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<document ID-DOI="10.11646/zootaxa.5249.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7690993" ID-ZooBank="51EE6A05-41C8-44F7-90C9-B785D6788E1B" approvalRequired="5" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="3" checkinTime="1677744808497" checkinUser="plazi" docAuthor="Ghosh, Aarini, Jaiswara, Ranjana, Sabharwal, Shagun, Dasoju, Vivek, Bhattacharjee, Anubhab &amp; Rajaraman, Bittu Kaveri" docDate="2023" docId="1461A42815694960EDFAFC99FDFEFA41" docLanguage="en" docName="zootaxa.5249.3.2.pdf" docOrigin="Zootaxa 5249 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexacentrus khasiensis Ghosh, Jaiswara &amp; Rajaraman 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="342" masterDocId="E858DC50156D4967ED6DFFCEFF87FFB0" masterDocTitle="Description of three new species of predatory Genus Hexacentrus (Orthoptera Tettigoniidae) from India, with bioacoustic and morphological characterizations" masterLastPageNumber="356" masterPageNumber="335" pageNumber="339" updateTime="1677745245794" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Description of three new species of predatory Genus Hexacentrus (Orthoptera Tettigoniidae) from India, with bioacoustic and morphological characterizations</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ghosh, Aarini</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4705-9846</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Ashoka University, Sonipat, Haryana, 131029, India. &amp; https: // orcid. org / 0000 - 0003 - 4705 - 9846</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaiswara, Ranjana</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2672-3093</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Panjab University, Chandigarh, 160014, India. &amp; https: // orcid. org / 0000 - 0002 - 2672 - 3093</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sabharwal, Shagun</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8828-5781</mods:nameIdentifier>
<mods:affiliation>School of Biological Science, NISER, Orissa, 752050, India. &amp; https: // orcid. org / 0000 - 0001 - 8828 - 5781</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dasoju, Vivek</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3815-9255</mods:nameIdentifier>
<mods:affiliation>Department of Psychology, Ashoka University, Sonipat, Haryana, 131029, India. &amp; https: // orcid. org / 0000 - 0003 - 3815 - 9255</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bhattacharjee, Anubhab</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1088-7157</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Ashoka University, Sonipat, Haryana, 131029, India. &amp; https: // orcid. org / 0000 - 0002 - 1088 - 7157</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rajaraman, Bittu Kaveri</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2435-7015</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Ashoka University, Sonipat, Haryana, 131029, India. &amp; Department of Psychology, Ashoka University, Sonipat, Haryana, 131029, India. &amp; https: // orcid. org / 0000 - 0003 - 2435 - 7015</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5249</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>335</mods:start>
<mods:end>356</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5249.3.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5249.3.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7690993</mods:identifier>
<mods:identifier type="ZooBank">51EE6A05-41C8-44F7-90C9-B785D6788E1B</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7691001" ID-Zenodo-Dep="7691001" LSID="urn:lsid:plazi:treatment:1461A42815694960EDFAFC99FDFEFA41" httpUri="http://treatment.plazi.org/id/1461A42815694960EDFAFC99FDFEFA41" lastPageId="7" lastPageNumber="342" pageId="4" pageNumber="339">
<subSubSection box="[151,938,855,882]" pageId="4" pageNumber="339" type="nomenclature">
<paragraph blockId="4.[151,938,855,882]" box="[151,938,855,882]" pageId="4" pageNumber="339">
<heading bold="true" box="[151,938,855,882]" fontSize="11" level="1" pageId="4" pageNumber="339" reason="1">
<emphasis bold="true" box="[151,938,855,882]" pageId="4" pageNumber="339">
<taxonomicName authority="Ghosh, Jaiswara &amp; Rajaraman" authorityName="Ghosh, Jaiswara &amp; Rajaraman" authorityYear="2023" box="[151,837,855,882]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis bold="true" box="[151,439,855,882]" italics="true" pageId="4" pageNumber="339">Hexacentrus khasiensis</emphasis>
Ghosh, Jaiswara &amp; Rajaraman
</taxonomicName>
<taxonomicNameLabel box="[845,938,855,881]" pageId="4" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="339" type="materials_examined">
<paragraph blockId="4.[151,1437,927,1998]" pageId="4" pageNumber="339">
<emphasis bold="true" box="[151,329,927,953]" pageId="4" pageNumber="339">Type material.</emphasis>
<materialsCitation collectingDate="2021-09-01" collectionCode="ZSI" country="India" elevation="1400" latitude="25.6166" location="Shillong" longLatPrecision="1" longitude="91.9007" municipality="East Khasi Hills" pageId="4" pageNumber="339" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis box="[340,443,927,953]" italics="true" pageId="4" pageNumber="339">
<typeStatus box="[340,443,927,953]" pageId="4" pageNumber="339">Holotype</typeStatus>
</emphasis>
,
<emphasis bold="true" box="[461,591,927,953]" pageId="4" pageNumber="339">
J,
<collectingCountry box="[500,587,927,953]" name="India" pageId="4" pageNumber="339">INDIA</collectingCountry>
:
</emphasis>
<collectingRegion box="[603,730,927,953]" country="India" name="Meghalaya" pageId="4" pageNumber="339">Meghalaya</collectingRegion>
,
<collectingMunicipality box="[745,938,927,954]" pageId="4" pageNumber="339">East Khasi Hills</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:1461A42815694960EDFAFC99FDFEFA41:991743E515694963EED4FC51FB9CFC09" box="[953,1051,927,953]" country="India" latitude="25.6166" longLatPrecision="1" longitude="91.9007" municipality="East Khasi Hills" name="Shillong" pageId="4" pageNumber="339" stateProvince="Meghalaya">Shillong</location>
, ~
<elevation box="[1080,1235,927,954]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="339" unit="m" value="1400.0">
<quantity box="[1080,1170,927,954]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="339" unit="m" value="1400.0">1400 m</quantity>
a.s.l.
