CF" type="Pensoft-Pub">1313-2970-893-91</mods:identifier>
<mods:identifier id="265A62928580FF6270BCA4538DAFDA79" type="ZooBank">4E267EC6A94F4227ADF5700DC17F0D87</mods:identifier>
<mods:identifier id="09E060A7855E2D6AB50BE06D5D3786FA" type="Pensoft-UUID">E8A0151ADE0F532CA16E0710F6F2BC50</mods:identifier>
</mods:mods>
<treatment id="1490FC06A592512B8B14F71B4E47B420" ID-GBIF-Taxon="160612157" LSID="urn:lsid:zoobank.org:act:9A3A7B46-6140-4865-AAFA-99E718FBA8D6" httpUri="http://treatment.plazi.org/id/1490FC06A592512B8B14F71B4E47B420" lastPageNumber="91" pageId="0" pageNumber="91" scope_class="Copepoda" scope_family="Caligidae" scope_order="Shiphonostomatoida">
<subSubSection id="B0DE19C07CD14659C6CF431C0136A184" pageId="0" pageNumber="91" type="nomenclature">
<paragraph id="0013D8F330D5428D2403880C3F315C0C" pageId="0" pageNumber="91">
<taxonomicName id="F88228B3F303394225A0B1E12C6DFD32" ID-CoL="7ZG3W" LSID="1490FC06-A592-512B-8B14-F71B4E47B420" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus kajii" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="kajii">Caligus kajii</taxonomicName>
<taxonomicNameLabel id="E3F033AB772937D8D6118A6F445517AA" pageId="0" pageNumber="91">sp. nov.</taxonomicNameLabel>
<figureCitation id="ABF028BFEE1B9BB54F1D0E9EC08B63C4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Figs 4</figureCitation>
,
<figureCitation id="635C72EA33C11FA9AC127916A0E67F23" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">5</figureCitation>
,
<figureCitation id="9EE398E21FBE89BA5D96711775778BD1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">6</figureCitation>
<tableCitation id="CA54667F530B557AFAC8BDB90F145C61" captionStartId="T2" httpUri="http://table.plazi.org/id/28BD004225F4678E2FF3806667A522D0" pageId="0" pageNumber="91" tableUuid="28BD004225F4678E2FF3806667A522D0">, Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="3E5770EEA9510FD48719A489461A5B98" pageId="0" pageNumber="91" type="materials_examined">
<paragraph id="E61057253B02C6B91D48B06E573AC263" pageId="0" pageNumber="91">Material examined.</paragraph>
<paragraph id="EF7A92B5ED296392FBB656136984889B" pageId="0" pageNumber="91">
JAPAN • 38 adult ♀♀ and 14 adult ♂♂; parasitic on body surface of
<taxonomicName id="7AAEEB0419AAB909FA59E4F890594D40" family="Platycephalidae" genus="Platycephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platycephalus" order="Scorpaeniformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis id="87569C26668C227DD658CA78CA9C8302" italics="true" pageId="0" pageNumber="91">Platycephalus</emphasis>
</taxonomicName>
sp. (total length 58 cm) collected from a depth of 15 m off Shimonoseki, Yamaguchi Prefecture (
<geoCoordinate id="DBE29E2CBB06870E572388D5A2FF59C7" degrees="34" direction="north" minutes="00.686" orientation="latitude" precision="1" value="34.011433">34°00.686'N</geoCoordinate>
,
<geoCoordinate id="6D42E376CE614C8ECC5AFB641BF6DFE9" degrees="130" direction="east" minutes="53.756" orientation="longitude" precision="1" value="130.89594">130°53.756'E</geoCoordinate>
); morning of August 24, 2016; S. Ohtsuka leg.
</paragraph>
</subSubSection>
<subSubSection id="B062C8704820034243CD1084A7819EDF" pageId="0" pageNumber="91" type="types">
<paragraph id="FCDA8A7BACBD73BD896590D83EA9BB61" pageId="0" pageNumber="91">Types.</paragraph>
<paragraph id="7E43BDC301DB64CA73B62E57F0487B9B" pageId="0" pageNumber="91">
<emphasis id="FAE3F803645DCBE74147476DEAC0D840" bold="true" pageId="0" pageNumber="91">
<emphasis id="4D039DAC3CBE05FD61044910D60657E4" italics="true" pageId="0" pageNumber="91">Holotype</emphasis>
.
</emphasis>
JAPAN •1 ovigerous adult ♀; parasitic on body surface of
<taxonomicName id="D9572E1D33310028A9A34E23D95E004B" family="Platycephalidae" genus="Platycephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platycephalus" order="Scorpaeniformes" pageId="0" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis id="6AC0C5316517F70F40B7D1F8FD0E8609" italics="true" pageId="0" pageNumber="91">Platycephalus</emphasis>
</taxonomicName>
sp. (total length 58 cm) collected from a depth of 15 m off Shimonoseki, Yamaguchi Prefecture (
<geoCoordinate id="8EAC771F0DF54EDAB74C83C93CC1E576" degrees="34" direction="north" minutes="00.686" orientation="latitude" precision="1" value="34.011433">34°00.686'N</geoCoordinate>
,
<geoCoordinate id="EFAAB68DCB0684BD69545F30AE6F3637" degrees="130" direction="east" minutes="53.756" orientation="longitude" precision="1" value="130.89594">130°53.756'E</geoCoordinate>
); morning of August 24, 2016; whole specimen (NSMT-Cr 26754); S. Ohtsuka leg.
<emphasis id="9E55A913CF9B5B94D1F2C1542DB2E309" bold="true" pageId="0" pageNumber="91">
<emphasis id="1DF89D074A30D4CB9AB77033126CECF0" italics="true" pageId="0" pageNumber="91">Allotype</emphasis>
.
