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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.564.6819" ID-GBIF-Dataset="2b9eca3b-58b9-4499-bbb4-64b8881e6a3f" ID-PMC="PMC4820093" ID-Pensoft-Pub="1313-2970-564-121" ID-PubMed="27081334" ID-ZBK="8D6563D67C4F4435BE6C19CCE2F9882F" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-564-121" ModsDocOrigin="ZooKeys 564" ModsDocTitle="Contributions to the knowledge of subterranean trechine beetles in southern Chinas karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)" checkinTime="1455646106532" checkinUser="pensoft" docAuthor="Tian, Mingyi, Huang, Sunbin, Wang, Xinhui &amp; Tang, Mingruo" docDate="2016" docId="147B9B414E2CA6FC065C7DE39A660FCB" docLanguage="en" docName="ZooKeys 564: 121-156" docOrigin="ZooKeys 564" docSource="http://dx.doi.org/10.3897/zookeys.564.6819" docTitle="Shiqianaphaenops Tian, gen. n." docType="treatment" docUuid="D7A2BFB8-59A2-4BA4-A26C-630F9E5B7488" docUuidSource="ZooBank" docVersion="5" lastPageNumber="127" masterDocId="646BFFE3E466300A6C0AFFE5FFBA1245" masterDocTitle="Contributions to the knowledge of subterranean trechine beetles in southern China's karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)" masterLastPageNumber="156" masterPageNumber="121" pageNumber="125" updateTime="1668162679916" updateUser="ExternalLinkService">
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<mods:title>Contributions to the knowledge of subterranean trechine beetles in southern China's karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Huang, Sunbin</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wang, Xinhui</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tang, Mingruo</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>564</mods:number>
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<mods:start>121</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.564.6819</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.564.6819</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-564-121</mods:identifier>
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<treatment ID-GBIF-Taxon="127880888" LSID="urn:lsid:zoobank.org:act:D7A2BFB8-59A2-4BA4-A26C-630F9E5B7488" httpUri="http://treatment.plazi.org/id/147B9B414E2CA6FC065C7DE39A660FCB" lastPageId="6" lastPageNumber="127" pageId="4" pageNumber="125">
<subSubSection pageId="4" pageNumber="125" type="multiple">
<paragraph pageId="4" pageNumber="125">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="nomenclature">
<paragraph pageId="4" pageNumber="125">
<taxonomicName LSID="http://zoobank.org/D7A2BFB8-59A2-4BA4-A26C-630F9E5B7488" authority="Tian" class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">Shiqianaphaenops Tian</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="125">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="type species">
<paragraph pageId="4" pageNumber="125">Type species.</paragraph>
<paragraph pageId="4" pageNumber="125">
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops majusculus" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="majusculus">Shenaphaenops majusculus</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999 (Cave Feng Dong, Shiqian, Guizhou).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="diagnosis">
<paragraph pageId="4" pageNumber="125">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="125">Medium-sized aphaenopsian trechine beetles, with sparsely pubescent body, elongated head, reduced frontal furrows, tridentate right mandible, 4-setose mentum, tumid propleura, widened 1st protarsomere and bisetose on each of abdominal ventrites.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="125" type="generic characteristics">
<paragraph pageId="4" pageNumber="125">Generic characteristics.</paragraph>
<paragraph pageId="4" pageNumber="125">Medium-sized aphaenopsian trechines, yet not too highly modified morphologically; eyeless, unpigmented and apterous; body slender and elongate, with slender and long appendages; covered with sparse pubescence, hairs being much longer on head and pronotum than on elytra; head elongate, longer than prothorax, much longer than wide; frontal furrows short, with two pairs of supra-orbital pores; right mandible tridentate; labial suture clear; mentum 4-setose, tooth simple and short, blunt at apex; submentum 9-setose; antennae long, nearly extending to elytral apex; prothorax longer than wide, propleura visible from above; pronotum elongate, longer than wide, widest near front; only anterior pairs of lateromarginal setae; elytra elongate-ovate, strongly convex, shoulders distinct and rounded, lateral margins ciliated throughout; striae faintly impressed; two dorsal and the pre-apical pores present on each elytron; humeral group of marginal umbilicate pores not aggregated; protibia without longitudinal groove externally; 1st protarsomere in both sexes widened and angularly produced externally, with a row of comb-like setae on ventral side; 1st and 2nd protarsomeres modified in male; ventrite VII with two pairs of setae in both sexes.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125" type="remarks">
<paragraph pageId="4" pageNumber="125">Remarks.</paragraph>
<paragraph lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125">
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