</elevation>
,
<date box="[1251,1381,927,953]" pageId="4" pageNumber="339" value="2021-09-01">
<collectingDate box="[1251,1381,927,953]" pageId="4" pageNumber="339" value="2021-09-01">01.IX.2021</collectingDate>
</date>
,
<geoCoordinate degrees="25" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="339" precision="1" seconds="59.76" value="25.6166">25° 36 59.76” N</geoCoordinate>
;
<geoCoordinate box="[318,501,963,990]" degrees="91" direction="east" minutes="54" orientation="longitude" pageId="4" pageNumber="339" precision="1" seconds="2.52" value="91.9007">91° 54 2.52” E</geoCoordinate>
, AG_SH1_M6 (
<collectionCode box="[689,736,963,989]" pageId="4" pageNumber="339">ZSI</collectionCode>
)
</materialsCitation>
.
<materialsCitation box="[757,1432,963,993]" collectingDate="2021-09-01" collectionCode="ZSI" country="India" elevation="1400" latitude="25.6166" location="Shillong" longLatPrecision="1" longitude="91.9007" municipality="East Khasi Hills" pageId="4" pageNumber="339" specimenCount="1" specimenCount-female="1" stateProvince="Meghalaya" typeStatus="allotype">
<emphasis box="[757,850,963,989]" italics="true" pageId="4" pageNumber="339">
<typeStatus box="[757,850,963,989]" pageId="4" pageNumber="339">Allotype</typeStatus>
</emphasis>
,
<emphasis bold="true" box="[867,895,965,993]" pageId="4" pageNumber="339">
<specimenCount box="[867,888,965,993]" pageId="4" pageNumber="339" type="female"></specimenCount>
,
</emphasis>
AG_SH1_F1, same locality as holotype (
<collectionCode box="[1377,1424,963,989]" pageId="4" pageNumber="339">ZSI</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="2021-08" collectingDateMax="2021-09-31" collectingDateMin="2021-08" collectionCode="AG, BNHS" country="India" pageId="4" pageNumber="339" specimenCode="SH1" specimenCount="3" specimenCount-female="3" stateProvince="Meghalaya" typeStatus="paratype">
<emphasis box="[151,265,1000,1025]" italics="true" pageId="4" pageNumber="339">
<typeStatus box="[151,265,1000,1025]" pageId="4" pageNumber="339">Paratypes</typeStatus>
</emphasis>
(4
<emphasis bold="true" box="[296,317,999,1025]" pageId="4" pageNumber="339">J</emphasis>
,
<specimenCount box="[331,366,999,1029]" pageId="4" pageNumber="339" type="female">
3
<emphasis bold="true" box="[345,366,1001,1029]" pageId="4" pageNumber="339"></emphasis>
</specimenCount>
), same information as
<typeStatus box="[626,727,999,1025]" pageId="4" pageNumber="339">holotype</typeStatus>
,
<date box="[738,897,999,1025]" pageId="4" pageNumber="339" value="2021-08" valueMax="2021-09-31" valueMin="2021-08-00">
<collectingDate box="[738,897,999,1025]" pageId="4" pageNumber="339" value="2021-08" valueMax="2021-09-31" valueMin="2021-08-00">VIIIIX.2021</collectingDate>
</date>
: 2
<emphasis bold="true" box="[922,959,999,1025]" pageId="4" pageNumber="339">J (</emphasis>
<collectionCode box="[959,1004,999,1025]" pageId="4" pageNumber="339">AG</collectionCode>
_
<specimenCode box="[1020,1067,999,1025]" collectionCode="AG" pageId="4" pageNumber="339">SH1</specimenCode>
_M1, AG_SH1_M2) 2
<emphasis bold="true" box="[1318,1339,999,1025]" pageId="4" pageNumber="339">J</emphasis>
ex-AG_ SH1_M3 (
<collectionCode box="[269,349,1035,1061]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="339">BNHS</collectionCode>
)
</materialsCitation>
, ex-AG_SH1_M4 (
<collectionCode box="[576,656,1035,1061]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="339">BNHS</collectionCode>
);
<materialsCitation box="[676,1388,1035,1065]" collectionCode="AG, BNHS" country="India" pageId="4" pageNumber="339" specimenCode="SH1" specimenCount="3" specimenCount-female="3" stateProvince="Meghalaya" typeStatus="paratype">
<specimenCount box="[676,711,1035,1065]" pageId="4" pageNumber="339" type="female">
2
<emphasis bold="true" box="[690,711,1037,1065]" pageId="4" pageNumber="339"></emphasis>
</specimenCount>
(
<collectionCode box="[728,772,1035,1061]" pageId="4" pageNumber="339">AG</collectionCode>
_
<specimenCode box="[787,833,1035,1061]" collectionCode="AG" pageId="4" pageNumber="339">SH1</specimenCode>
_F2, AG_SH1_F3),
<specimenCount box="[1059,1094,1035,1065]" pageId="4" pageNumber="339" type="female">
1
<emphasis bold="true" box="[1073,1094,1037,1065]" pageId="4" pageNumber="339"></emphasis>
</specimenCount>
ex-AG_SH1_F4 (