</emphasis>
JAPAN•1 adult ♂, same data as in holotype; partly dissected on 1 slide, body in vial (NSMT-Cr 26755); S. Ohtsuka leg.
<emphasis id="7463C3F19E6F1A5D564F92C0670C5831" bold="true" pageId="0" pageNumber="91">
<emphasis id="95D6F039C75FA9131EDC1C6443D157BC" italics="true" pageId="0" pageNumber="91">Paratypes</emphasis>
.
</emphasis>
JAPAN•1♀, same data as in holotype; partly dissected and bodies in vials (NSMT-Cr 26756); 36♀♀ and 13♂♂, same data as in holotype; whole specimens (NSMT-Cr 26757); S. Ohtsuka leg.
</paragraph>
</subSubSection>
<subSubSection id="40B30EC351E7870A8E0FDA5E7B648B4E" pageId="0" pageNumber="91" type="description">
<paragraph id="35306E13B77EF3D7F3A7BA4F9BCD1FFC" pageId="0" pageNumber="91">Description.</paragraph>
<paragraph id="0260FA7B9EA993C1C00D03EB7C4F6E3E" pageId="0" pageNumber="91">
<emphasis id="31330167F91AAD39A2C3D0A0AC41A65A" bold="true" pageId="0" pageNumber="91">Female.</emphasis>
Body length of holotype 6.16 mm, 4.86-6.16 mm in holotype and female paratypes (mean
<normalizedToken id="986EA003B77E12F6F0215DBCF497D85C" originalValue="±">+/-</normalizedToken>
standard deviation = 5.49
<normalizedToken id="AF22F5EBDC7EA293CE9C93670D98135D" originalValue="±">+/-</normalizedToken>
0.32 mm,
<emphasis id="5629694451B2FD17608FDB4AD51F9D41" italics="true" pageId="0" pageNumber="91">N</emphasis>
= 38). Dorsal cephalothoracic shield subcircular, almost as long as wide (
<figureCitation id="7334E212F1F44A3CF087B0E71E0CCF69" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4A</figureCitation>
). Lunules (
<figureCitation id="731958CBC56E01D9F8B2FE8DB9567885" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4A</figureCitation>
) relatively small. Pediger 4 almost completely fused to genital complex. Genital complex (
<figureCitation id="D049E7BBCA30307ACF307AFF9DCD04B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4A</figureCitation>
) subquadrate, about 1.14 times longer than wide, produced posteroventrally into pair of rounded processes between which paired copulatory pores located (
<figureCitation id="448AF7179CE0A1DC3B55A68424774442" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4B</figureCitation>
). Spermatophores (
<figureCitation id="94CB7BED0E6B8077E3D73430C731A320" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4B</figureCitation>
) attached to copulatory pores via fine tubules; spermatophore proper globular, ca. 0.12 mm in diameter. Paired egg strings of holotype female containing 22 and 24 eggs. Abdomen (
<figureCitation id="06967420DE6E3F2A1D0082AC78FD718E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4A, B</figureCitation>
) 1-segmented, about as long as wide. Caudal ramus (
<figureCitation id="5BCBA5622F8880F9DD5B1330F27977CB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4A, B</figureCitation>
) furnished with rows of setules along inner and outer margins; seta II minute, located near base of seta III on subterminal ventral surface, seta III subterminal, setae IV-VI terminal, well developed, seta VII minute, located at inner distal corner.
</paragraph>
<caption id="80FFB195FF8CB2B9E7BC21E40BBEF0A6" doi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" start="Figure 4" startId="F4">
<paragraph id="9312F5B03C799467E5A4180F8172B435" pageId="0" pageNumber="91">
<emphasis id="879A795DE22534854464AD6E31AC534F" bold="true" pageId="0" pageNumber="91">Figure 4.</emphasis>
<taxonomicName id="A5CFF48233E0AF0056B1F6AA9C5FCC84" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus kajii" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="kajii">
<emphasis id="84DD68129B5B5E08357274E51A6A09FE" italics="true" pageId="0" pageNumber="91">Caligus kajii</emphasis>
</taxonomicName>
sp. nov., adult female, paratype
<emphasis id="CBC2C5D471D1D6FC6B81C376B794FF36" bold="true" pageId="0" pageNumber="91">A</emphasis>
habitus, dorsal view
<emphasis id="B1606C4AFF5D6B4D5876BE3E2A032CCD" bold="true" pageId="0" pageNumber="91">B</emphasis>
posterior end of genital complex plus abdomen, ventral view
<emphasis id="CC6DF73E07447A769A5DFA426DDE9C15" bold="true" pageId="0" pageNumber="91">C</emphasis>
antennule
<emphasis id="B34E9C6E5E69466796F29BF00848179F" bold="true" pageId="0" pageNumber="91">D</emphasis>
antenna
<emphasis id="1C0154CEC7A9903148C59526855C04FA" bold="true" pageId="0" pageNumber="91">E</emphasis>
postantennal process
<emphasis id="AA2D75E8FEEF6C5AB615861785C5F110" bold="true" pageId="0" pageNumber="91">F</emphasis>
mandible
<emphasis id="EEFA484B789F18B9E01053D8C2C851D8" bold="true" pageId="0" pageNumber="91">G</emphasis>
maxillule
<emphasis id="50E7E7E5D10083C7B3D165F5DC63F2F9" bold="true" pageId="0" pageNumber="91">H</emphasis>
maxilla
<emphasis id="4A4BAE79F9807AAB82ED63FEBB9EDC3D" bold="true" pageId="0" pageNumber="91">I</emphasis>
maxilliped
<emphasis id="ED4083994BFA2075CD66DAFCD142C1E5" bold="true" pageId="0" pageNumber="91">J</emphasis>
sternal furca.