(1999) hesitantly treated the two trechine species found in Shiqian County, eastern Guizhou as members of the genus
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999, which had been set up based on a single female collected in Shuicheng County, northwestern Guizhou. However, he realized some striking differences and the geographical gap between the Shiqian species and the type species
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999. Apart from the somewhat similar general body configurations in both
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
gen. n. (Fig. 1) and
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
(Fig. 2), many character states of generic importance are
<pageBreakToken pageId="5" pageNumber="126" start="start">different</pageBreakToken>
, such as: (1) 1st protarsomere in both sexes widened apically and distinctly produced externally, covered with a ctenidium structure ventrally in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
which, amongst trechines, is only found in this genus; (2) right mandible tridentate in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, versus bidentate in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (3) labial suture clear in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, but completely missing in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (4) ventrite VII in male bisetose in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, versus 4-setose in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (5) in male, both 1st and 2nd protarsomeres modified in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, versus only 1st protarsomere being modified in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (6) pubescence weaker and sparser, hairs much longer on head and pronotum than on elytra in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, versus generally denser and hairs being of same length in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (7) base of pronotum much narrower than front in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, versus only slightly narrower in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (8) elytra broader, with humeral angles rounded in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
(Fig. 3A), versus more slender, with humeral angles distinctly angular in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
(Fig. 3B); (9) marginal sides of elytra ciliate throughout, versus smooth in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
; (10) in
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shiqianaphaenops</taxonomicName>
, median lobe of aedeagus stout and thick, broadly rounded dorsally, basal part comparatively small, with an indistinct sagittal aileron, inner sac armed with a very large copulatory piece; apical lobe contracted at apex, roundly blunt in lateral view, but pointed in dorsal view; parameres thin, much shorter than median lobe, each bearing three long setae at apex (Fig. 4A, B). In contrast, median lobe of aedeagus in
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Shenaphaenops</taxonomicName>
slender and thin, gently expanded dorsally, basal part comparatively large, with a small but distinct sagittal aileron; apical lobe indistinctly contracted at apex, obtuse in lateral view, broad in dorsal view; inner sac armed with a large copulatory piece; parameres shorter than median lobe, left one slightly longer than right one, each bearing two long setae at apex (Fig. 4C, D).
</paragraph>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 1. Habitus of
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n., male Scale bar: 2.0 mm.
</paragraph>
</caption>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 2. Habitus of
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999, male Scale bar: 2.0 mm.
</paragraph>
</caption>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 3. Elytral chaetotaxy A
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n. B
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999 C
<taxonomicName class="Insecta" family="Carabidae" genus="Dianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dianotrechus gueorguievi" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="gueorguievi">Dianotrechus gueorguievi</taxonomicName>
Tian, gen. n., sp. n. D
<taxonomicName class="Insecta" family="Carabidae" genus="Tianeotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tianeotrechus trisetosus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="trisetosus">Tianeotrechus trisetosus</taxonomicName>
Tian &amp; Tang, gen. n., sp. n. E
<taxonomicName class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes tujiaphilus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="tujiaphilus">Huoyanodytes tujiaphilus</taxonomicName>
Tian &amp; Huang, gen. n., n. sp. F
<taxonomicName class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes tujiaphilus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="tujiaphilus">Huoyanodytes tujiaphilus</taxonomicName>
Tian &amp; Huang, gen. n., sp. n. G
<taxonomicName class="Insecta" family="Carabidae" genus="Wanhuaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wanhuaphaenops zhangi" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="zhangi">Wanhuaphaenops zhangi</taxonomicName>
Tian &amp; Wang, sp. n.