<collectionCode box="[1300,1381,1035,1061]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="339">BNHS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="4.[151,1437,927,1998]" box="[199,952,1071,1098]" pageId="4" pageNumber="339">
<materialsCitation box="[199,947,1071,1098]" country="India" location="East Khasi Hill" municipality="Shillong" pageId="4" pageNumber="339" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<emphasis bold="true" box="[199,358,1071,1097]" pageId="4" pageNumber="339">
<typeStatus box="[199,258,1071,1097]" pageId="4" pageNumber="339">Type</typeStatus>
locality.
</emphasis>
<location LSID="urn:lsid:plazi:treatment:1461A42815694960EDFAFC99FDFEFA41:991743E515694963EC01FBFEFD9DFBF9" box="[364,538,1071,1098]" country="India" municipality="Shillong" name="East Khasi Hill" pageId="4" pageNumber="339" stateProvince="Meghalaya">East Khasi Hill</location>
,
<collectingMunicipality box="[549,647,1071,1097]" pageId="4" pageNumber="339">Shillong</collectingMunicipality>
,
<collectingRegion box="[658,785,1071,1097]" country="India" name="Meghalaya" pageId="4" pageNumber="339">Meghalaya</collectingRegion>
,
<collectingCountry box="[797,855,1071,1097]" name="India" pageId="4" pageNumber="339">India</collectingCountry>
(
<figureCitation box="[870,935,1071,1097]" captionStart="FIGURE 1" captionStartId="2.[152,255,1205,1229]" captionTargetBox="[151,1436,270,1179]" captionTargetId="figure-66@2.[151,1436,270,1179]" captionTargetPageId="2" captionText="FIGURE 1. India map showing distributions for all three species including their type localities. The light green colour shows Haryana state, and the dark green colour shows Meghalaya state with other colours highlighting the distribution of H. ashoka and H. tiddae in Sonipat district (dark blue); H. khasiensis distribution in the East-Khasi hills (yellow), West-Khasi hills (red), Ri-Bhoi (sky blue), South Garo hills (purple)." figureDoi="http://doi.org/10.5281/zenodo.7690997" httpUri="https://zenodo.org/record/7690997/files/figure.png" pageId="4" pageNumber="339">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="339" type="distribution">
<paragraph blockId="4.[151,1437,927,1998]" pageId="4" pageNumber="339">
<emphasis bold="true" box="[199,353,1107,1133]" pageId="4" pageNumber="339">Distribution.</emphasis>
In addition to
<typeStatus box="[522,570,1108,1134]" pageId="4" pageNumber="339">type</typeStatus>
locality, this new species has been heard in different parts of the Khasi hills including the Ri-Bhoi and West Khasi Hill districts of
<collectingRegion box="[740,868,1143,1169]" country="India" name="Meghalaya" pageId="4" pageNumber="339">Meghalaya</collectingRegion>
. The same call
<typeStatus box="[1032,1080,1144,1170]" pageId="4" pageNumber="339">type</typeStatus>
has been recorded in Balpakram National Park in the South Garo hill district of
<collectingRegion box="[678,802,1179,1205]" country="India" name="Meghalaya" pageId="4" pageNumber="339">Meghalaya</collectingRegion>
(
<figureCitation box="[818,879,1179,1205]" captionStart="FIGURE 1" captionStartId="2.[152,255,1205,1229]" captionTargetBox="[151,1436,270,1179]" captionTargetId="figure-66@2.[151,1436,270,1179]" captionTargetPageId="2" captionText="FIGURE 1. India map showing distributions for all three species including their type localities. The light green colour shows Haryana state, and the dark green colour shows Meghalaya state with other colours highlighting the distribution of H. ashoka and H. tiddae in Sonipat district (dark blue); H. khasiensis distribution in the East-Khasi hills (yellow), West-Khasi hills (red), Ri-Bhoi (sky blue), South Garo hills (purple)." figureDoi="http://doi.org/10.5281/zenodo.7690997" httpUri="https://zenodo.org/record/7690997/files/figure.png" pageId="4" pageNumber="339">Fig.1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,1112,1215,1242]" pageId="4" pageNumber="339" type="etymology">
<paragraph blockId="4.[151,1437,927,1998]" box="[199,1112,1215,1242]" pageId="4" pageNumber="339">
<emphasis bold="true" box="[199,328,1215,1241]" pageId="4" pageNumber="339">Etymology</emphasis>
. The species is named after the
<typeStatus box="[687,735,1216,1242]" pageId="4" pageNumber="339">type</typeStatus>
locality Khasi hill of