</paragraph>
</caption>
<paragraph id="E415B0CACE5B6643B34F6A207F74383A" pageId="0" pageNumber="91">
Antennule (
<figureCitation id="6D99441BF697ABDE81EEF3B14378523C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4C</figureCitation>
) 2-segmented; proximal segment bearing 26 setae on anteroventral surface; distal segment with 1 subterminal and 11 terminal setae and 2 short aesthetascs. Antenna (
<figureCitation id="BFC16747E5CD13AE4AF151433AB952D8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4D</figureCitation>
) 3-segmented, heavily sclerotized; proximal segment with acutely pointed posterior process; middle segment subrectangular, unarmed; distal segment abruptly curved inward at distal quarter, armed with proximal seta and minute middle seta. Postantennal process (
<figureCitation id="A51C6E4794B5EA15BA09A7AF74CBE4F9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4E</figureCitation>
) as long as distal segment of antenna, smoothly curved inward, with 2 multisensillate papillae basally; similar multisensillate papilla located adjacent to base of process. Mandible (
<figureCitation id="CE5FBDA7E60F60A33B10433F2312B06D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4F</figureCitation>
) with distal part bearing 12 teeth. Maxillule (
<figureCitation id="5C2C77D8EC9DA9EBC05509BCDBEAF8EB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4G</figureCitation>
) consisting of anterior papilla bearing 3 setae of unequal length and triangular, plate-like posterior process. Maxilla (
<figureCitation id="CD6016D9CFCDC6BD3B69BF9D04FAF6D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4H</figureCitation>
) 2-segmented, lacertus (syncoxa) unarmed; brachium (basis) ca. 1.5 times longer than lacertus, with flabellum at about anterior one-third of length;
<taxonomicName id="94F46FF9325F2EB18C333B3224F19F97" lsidName="calamus" pageId="0" pageNumber="91" rank="species" species="calamus">calamus</taxonomicName>
about 1.5 times longer than canna. Maxilliped (
<figureCitation id="62A90BDE71444C6F1B6A3E74A89D2646" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4I</figureCitation>
) heavily sclerotized; corpus (protopod) elongate, about 1.7 times as long as subchela, with low proximal lobe located at about 30% along myxal margin, plus minute setule in distal quarter of corpus; shaft (endopod) and claw incompletely fused; claw with antero-proximal barb reaching beyond mid-length. Sternal furca (
<figureCitation id="D05EC9E2D456913939D6517FC77CC990" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4J</figureCitation>
) with tines widely separated at base and bluntly pointed.
</paragraph>
<paragraph id="2F7372ED23F9084465AC021B8AFA45AB" pageId="0" pageNumber="91">
Armature and elements of legs 1-4 as in Table
<tableCitation id="8A0A0E532C7C5ACF5A4146C830B95A47" captionStartId="T2" httpUri="http://table.plazi.org/id/28BD004225F4678E2FF3806667A522D0" pageId="0" pageNumber="91" tableUuid="28BD004225F4678E2FF3806667A522D0">2</tableCitation>
.
<emphasis id="FD1458665159F021A17DF14096629E1D" bold="true" italics="true" pageId="0" pageNumber="91">Leg 1</emphasis>
(
<figureCitation id="579BE032885AEBA5DEE12D8177411BE9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">
Fig. 5
<normalizedToken id="459AA40D7879B25A31CBA30DA82AE6EC" originalValue="A–C">A-C</normalizedToken>
</figureCitation>
) with protopod bearing 1 marginal bifurcate setule and 2 surface setae; endopod reduced to small knob with vestigial element at tip (
<figureCitation id="9FA209E896E8F21DC64BB8BB5F79F182" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5B</figureCitation>
); exopod 2-segmented, proximal segment with row of setules along inner margin and outer distal seta, distal segment with 3 plumose setae along inner margin and 1 naked outer spine, 2 terminal spines each with accessory process (
<figureCitation id="E87F575EB6F5E77A4DC51828658DC1F5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5C</figureCitation>
), plus long spinulose seta (seta 4) terminally.
<emphasis id="988474CF8298738EC75F9CCBFEE8154D" bold="true" italics="true" pageId="0" pageNumber="91">Leg 2</emphasis>
(
<figureCitation id="6FF3D694C5D7C1BA35CA36E498EB59F8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5D, E</figureCitation>
) with intercoxal sclerite bearing semi-circular marginal membrane along posterior margin; coxa with large plumose seta at posterior corner and setule on anterior surface; basis ornamented with marginal membrane along both inner and outer edges and long setule near midpoint of inner margin; armed with minute seta at distal outer corner; rami 3-segmented; first endopod segment with notch bearing tuft of setules, second segment furnished with row of setules along outer margin, third segment with tuft of setules near base of proximal outer seta; first exopod segment with anterior marginal membrane reflexed dorsally over segment surface and long, stout outer spine directed obliquely across surface of second segment; second segment with smaller outer spine than in first segment; third segment armed with 1 reduced spine (arrowed in
<figureCitation id="219E5F88C72D77BF9E0B3CCEB04EB16D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5E</figureCitation>
) and 1 moderate-size outer spine.
<emphasis id="6C25FE7A63DF6F41DB248766DAF55153" bold="true" italics="true" pageId="0" pageNumber="91">Leg 3</emphasis>
(
<figureCitation id="EB706983ECA8832326552957A928A2CA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">
Fig. 5
<normalizedToken id="BB90B77499234F9AFB7E68D59065A42C" originalValue="F–H">F-H</normalizedToken>
</figureCitation>
) apron (protopod) with no distinct ornamentation on surface, armed with 1 long inner seta and 1 small outer seta terminally; endopod 2-segmented, proximal segment reduced, velum developed, decorated entirely with row of setules along free posterior margin; distal segment with outer margin expanded and hirsute
<figureCitation id="4F5D0E5C66BEC3255CFF07885A3280C9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5G</figureCitation>
; exopod 3-segmented, first segment with strong, slightly curved, inward-directed outer spine not reaching distal margin of second segment, second segment with expanded, hirsute outer margin and 1 minute outer seta (arrowed in
<figureCitation id="0B58A0ABFF382DA4E3DD70410A22C777" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5H</figureCitation>
), third segment with 3 small spines plus 4 inner setae.