</paragraph>
</caption>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 4. Male genitalia A and C median lobe, lateral view B and D apical lobe, dorsal view A and B
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n. C and D
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="etymology">
<paragraph pageId="5" pageNumber="126">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="126">Shiqian+Aphaenops, to indicate that both known members of this genus occur in Shiqian County, eastern Guizhou Province. Gender masculine.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="range">
<paragraph pageId="5" pageNumber="126">Range.</paragraph>
<paragraph pageId="5" pageNumber="126">China (eastern Guizhou) (Fig. 5c).</paragraph>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 5. Distribution of the trechine beetles a
<taxonomicName class="Insecta" family="Carabidae" genus="Dianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dianotrechus gueorguievi" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="gueorguievi">Dianotrechus gueorguievi</taxonomicName>
Tian, gen. n., sp. n. b
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999 c
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n. d
<taxonomicName class="Insecta" family="Carabidae" genus="Tianeotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tianeotrechus trisetosus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="trisetosus">Tianeotrechus trisetosus</taxonomicName>
Tian &amp; Tang, gen. n., sp. n. e
<taxonomicName class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes tujiaphilus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="tujiaphilus">Huoyanodytes tujiaphilus</taxonomicName>
Tian &amp; Huang, sp. n. f
<taxonomicName class="Insecta" family="Carabidae" genus="Wanhuaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wanhuaphaenops zhangi" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="zhangi">Wanhuaphaenops zhangi</taxonomicName>
Tian &amp; Wang, gen. n., sp. n.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="126">
Known so far by two very similar species from two caves in Shiqian County: Shenxian Dong and Feng Dong. Since Shenxian Dong lies very close to Feng Dong, only about one kilometre in distance across a shallow valley, with still another cave just between them, all three caves may prove belong to a same cave system. Perhaps this is why both
<normalizedToken originalValue="Uénos">Ueno's</normalizedToken>
species from Shiqian County are not too different from each other. Feng Dong is a large and beautiful cave (Fig. 6A, B). The beetle (Fig. 6C) is sympatric with millipedes, crickets and frogs.
</paragraph>
<caption pageId="5" pageNumber="126">
<paragraph pageId="5" pageNumber="126">
Figure 6. Cave Feng Dong, type locality of
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n. A entrance B cave chamber, to show where the beetles were collected C a wandering individual in cave.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="127" pageId="5" pageNumber="126" type="materials_examined">
<paragraph pageId="5" pageNumber="126">Material examined.</paragraph>
<paragraph pageId="5" pageNumber="126">
<taxonomicName class="Insecta" family="Carabidae" genus="Shenaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shenaphaenops humeralis" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="humeralis">Shenaphaenops humeralis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999: a male, Cave Shendong Migong (Shendong in the original description), Muqiao, Laoyingshan, Shuicheng County, northwestern Guizhou,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="26.587502">26°35'15&quot;N</geoCoordinate>
/
<geoCoordinate direction="east" orientation="longitude" precision="15" value="104.99638">104°59'47&quot;E</geoCoordinate>
, 1910 m in altitude, VIII-22-2014, Mingyi Tian leg., deposited in the insect collections of South China Agricultural University (SCAU).
</paragraph>
<paragraph pageId="5" pageNumber="126">
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops majusculus" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="majusculus">Shiqianaphaenops majusculus</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n.: 3 males, V-1-2015, Cave Feng Dong, Tangshan, Shiqian County,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="27.48611">27°29'10&quot;N</geoCoordinate>
/
<geoCoordinate direction="east" orientation="longitude" precision="15" value="108.256386">108°15'23&quot;E</geoCoordinate>
, 700 m in altitude, Mingyi Tian &amp; Jingli Cheng leg., in SCAU; 6 males &amp; 4 females, ibid., VIII-1-2015, same collectors, in SCAU.
</paragraph>
<paragraph pageId="6" pageNumber="127">
<taxonomicName class="Insecta" family="Carabidae" genus="Shiqianaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shiqianaphaenops cursor" order="Coleoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cursor">
<pageBreakToken pageId="6" pageNumber="127" start="start">Shiqianaphaenops</pageBreakToken>
cursor
</taxonomicName>
(
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
, 1999), comb. n.: We have not seen any material. We visited Cave Shenxian Dong, the type locality, in August 2015, but failed to catch anything. The cave is badly disturbed by human activities.
</paragraph>
</subSubSection>
</treatment>
</document>