<collectingRegion box="[980,1108,1215,1241]" country="India" name="Meghalaya" pageId="4" pageNumber="339">Meghalaya</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="339" type="diagnosis">
<paragraph blockId="4.[151,1437,927,1998]" pageId="4" pageNumber="339">
<emphasis bold="true" box="[199,322,1251,1277]" pageId="4" pageNumber="339">Diagnosis.</emphasis>
The species is somewhat similar to
<taxonomicName authority="Chen &amp; He, 2021" authorityName="Chen &amp; He" authorityYear="2021" box="[733,1107,1251,1278]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis box="[733,899,1251,1277]" italics="true" pageId="4" pageNumber="339">H. formosanus</emphasis>
<bibRefCitation author="Chen, P. W. &amp; Xie, H. C. &amp; Wu, X. &amp; Shen, C. Z. &amp; He, Z. Q." box="[907,1107,1251,1278]" pageId="4" pageNumber="339" pagination="543 - 556" refId="ref7368" refString="Chen, P. W., Xie, H. C., Wu, X., Shen, C. Z. &amp; He, Z. Q. (2021) A new species of genus Hexacentrus Serville, 1831 from Taiwan (Orthoptera: Tettigoniidae: Hexacentrinae). Zootaxa, 4933 (4), 543 - 556. https: // doi. org / 10.11646 / zootaxa. 4933.4.5" type="journal article" year="2021">Chen &amp; He, 2021</bibRefCitation>
</taxonomicName>
with respect to the marks on the disc of pronotum however it differs as follows: In
<taxonomicName authorityName="Chen &amp; He" authorityYear="2021" box="[760,925,1287,1313]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis box="[760,925,1287,1313]" italics="true" pageId="4" pageNumber="339">H. formosanus</emphasis>
</taxonomicName>
the lateral margin of the pronotum is straight but in
<taxonomicName authority="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman, 2023" authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[221,370,1323,1349]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis box="[221,370,1323,1349]" italics="true" pageId="4" pageNumber="339">H. khasiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[376,465,1323,1349]" pageId="4" pageNumber="339">
<taxonomicNameLabel box="[376,465,1323,1349]" pageId="4" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it is slightly concave. Teeth in the middle region of the stridulatory file in
<taxonomicName authority="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman, 2023" authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[1288,1436,1323,1349]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis box="[1288,1436,1323,1349]" italics="true" pageId="4" pageNumber="339">H. khasiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[151,240,1359,1385]" pageId="4" pageNumber="339">
<taxonomicNameLabel box="[151,240,1359,1385]" pageId="4" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are much wider than the lateral teeth. Stridulatory teeth are more in number in
<taxonomicName authority="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman, 2023" authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[1111,1260,1359,1385]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis box="[1111,1260,1359,1385]" italics="true" pageId="4" pageNumber="339">H. khasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1266,1356,1359,1386]" pageId="4" pageNumber="339" rank="species">
<emphasis bold="true" box="[1266,1349,1359,1385]" pageId="4" pageNumber="339">sp. nov</emphasis>
.
</taxonomicNameLabel>
Apical third of cerci in
<taxonomicName authority="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman, 2023" authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[332,481,1395,1421]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis box="[332,481,1395,1421]" italics="true" pageId="4" pageNumber="339">H. khasiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[488,577,1395,1421]" pageId="4" pageNumber="339">
<taxonomicNameLabel box="[488,577,1395,1421]" pageId="4" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
longer and styli narrower. Female ovipositor is much narrower.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="342" pageId="4" pageNumber="339" type="description">
<paragraph blockId="4.[151,1437,927,1998]" pageId="4" pageNumber="339">
<emphasis bold="true" box="[199,413,1431,1457]" pageId="4" pageNumber="339">Description. Male</emphasis>
. Body medium sized and slender (
<figureCitation box="[801,925,1431,1457]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="4" pageNumber="339">Fig. 3AB</figureCitation>
).