</paragraph>
<caption id="02364AB2F7152DC066972982D89E4340" doi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" start="Figure 5" startId="F5">
<paragraph id="76BB94E64F1C7037A807A1AC8DBE2831" pageId="0" pageNumber="91">
<emphasis id="C9CBE7FF4BAEB0A1ABBFBB96F30EEAF9" bold="true" pageId="0" pageNumber="91">Figure 5.</emphasis>
<taxonomicName id="50556CA2FACD3A8326D4A0BC23A4A3CE" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus kajii" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="kajii">
<emphasis id="26E6448BBDB0BE3445DCC249FC99C5C3" italics="true" pageId="0" pageNumber="91">Caligus kajii</emphasis>
</taxonomicName>
sp. nov., adult female, paratype
<emphasis id="CEB0D451C456A0F2B62629A437972922" bold="true" pageId="0" pageNumber="91">A</emphasis>
leg 1
<emphasis id="628E938C96CEBC222A129F6EDEF1A432" bold="true" pageId="0" pageNumber="91">B</emphasis>
endopod of leg 1
<emphasis id="07790DBB97EA330E5D2138E82A5D4D75" bold="true" pageId="0" pageNumber="91">C</emphasis>
two terminal spines of second exopod segment of leg 1
<emphasis id="989736FBC48EC03E6FF6DC27D7BD26D0" bold="true" pageId="0" pageNumber="91">D</emphasis>
leg 2
<emphasis id="D587F42DC2852DFF259A206E2FBE1567" bold="true" pageId="0" pageNumber="91">E</emphasis>
outer spines of terminal exopod segment of leg 2, reduced spine arrowed
<emphasis id="FE9EE50C269A29EEE8701F4D3E45CF0E" bold="true" pageId="0" pageNumber="91">F</emphasis>
leg 3
<emphasis id="B57BA60AFF659F86EB9E698B3F246206" bold="true" pageId="0" pageNumber="91">G</emphasis>
endopod of leg 3
<emphasis id="933E75C9536B809B5D59A065F5C473B0" bold="true" pageId="0" pageNumber="91">H</emphasis>
exopod of leg 3
<emphasis id="FDF8FD6FB3780808FAC8FAEDFCCB8BF1" bold="true" pageId="0" pageNumber="91">I</emphasis>
leg 4
<emphasis id="A724F6347C481004FA796E8CF90AE481" bold="true" pageId="0" pageNumber="91">J</emphasis>
terminal part of leg 4.
</paragraph>
</caption>
<paragraph id="C53989D3E4F10AD62E864902012B3C6E" pageId="0" pageNumber="91">
<emphasis id="8AAB96CB81A689EF73F4C95479DD216D" bold="true" italics="true" pageId="0" pageNumber="91">Leg 4</emphasis>
(
<figureCitation id="F08E79F5BEAEFB35F4B54FA05B826593" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Caligus kajii sp. nov., adult female, paratype A leg 1 B endopod of leg 1 C two terminal spines of second exopod segment of leg 1 D leg 2 E outer spines of terminal exopod segment of leg 2, reduced spine arrowed F leg 3 G endopod of leg 3 H exopod of leg 3 I leg 4 J terminal part of leg 4." figureDoi="10.3897/zookeys.893.46923.figure5" httpUri="https://binary.pensoft.net/fig/361323" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure5">Fig. 5I, J</figureCitation>
) protopod slightly shorter than exopod, bearing minute plumose seta at outer distal corner; exopod 2-segmented, with pecten at base of each exopodal spine; first exopodal segment bearing spinulose spine reaching nearly half of distance to proximalmost outer spine on second segment; innermost terminal spine more than 3 times longer than middle spine. Leg 5 (
<figureCitation id="42E5E5C71DDE3B432E3822AB1EAA5B3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Caligus kajii sp. nov., adult female, paratype A habitus, dorsal view B posterior end of genital complex plus abdomen, ventral view C antennule D antenna E postantennal process F mandible G maxillule H maxilla I maxilliped J sternal furca." figureDoi="10.3897/zookeys.893.46923.figure4" httpUri="https://binary.pensoft.net/fig/361322" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure4">Fig. 4B</figureCitation>
) represented by small outer knob bearing protopodal seta and inner (exopodal) knob bearing 2 setae.
</paragraph>
<paragraph id="0E4C5DC3A3A9A9E2432A4C8203C95C8C" pageId="0" pageNumber="91">
<emphasis id="987D3CD7E6376659E9DC87F9333D6014" bold="true" pageId="0" pageNumber="91">Male.</emphasis>
Body length of allotype 4.36 mm, range 4.09-5.73 mm long in allotype plus all male paratypes (4.69
<normalizedToken id="A39385D1E5B231035C4F5023898C27F6" originalValue="±">+/-</normalizedToken>
0.51 mm long,
<emphasis id="847B831B9C058F12352CDE1294A03309" italics="true" pageId="0" pageNumber="91">N</emphasis>
= 14). Cephalothorax (
<figureCitation id="39C34EC883EDFD4CD2812221DAE5AB04" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6A</figureCitation>
) as in female. Pediger 4 (
<figureCitation id="7B1712BF12897C31E0120F861FB2DA35" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6A</figureCitation>
) separate from genital complex, wider than long. Genital complex (
<figureCitation id="A9A2A28D00F4F893AE629962EF297D97" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6A, B</figureCitation>
) about 1.4 times wider than long and about 1.2 times longer than abdominal somites combined, expanded posterolaterally into 2 knobs (leg 5), outer knob bearing 1 seta, inner (exopodal) knob with 2 setae; paired genital opercula representing leg 6 (
<figureCitation id="5F8BD31806E72F4DF6DD8BB2D28D8F65" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6B</figureCitation>
), each bearing 1 small terminal seta. Abdomen (
<figureCitation id="2E33842C986B98FA7379418E69FB8BAA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6A</figureCitation>
) 2-segmented, first segment small, second segment ca. 4.8 times longer than first. Caudal ramus (
<figureCitation id="7C4B0255F384DF987BC30D0788378FE7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6C</figureCitation>
) as in female.
</paragraph>
<caption id="AB4E886F00939FCBD21908F07F3F040D" doi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" start="Figure 6" startId="F6">
<paragraph id="73B2C581B5E3C9635F277DEC96290019" pageId="0" pageNumber="91">
<emphasis id="70D161D2DC3C6E2C87C6633740F95191" bold="true" pageId="0" pageNumber="91">Figure 6.</emphasis>
<taxonomicName id="88D5C9FD1F634709AFB81019AAC3ADE4" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus kajii" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="kajii">
<emphasis id="BFD871159F094A090C3D3A4F9AAB53EE" italics="true" pageId="0" pageNumber="91">Caligus kajii</emphasis>
</taxonomicName>
sp. nov., adult male, allotype
<emphasis id="DE24D8079B829C4D578F452C1050FA77" bold="true" pageId="0" pageNumber="91">A</emphasis>
habitus, dorsal view
<emphasis id="0A3979711F167AC01664566846789F3A" bold="true" pageId="0" pageNumber="91">B</emphasis>
gonopores, ventral view
<emphasis id="4F4E21951ED2D32F9B6AAF401BAB116C" bold="true" pageId="0" pageNumber="91">C</emphasis>
tip of caudal ramus, dorsal view
<emphasis id="86FA690EE6527BD6D300526C31795A0F" bold="true" pageId="0" pageNumber="91">D</emphasis>
antenna
<emphasis id="BEFA811030B084A6A021324F99491774" bold="true" pageId="0" pageNumber="91">E</emphasis>
postantennal process
<emphasis id="C52FAA444E24A600A63647158C0370C6" bold="true" pageId="0" pageNumber="91">F</emphasis>
maxillule
<emphasis id="7523C3FBB231EFEB916A1C8A19B8AFBF" bold="true" pageId="0" pageNumber="91">G</emphasis>
maxilliped, rounded knob on posterior surface of corpus arrowed
<emphasis id="70191C65F6C50B8CC5B9A0FB01BFF3FA" bold="true" pageId="0" pageNumber="91">H</emphasis>
sternal furca
<emphasis id="390E55A2CAEB1DCB71181DB2B217C461" bold="true" pageId="0" pageNumber="91">I</emphasis>
leg 4.
</paragraph>
</caption>
<paragraph id="C62506F4B5F47F6A598108CEC3027CA7" pageId="0" pageNumber="91">
Antennule, mandible, maxillule (
<figureCitation id="5B34C8A240CB373249798A302BB0FB14" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6F</figureCitation>
) and maxilla as in female. Antenna (
<figureCitation id="AC9F876847EB86B0CD9D1264E5904AF7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6D</figureCitation>
) 3-segmented; proximal segment long, unarmed, with corrugated pad distally; middle segment massive, with 1 proximal, 2 middle and 2 subterminal corrugated pads; distal segment short, with 2 fine setae at mid-length, 1 bluntly pointed process terminally and 1 membranous truncate projection subterminally. Postantennal process (
<figureCitation id="316D94F078FC8DA63F19C4BA800533C1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6E</figureCitation>
) similar to that of female, but more abruptly curved inward than in female. Maxilliped (
<figureCitation id="5ADF2D1B4BCB744CCBDDD8B3F4A02F15" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6G</figureCitation>
) stouter than in female; corpus with rounded knob proximally on posterior surface (arrowhead in
<figureCitation id="F6FE250035FD72771587301303E1FF4A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Caligus kajii sp. nov., adult male, allotype A habitus, dorsal view B gonopores, ventral view C tip of caudal ramus, dorsal view D antenna E postantennal process F maxillule G maxilliped, rounded knob on posterior surface of corpus arrowed H sternal furca I leg 4." figureDoi="10.3897/zookeys.893.46923.figure6" httpUri="https://binary.pensoft.net/fig/361324" pageId="0" pageNumber="91" tableDoi="10.3897/zookeys.893.46923.figure6">Fig. 6G</figureCitation>
) and 3 distinct processes in myxal area, proximalmost process socket-like, to receive distal tip of subchela; shaft and claw partly fused, ca. 0.67 times as long as corpus; barb about half length of claw.