<emphasis bold="true" box="[944,1014,1431,1457]" pageId="4" pageNumber="339">Head.</emphasis>
Fastigium verticis triangular, narrow, laterally compressed with median longitudinal sulcus. Apex narrow, tapering forward, rounded in lateral view (
<figureCitation captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="4" pageNumber="339">Fig. 3C</figureCitation>
). Base of fastigium 1.25 times as wide as scapus. Fastigium verticis separated from fastigium frontis by a furrow. Eyes globular.
<emphasis bold="true" box="[322,450,1539,1565]" pageId="4" pageNumber="339">Pronotum.</emphasis>
Saddle-shaped, longer than broad, and expanded posteriorly. Anterior margin straight. Posterior margin slightly concave and the pronotal disc flat expanded and rounded in posterior region (
<figureCitation box="[1296,1384,1575,1601]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="4" pageNumber="339">Fig. 3C</figureCitation>
). An hourglass-like band covers whole pronotum. Median carina faintly visible. Slightly depressed behind first transverse sulcus; with three transverse sulci. A V-shaped sulcus is present between first and second sulci (
<figureCitation box="[1244,1333,1647,1673]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="4" pageNumber="339">Fig. 3C</figureCitation>
). Second and third transverse sulci restricted to disk. Ventral margin of the lateral lobe inclined downwards longer than high, humeral sinus absent. Anterior angle obtuse, and posterior angle rounded. Posterior part forming a straight slope, and anterior part straight (
<figureCitation box="[452,542,1755,1781]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="4" pageNumber="339">Fig. 3E</figureCitation>
). Thoracic auditory spiracle large and oval, party hidden under lateral lobe of pronotum.
<emphasis bold="true" box="[277,340,1791,1817]" pageId="4" pageNumber="339">Legs.</emphasis>
Fore coxae with a forward outward projecting spine. FI dorsally unarmed and ventrally armed with 34 subapical spurs present on inner margin with 45 spines present in between subapical spurs and outer margin bearing 49 spines, no subapical spurs. TI armed with 6 long subapical spurs, first subapical spur slightly shorter than 2
<superScript attach="left" box="[303,319,1899,1913]" fontSize="6" pageId="4" pageNumber="339">nd</superScript>
subapical spur and then decreasing in length apically on both sides. TII armed with 6 subapical spurs on inner and outer margins. TIII dorsally armed with 2234 spines on the inner margin and 2033 spines on outer margin, ventrally armed with 1115 and 1013 subapical spurs variable in length on outer and inner margin.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7691003" ID-Zenodo-Dep="7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="5" pageNumber="340" startId="5.[152,255,1943,1967]" targetBox="[226,1373,181,1907]" targetPageId="5">
<paragraph blockId="5.[151,1437,1943,2040]" pageId="5" pageNumber="340">
<emphasis bold="true" box="[152,280,1943,1967]" pageId="5" pageNumber="340">FIGURE 3.</emphasis>
<taxonomicName authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[286,524,1944,1968]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="khasiensis">
<emphasis box="[286,524,1944,1968]" italics="true" pageId="5" pageNumber="340">Hexacentrus khasiensis</emphasis>
</taxonomicName>
male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7691005" ID-Zenodo-Dep="7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="6" pageNumber="341" startId="6.[152,255,1943,1967]" targetBox="[206,1381,181,1920]" targetPageId="6">
<paragraph blockId="6.[151,1437,1943,2040]" pageId="6" pageNumber="341">
<emphasis bold="true" box="[152,279,1943,1967]" pageId="6" pageNumber="341">FIGURE 4.</emphasis>
<taxonomicName authorityName="Ghosh &amp; Jaiswara &amp; Sabharwal &amp; Dasoju &amp; Bhattacharjee &amp; Rajaraman" authorityYear="2023" box="[284,520,1944,1968]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="khasiensis">
<emphasis box="[284,520,1944,1968]" italics="true" pageId="6" pageNumber="341">Hexacentrus khasiensis</emphasis>
</taxonomicName>
female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm.
</paragraph>
</caption>
<paragraph blockId="7.[151,1437,151,681]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[199,267,151,177]" pageId="7" pageNumber="342">Male.</emphasis>
Tegmen of medium size, broad in the middle, apical margin not oval in lateral view (
<figureCitation box="[1230,1318,151,177]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="7" pageNumber="342">Fig. 3B</figureCitation>
), reaching middle of hind tibia. Left tegmen long, stridulatory file with 3740 teeth; 1
<superScript attach="left" box="[1002,1013,187,201]" fontSize="6" pageId="7" pageNumber="342">st</superScript>
anal vein teeth broader and elevated, followed by narrow teeth. Mirror longer than wide. Media slightly curved, Cubitus posterior vein slightly bent in middle, space between CuP and CuA slightly wider than M and CuA, 1
<superScript attach="left" box="[944,955,259,273]" fontSize="6" pageId="7" pageNumber="342">st</superScript>
anal vein bearing stridulatory file inclined,
<quantity box="[151,237,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="342" unit="mm" value="1.3">1.3 mm</quantity>
long with 3740 teeth (
<figureCitation box="[504,608,295,321]" captionStart="FIGURE 14" captionStartId="19.[152,255,1981,2005]" captionTargetBox="[189,1399,181,1956]" captionTargetId="figure-19@19.[189,1399,181,1956]" captionTargetPageId="19" captionText="FIGURE 14. Stridulatory apparatus structure—ventral side of left wing with file and scraper on the dorsal side of right wing (AB) of H. khasiensis; (CD) of H. ashoka; (EF) of H. tiddae." figureDoi="http://doi.org/10.5281/zenodo.7691029" httpUri="https://zenodo.org/record/7691029/files/figure.png" pageId="7" pageNumber="342">Fig. 14A</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,681]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[199,395,330,357]" pageId="7" pageNumber="342">Male. Genitalia.</emphasis>
Supra-anal plate triangular with a dorsal groove, apex round. Cerci long cylindrical, then slowly narrowing to a long digitiform apical appendage with incurved apex (
<figureCitation box="[1035,1127,367,393]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="7" pageNumber="342">Fig. 3D</figureCitation>
). Subgenital plate elongate with a median longitudinal furrow, lateral ridges well developed, apical margin with deeply curved (
<figureCitation box="[1274,1359,403,429]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[226,1373,181,1907]" captionTargetId="figure-19@5.[214,1373,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Hexacentrus khasiensis male: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum. Scale 5mm, (D) Supra-anal plate, (E) Lateral view of head and pronotum. Scale 5mm, (FG) Dorsal and Lateral view of subgenital plate, Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691003" httpUri="https://zenodo.org/record/7691003/files/figure.png" pageId="7" pageNumber="342">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,681]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[199,293,439,465]" pageId="7" pageNumber="342">Female.</emphasis>
Small body.
<emphasis bold="true" box="[446,567,439,465]" pageId="7" pageNumber="342">Pronotum</emphasis>
. Lateral lobe straight in posterior margin (
<figureCitation box="[1057,1144,439,465]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[206,1381,181,1920]" captionTargetId="figure-27@6.[206,1381,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hexacentrus khasiensis female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="7" pageNumber="342">Fig. 4F</figureCitation>
).
<emphasis bold="true" box="[1166,1242,438,464]" pageId="7" pageNumber="342">Wings</emphasis>
. FW narrow and short, reaching mid ovipositor (
<figureCitation box="[510,600,475,501]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[206,1381,181,1920]" captionTargetId="figure-27@6.[206,1381,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hexacentrus khasiensis female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="7" pageNumber="342">Fig. 4B</figureCitation>
).
<emphasis bold="true" box="[621,826,474,501]" pageId="7" pageNumber="342">Female Genitalia</emphasis>
. Supra-anal plate triangular with dorsal groove, short in length and broad basally, apex rounded (
<figureCitation box="[655,749,511,537]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[206,1381,181,1920]" captionTargetId="figure-27@6.[206,1381,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hexacentrus khasiensis female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="7" pageNumber="342">Fig. 4H</figureCitation>
). Cerci conical, slightly upcurved, and apex diverging. The subgenital plate is triangular and wide (
<figureCitation box="[588,667,547,573]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[206,1381,181,1920]" captionTargetId="figure-27@6.[206,1381,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hexacentrus khasiensis female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="7" pageNumber="342">Fig. 4J</figureCitation>
). Apex with a median excision. Ovipositor broad at the base, straight, and with pointed end (
<figureCitation box="[402,481,583,609]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[206,1381,181,1920]" captionTargetId="figure-27@6.[206,1381,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hexacentrus khasiensis female: (AB) Dorsal and Lateral view. Scale 5mm, (C) Dorsal view of head and pronotum, (D) Ventral view of Specimen, (E) Face, (F) Maxillary palpi. Scale 1mm, (G) Lateral view of head and pronotum, (H) Dorsal view of the subgenital plate. Scale 1mm, (I) Ovipositor in lateral view, (J) Supra-anal plate. Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.7691005" httpUri="https://zenodo.org/record/7691005/files/figure.png" pageId="7" pageNumber="342">Fig. 4I</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,151,681]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[199,328,618,644]" pageId="7" pageNumber="342">Coloration</emphasis>
. Green when alive (
<figureCitation box="[551,681,619,645]" captionStart="FIGURE 2" captionStartId="3.[152,255,1966,1990]" captionTargetBox="[151,1436,561,1943]" captionTargetId="figure-96@3.[151,1436,561,1943]" captionTargetPageId="3" captionText="FIGURE 2. Picture of all 3 species in their natural habitat—H. khasiensis male (A) and female (B), H. ashoka male (C), H. tiddae male (D)." figureDoi="http://doi.org/10.5281/zenodo.7690999" httpUri="https://zenodo.org/record/7690999/files/figure.png" pageId="7" pageNumber="342">Fig. 2B&amp;D</figureCitation>
). Antennae with regularly spaced blackish brown annules. Male FW with stridulatory apparatus and apical field rusty brown. The dorsal part of the female FW is rusty brown.