</paragraph>
<caption id="28BD004225F4678E2FF3806667A522D0" ID-Table-UUID="28BD004225F4678E2FF3806667A522D0" httpUri="http://table.plazi.org/id/28BD004225F4678E2FF3806667A522D0" pageId="0" pageNumber="91" start="Table 2" startId="T2">
<paragraph id="2E41D321173FC77EFA1EB5A63F2EE83E" pageId="0" pageNumber="91">
<emphasis id="8B3945E5DA6D56F6EEE2FB7F20CC56BE" bold="true" pageId="0" pageNumber="91">Table 2.</emphasis>
Armature and elements of legs 1 to 4 of
<taxonomicName id="DB70287DAE1AE0600DF6DA5A4EFC7883" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus kajii" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="kajii">
<emphasis id="99EDDE5E1901E6187C897FDCA088882D" italics="true" pageId="0" pageNumber="91">Caligus kajii</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph id="5077EB80FF2C3B7938B4AD9DC9849B7E" pageId="0" pageNumber="91">
<table id="F50176402FCA45FEDDA8BA96A2DF76B8" pageId="0" pageNumber="91">
<tr id="8128D4E72C7AFAD6479E351C834A1E50" pageId="0" pageNumber="91">
<th id="5B092A09508B5D7B6C18075D3ACF3687" colspan="1" isEmpty="true" pageId="0" pageNumber="91" rowspan="1">-</th>
<th id="98D05569EFFB362C526BA4492033BB15" colspan="1" pageId="0" pageNumber="91" rowspan="1">Protopod (coxa; basis)</th>
<th id="9635065F01EC5537D95E3E8E23A55BCD" colspan="1" pageId="0" pageNumber="91" rowspan="1">Endopod</th>
<th id="9AA4B2C1F0A41AAE6014AC35F2DB3D80" colspan="1" pageId="0" pageNumber="91" rowspan="1">Exopod</th>
</tr>
<tr id="3AEC382B5F06529BC42F6447175F612C" pageId="0" pageNumber="91">
<td id="EA3A9D34EF3173E726509B6A5C828C31" colspan="1" pageId="0" pageNumber="91" rowspan="1">Leg 1</td>
<td id="99A3B478D6DBEEDFB76F360931C3F41A" colspan="1" pageId="0" pageNumber="91" rowspan="1">1-1</td>
<td id="1DD6F6AB405E1D9DEDC0CDC4D49FE9DF" colspan="1" pageId="0" pageNumber="91" rowspan="1">(vestigial)</td>
<td id="B2C8A410B17FB5C3E40B373BAB10A1AE" colspan="1" pageId="0" pageNumber="91" rowspan="1">1-0; III, 1, 3</td>
</tr>
<tr id="5D6A2109F368504F52B3CC835E44FDA1" pageId="0" pageNumber="91">
<td id="1BC18CA9416AB97485DC3FBAF347F2F2" colspan="1" pageId="0" pageNumber="91" rowspan="1">Leg 2</td>
<td id="CBE2C36BD66E3FC2F060E09D55A0BA89" colspan="1" pageId="0" pageNumber="91" rowspan="1">0-1; 1-0</td>
<td id="72BD5B834B3127037F870F1C86297F06" colspan="1" pageId="0" pageNumber="91" rowspan="1">0-1; 0-2; 6</td>
<td id="13AFAB6599B9B8A19FB10C36E6A6A391" colspan="1" pageId="0" pageNumber="91" rowspan="1">I-1; I-1; II, I,5</td>
</tr>
<tr id="9A3F967C04982871F66779ACE8CD101B" pageId="0" pageNumber="91">
<td id="6F96AF2D92E7AA363289A054BDDA28DE" colspan="1" pageId="0" pageNumber="91" rowspan="1">Leg 3</td>
<td id="357338984C16A14CF779041481DD383D" colspan="1" pageId="0" pageNumber="91" rowspan="1">1-1</td>
<td id="C8C8B5BFE0FAC931D495E3CAC4451B5A" colspan="1" pageId="0" pageNumber="91" rowspan="1">0-1; 6</td>
<td id="1E40FEFCF9B292EA6E4902BD3D7CF2F9" colspan="1" pageId="0" pageNumber="91" rowspan="1">I-0; 1(vestigial)-1; III, 4</td>
</tr>
<tr id="C33B49F0F24BB8867A998049F781739A" pageId="0" pageNumber="91">
<td id="CE292B5B6A483529F6E891E42DCA30CB" colspan="1" pageId="0" pageNumber="91" rowspan="1">Leg 4</td>
<td id="D1405D79AD616562094DAAB7AEF7E921" colspan="1" pageId="0" pageNumber="91" rowspan="1">1-0</td>
<td id="F2E700DEA63410CE53E0FB62990C696B" colspan="1" pageId="0" pageNumber="91" rowspan="1">(absent)</td>
<td id="A76B20F2BDAD832D1CAC3BADDF3B16F1" colspan="1" pageId="0" pageNumber="91" rowspan="1">1-0; I, III</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="A7CC8077C45FA87891365F2B267F91E9" pageId="0" pageNumber="91" type="remarks">
<paragraph id="A9B515C6C52654A598DB2D2D2C8A2A70" pageId="0" pageNumber="91">Remarks.</paragraph>
<paragraph id="0CED29AD6AAF7D1D1D63B3771AC513A0" pageId="0" pageNumber="91">
The female of the new species most closely resembles
<taxonomicName id="3506C4DD8CDA7B27615B71AFD4FDA0F4" class="Copepoda" family="Caligidae" genus="Caligus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caligus bifurcus" order="Siphonostomatoida" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bifurcus">
<emphasis id="2AE1BF1573D12F0D6D87328FDD6C59A7" italics="true" pageId="0" pageNumber="91">Caligus bifurcus</emphasis>
</taxonomicName>
,
<taxonomicName id="D150C6760E7AD0EC2A104FAB17364AC0" lsidName="C. musaicus" pageId="0" pageNumber="91" rank="species" species="musaicus">
<emphasis id="E34B2A4DEB5281EA969113E9486F6F2D" italics="true" pageId="0" pageNumber="91">C. musaicus</emphasis>
</taxonomicName>
Cavaleiro, Santos &amp; Ho, 2010,
<taxonomicName id="930431C489A9BFFE77068E15519463F6" lsidName="C. pectinatus" pageId="0" pageNumber="91" rank="species" species="pectinatus">
<emphasis id="171A5E08ADC03BE54AC354A40FDDAEB7" italics="true" pageId="0" pageNumber="91">C. pectinatus</emphasis>
</taxonomicName>
Shiino, 1965,
<taxonomicName id="8E704C936AD984D3BF134BE869771E6A" lsidName="C. pseudorhombi" pageId="0" pageNumber="91" rank="species" species="pseudorhombi">
<emphasis id="3859429B8BC3B5F2DC96F3A630D05F5A" italics="true" pageId="0" pageNumber="91">C. pseudorhombi</emphasis>
</taxonomicName>
Boxshall, 2018,
<taxonomicName id="09E94D9E59E81AD5876A1A7328B9337B" lsidName="C. pterois" pageId="0" pageNumber="91" rank="species" species="pterois">
<emphasis id="307AA1A5EA3F455934FB0ADB73209921" italics="true" pageId="0" pageNumber="91">C. pterois</emphasis>
</taxonomicName>
Kurian, 1949 and
<taxonomicName id="22D8D13DA281BA1F07C7D29C8A2E4367" lsidName="C. xystercus" pageId="0" pageNumber="91" rank="species" species="xystercus">
<emphasis id="FD2B97834876E4282B4AAD3FEFA8BB2E" italics="true" pageId="0" pageNumber="91">C. xystercus</emphasis>
</taxonomicName>
Cressey, 1991. All these species share a 2-segmented exopod on leg 4 armed with 4 spines on the distal exopodal segment, the female genital complex is nearly as long as wide and about twice as long as the abdomen, and the abdomen is about as long as wide. However, the present new species is distinguished from these species by the combination of the following characteristics: (1) the genital complex is as long as wide (cf. wider than long in
<taxonomicName id="63D4EB0D757DBA37CFA6D7306F29E9A9" lsidName="C. bifurcus" pageId="0" pageNumber="91" rank="species" species="bifurcus">
<emphasis id="1E2501B17F249EB3BE3491E8DF8BF6CE" italics="true" pageId="0" pageNumber="91">C. bifurcus</emphasis>
</taxonomicName>
,
<taxonomicName id="CBAC5499834859C5EC7044283DFC0651" lsidName="C. musaicus" pageId="0" pageNumber="91" rank="species" species="musaicus">
<emphasis id="54046A3A38AB460BADB40C1034706F95" italics="true" pageId="0" pageNumber="91">C. musaicus</emphasis>
</taxonomicName>
and
<taxonomicName id="C57D39066F3FEB5410B38AF8E97BBC5E" lsidName="C. pterois" pageId="0" pageNumber="91" rank="species" species="pterois">
<emphasis id="3011F01E222DE77AF604D01E79C77078" italics="true" pageId="0" pageNumber="91">C. pterois</emphasis>
</taxonomicName>
; slightly longer than wide in
<taxonomicName id="91C61DBC9C16E7023C4EFBD4E17BA51B" lsidName="C. xystercus" pageId="0" pageNumber="91" rank="species" species="xystercus">
<emphasis id="F51BACBE55B088F9AB8C05B27F0C35E7" italics="true" pageId="0" pageNumber="91">C. xystercus</emphasis>
</taxonomicName>
); (2) the genital complex is about 2.1 times longer than the abdomen (cf. 1.2 times longer in
<taxonomicName id="A0756811D138614265A8BBFF6C59B873" lsidName="C. bifurcus" pageId="0" pageNumber="91" rank="species" species="bifurcus">
<emphasis id="494316477D41AAB0B7D626AD4B07CCE9" italics="true" pageId="0" pageNumber="91">C. bifurcus</emphasis>
</taxonomicName>
; 2.2 in
<taxonomicName id="5BFBFFF139296694A3DEA911E9786A51" lsidName="C. musaicus" pageId="0" pageNumber="91" rank="species" species="musaicus">
<emphasis id="38D4DF192F180D6BF77F2B3ED8118731" italics="true" pageId="0" pageNumber="91">C. musaicus</emphasis>
</taxonomicName>
; 2.1 in
<taxonomicName id="E5628F8A38E6BC3C59F8A9417CC57CF8" lsidName="C. pseudorhombi" pageId="0" pageNumber="91" rank="species" species="pseudorhombi">
<emphasis id="0A10D23402B43E17392A82E81950843E" italics="true" pageId="0" pageNumber="91">C. pseudorhombi</emphasis>
</taxonomicName>
; 1.6 in
<taxonomicName id="4157DFF5A04C8602E319E8B4A338085A" lsidName="C. pterois" pageId="0" pageNumber="91" rank="species" species="pterois">
<emphasis id="62052CA1DF04663A9D715A8EC901CF70" italics="true" pageId="0" pageNumber="91">C. pterois</emphasis>
</taxonomicName>
; 3.6 in
<taxonomicName id="7645D2E0B37550E33FB283C130573677" lsidName="C. xystercus" pageId="0" pageNumber="91" rank="species" species="xystercus">
<emphasis id="3A5EF3DA0182E16ED7D5AF7E7FB39325" italics="true" pageId="0" pageNumber="91">C. xystercus</emphasis>
</taxonomicName>
); (3) the corpus of the maxilliped lacks processes (cf. ridge-like process present in
<taxonomicName id="1B5F79388D1F02D643B31E3E34B634B4" lsidName="C. pseudorhombi" pageId="0" pageNumber="91" rank="species" species="pseudorhombi">
<emphasis id="2835C9363E5C17DAD0B5AF79948EE0D2" italics="true" pageId="0" pageNumber="91">C. pseudorhombi</emphasis>
</taxonomicName>
and
<taxonomicName id="D8B7C6D45A082F7DC5C0E7C84D7DDA35" lsidName="C. pterois" pageId="0" pageNumber="91" rank="species" species="pterois">
<emphasis id="73E4BA54F9921DAF283DE5BE46908E1D" italics="true" pageId="0" pageNumber="91">C. pterois</emphasis>
</taxonomicName>
); (4) the tines of the sternal furca taper distally (cf. uniform in width and with a truncate tip in