</paragraph>
<caption pageId="7" pageNumber="342" startId="7.[152,244,728,752]">
<paragraph blockId="7.[151,1436,728,788]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[152,271,728,752]" pageId="7" pageNumber="342">TABLE 2.</emphasis>
Measurements of male and female
<taxonomicName authority="Ghosh, Jaiswara &amp; Rajaraman" authorityName="Ghosh, Jaiswara &amp; Rajaraman" authorityYear="2023" box="[651,1124,728,753]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="342" phylum="Arthropoda" rank="species" species="khasiensis" status="sp. nov.">
<emphasis box="[651,794,728,752]" italics="true" pageId="7" pageNumber="342">H. khasiensis</emphasis>
Ghosh, Jaiswara &amp; Rajaraman
</taxonomicName>
<emphasis bold="true" box="[1130,1216,729,753]" pageId="7" pageNumber="342">
<taxonomicNameLabel box="[1130,1216,729,753]" pageId="7" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations listed in material and method section. Values in mm.
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="342">
<table box="[151,1399,804,1305]" gridcols="13" gridrows="12" pageId="7" pageNumber="342">
<tr box="[151,1399,804,828]" gridrow="0" pageId="7" pageNumber="342" rowspan-7="1">
<th box="[151,350,804,828]" gridcol="0" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[166,231,804,828]" pageId="7" pageNumber="342">Males</emphasis>
</th>
<th box="[402,475,804,828]" gridcol="1" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[405,475,804,828]" pageId="7" pageNumber="342">FW_L</emphasis>
</th>
<th box="[504,681,804,828]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[545,649,804,828]" pageId="7" pageNumber="342">BL_FWL</emphasis>
</th>
<th box="[707,789,804,828]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[707,742,804,828]" pageId="7" pageNumber="342">PH</emphasis>
</th>
<th box="[811,849,804,828]" gridcol="6" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[811,843,804,828]" pageId="7" pageNumber="342">PL</emphasis>
</th>
<th box="[923,1032,804,828]" gridcol="8" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[923,978,804,828]" pageId="7" pageNumber="342">FI_L</emphasis>
</th>
<th box="[1046,1145,804,828]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[1046,1102,804,828]" pageId="7" pageNumber="342">TI_L</emphasis>
</th>
<th box="[1171,1281,804,828]" gridcol="11" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[1171,1245,804,828]" pageId="7" pageNumber="342">FIII_L</emphasis>
</th>
<th box="[1311,1399,804,828]" gridcol="12" gridrow="0" pageId="7" pageNumber="342">
<emphasis bold="true" box="[1311,1387,804,828]" pageId="7" pageNumber="342">TIII_L</emphasis>
</th>
</tr>
<tr box="[151,1399,844,869]" gridrow="1" pageId="7" pageNumber="342" rowspan-10="1" rowspan-5="1" rowspan-7="1">
<th box="[151,350,844,869]" gridcol="0" gridrow="1" pageId="7" pageNumber="342">
<emphasis bold="true" box="[166,265,844,868]" pageId="7" pageNumber="342">Holotype</emphasis>
</th>
<td box="[402,475,844,869]" gridcol="1" gridrow="1" pageId="7" pageNumber="342">21.4</td>
<td box="[504,576,844,869]" gridcol="2" gridrow="1" pageId="7" pageNumber="342">30</td>
<td box="[625,739,844,869]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="7" pageNumber="342">2.9</td>
<td box="[811,849,844,869]" gridcol="6" gridrow="1" pageId="7" pageNumber="342">6.5</td>
<td box="[923,1032,844,869]" gridcol="8" gridrow="1" pageId="7" pageNumber="342">6.5</td>
<td box="[1046,1084,844,869]" gridcol="9" gridrow="1" pageId="7" pageNumber="342">7.3</td>
<td box="[1171,1281,844,869]" gridcol="11" gridrow="1" pageId="7" pageNumber="342">17.4</td>
<td box="[1311,1399,844,869]" gridcol="12" gridrow="1" pageId="7" pageNumber="342">17.3</td>
</tr>
<tr box="[151,1399,885,909]" gridrow="2" pageId="7" pageNumber="342" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th box="[151,350,885,909]" gridcol="0" gridrow="2" pageId="7" pageNumber="342">M_1 paratype dry</th>
<td box="[402,475,885,909]" gridcol="1" gridrow="2" pageId="7" pageNumber="342">21.3</td>
<td box="[504,576,885,909]" gridcol="2" gridrow="2" pageId="7" pageNumber="342">30</td>