<taxonomicName id="584F631EED5551E24F5F8D12DCE48338" lsidName="C. pectinatus" pageId="0" pageNumber="91" rank="species" species="pectinatus">
<emphasis id="81D98E845917979D1A58B516AED6A6F6" italics="true" pageId="0" pageNumber="91">C. pectinatus</emphasis>
</taxonomicName>
); (5) the terminal exopod segment of leg 1 is furnished with 3 large spines terminally (2 in
<taxonomicName id="282916BD2DF26B33CE2B9628ABBCA16E" lsidName="C. pseudorhombi" pageId="0" pageNumber="91" rank="species" species="pseudorhombi">
<emphasis id="0C67112E2FF6645343A19ED9FBD5F4B5" italics="true" pageId="0" pageNumber="91">C. pseudorhombi</emphasis>
</taxonomicName>
); and (6) the maxillipedal subchela is more than half the length of the corpus (cf. much shorter in
<taxonomicName id="0D5C563FDCF2867AC609B1BE73E98156" lsidName="C. musaicus" pageId="0" pageNumber="91" rank="species" species="musaicus">
<emphasis id="AE98D94122728AF3C757D6FC75A61A2E" italics="true" pageId="0" pageNumber="91">C. musaicus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3FFE0D9B8329EE90564FA51ACB3EC4CF" pageId="0" pageNumber="91">
In males, the new species is most similar to
<taxonomicName id="B91FDC9CD74F9483348547D821D05E8A" lsidName="C. musaicus" pageId="0" pageNumber="91" rank="species" species="musaicus">
<emphasis id="482369C334F7A15684566D78D3EDC497" italics="true" pageId="0" pageNumber="91">C. musaicus</emphasis>
</taxonomicName>
,
<taxonomicName id="D711F5361C0C535485324CCD584EEED4" lsidName="C. nuenonnae" pageId="0" pageNumber="91" rank="species" species="nuenonnae">
<emphasis id="48F3B1A3E28C79B9D41CDB352B631159" italics="true" pageId="0" pageNumber="91">C. nuenonnae</emphasis>
</taxonomicName>
Andrews, Bott, Battaglene &amp; Nowak, 2009,
<taxonomicName id="9BF6BE591A607FF90DFBAAD80B4E7772" lsidName="C. pterois" pageId="0" pageNumber="91" rank="species" species="pterois">
<emphasis id="56888821D58FC09CB341A359E08276A1" italics="true" pageId="0" pageNumber="91">C. pterois</emphasis>
</taxonomicName>
, and
<taxonomicName id="E2C9BD18BE82FFDEB96151378B320BC5" lsidName="C. priacanthi" pageId="0" pageNumber="91" rank="species" species="priacanthi">
<emphasis id="A4FA400A6B8C31BF88BD04148EA505A6" italics="true" pageId="0" pageNumber="91">C. priacanthi</emphasis>
</taxonomicName>
Pillai, 1962. These five species share the following characteristics: (1) the genital complex is laterally expanded and produced into 2 posterolateral protuberances representing leg 5, armed with 1 (outer lobe) and 2 setae (inner lobe); (2) the abdomen is completely or incompletely 2-segmented and shorter than the genital complex; (3) the maxillipedal corpus is well developed and carries anteriorly-produced processes on the myxal surface. However, the new species is easily distinguishable from these congeners by the combination of the following features: (1) the maxillipedal corpus has a rounded process on the posterior surface (absent in the other species); (2) the sternal furca has pointed tines which are widely separated (tines that are close at base and with rounded tips in
<taxonomicName id="42999D349F7DC9E3471B685B3A46B29E" lsidName="C. priacanthi" pageId="0" pageNumber="91" rank="species" species="priacanthi">
<emphasis id="5DD0C0667325105F5BC83DB19DC32033" italics="true" pageId="0" pageNumber="91">C. priacanthi</emphasis>
</taxonomicName>
; rounded in
<taxonomicName id="D7E794FFC5D205561888CA74DB823AD7" lsidName="C. nuenonnae" pageId="0" pageNumber="91" rank="species" species="nuenonnae">
<emphasis id="190E3D22886D88F18C0FEBDDC2FAFBF9" italics="true" pageId="0" pageNumber="91">C. nuenonnae</emphasis>
</taxonomicName>
); (3) the mxyal surface of the maxilliped carries 3 large, rounded processes along the margin, (cf. the processes are different in shape and number in the other species); (4) the posterior dentiform process of the maxillule lacks a surface ornamentation of minute prominences (present in
<taxonomicName id="1EED06506B172DF01E2288F76A4C64E9" lsidName="C. nuenonnae" pageId="0" pageNumber="91" rank="species" species="nuenonnae">
<emphasis id="474D8CB64A5377DE8AE4CBEFEA0BDAF0" italics="true" pageId="0" pageNumber="91">C. nuenonnae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="0CC69AD1E8FC153A1570703907E278AA" pageId="0" pageNumber="91" type="etymology">
<paragraph id="5EF9DC63329EBCCA65EEFF2AFB3AE18E" pageId="0" pageNumber="91">Etymology.</paragraph>
<paragraph id="97E06C9875490B254FB2D88C720EF69D" pageId="0" pageNumber="91">The new species is named in honor of the late, supremely talented carcinologist Tomonari Kaji who passed away in May 2019.</paragraph>
</subSubSection>
</treatment>
</document>
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