<td box="[707,739,885,909]" gridcol="4" gridrow="2" pageId="7" pageNumber="342">4</td>
<td box="[811,849,885,909]" gridcol="6" gridrow="2" pageId="7" pageNumber="342">6.4</td>
<td box="[923,1032,885,909]" gridcol="8" gridrow="2" pageId="7" pageNumber="342">6.7</td>
<td box="[1046,1084,885,909]" gridcol="9" gridrow="2" pageId="7" pageNumber="342">5.7</td>
<td box="[1171,1281,885,909]" gridcol="11" gridrow="2" pageId="7" pageNumber="342">15.8</td>
<td box="[1311,1399,885,909]" gridcol="12" gridrow="2" pageId="7" pageNumber="342">15.2</td>
</tr>
<tr box="[151,1399,925,949]" gridrow="3" pageId="7" pageNumber="342" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th box="[151,350,925,949]" gridcol="0" gridrow="3" pageId="7" pageNumber="342">M_2 paratype dry</th>
<td box="[402,475,925,949]" gridcol="1" gridrow="3" pageId="7" pageNumber="342">21.5</td>
<td box="[504,576,925,949]" gridcol="2" gridrow="3" pageId="7" pageNumber="342">32</td>
<td box="[707,739,925,949]" gridcol="4" gridrow="3" pageId="7" pageNumber="342">3.9</td>
<td box="[811,849,925,949]" gridcol="6" gridrow="3" pageId="7" pageNumber="342">6.5</td>
<td box="[923,1032,925,949]" gridcol="8" gridrow="3" pageId="7" pageNumber="342">6.5</td>
<td box="[1046,1084,925,949]" gridcol="9" gridrow="3" pageId="7" pageNumber="342">7.5</td>
<td box="[1171,1281,925,949]" gridcol="11" gridrow="3" pageId="7" pageNumber="342">16</td>
<td box="[1311,1399,925,949]" gridcol="12" gridrow="3" pageId="7" pageNumber="342">17</td>
</tr>
<tr box="[151,1399,965,990]" gridrow="4" pageId="7" pageNumber="342" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th box="[151,350,965,990]" gridcol="0" gridrow="4" pageId="7" pageNumber="342">M_3 paratype wet</th>
<td box="[402,475,965,990]" gridcol="1" gridrow="4" pageId="7" pageNumber="342">21.3</td>
<td box="[504,576,965,990]" gridcol="2" gridrow="4" pageId="7" pageNumber="342">31</td>
<td box="[707,739,965,990]" gridcol="4" gridrow="4" pageId="7" pageNumber="342">3.1</td>
<td box="[811,849,965,990]" gridcol="6" gridrow="4" pageId="7" pageNumber="342">6.3</td>
<td box="[923,1032,965,990]" gridcol="8" gridrow="4" pageId="7" pageNumber="342">6.3</td>
<td box="[1046,1084,965,990]" gridcol="9" gridrow="4" pageId="7" pageNumber="342">8</td>
<td box="[1171,1281,965,990]" gridcol="11" gridrow="4" pageId="7" pageNumber="342">15.7</td>
<td box="[1311,1399,965,990]" gridcol="12" gridrow="4" pageId="7" pageNumber="342">17.3</td>
</tr>
<tr box="[151,1399,1005,1030]" gridrow="5" pageId="7" pageNumber="342" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
<th box="[151,350,1005,1030]" gridcol="0" gridrow="5" pageId="7" pageNumber="342">M_4 paratype wet</th>
<td box="[402,475,1005,1030]" gridcol="1" gridrow="5" pageId="7" pageNumber="342">23.4</td>
<td box="[504,576,1005,1030]" gridcol="2" gridrow="5" pageId="7" pageNumber="342">35</td>
<td box="[707,739,1005,1030]" gridcol="4" gridrow="5" pageId="7" pageNumber="342">3.7</td>
<td box="[811,849,1005,1030]" gridcol="6" gridrow="5" pageId="7" pageNumber="342">7</td>
<td box="[923,1032,1005,1030]" gridcol="8" gridrow="5" pageId="7" pageNumber="342">6.3</td>
<td box="[1046,1084,1005,1030]" gridcol="9" gridrow="5" pageId="7" pageNumber="342">7.8</td>
<td box="[1171,1281,1005,1030]" gridcol="11" gridrow="5" pageId="7" pageNumber="342">17.8</td>
<td box="[1311,1399,1005,1030]" gridcol="12" gridrow="5" pageId="7" pageNumber="342">17.8</td>
</tr>
<tr box="[151,1399,1079,1103]" gridrow="6" pageId="7" pageNumber="342">
<th box="[151,1399,1079,1103]" colspan="13" colspanRight="12" gridcol="0" gridrow="6" pageId="7" pageNumber="342">Continued.</th>
</tr>
<tr box="[151,1399,1119,1143]" gridrow="7" pageId="7" pageNumber="342" rowspan-6="1" rowspan-9="1">
<th box="[151,350,1119,1143]" gridcol="0" gridrow="7" pageId="7" pageNumber="342">Females</th>
<td box="[402,475,1119,1143]" gridcol="1" gridrow="7" pageId="7" pageNumber="342">BL</td>
<td box="[504,576,1119,1143]" gridcol="2" gridrow="7" pageId="7" pageNumber="342">FW_L</td>
<td box="[625,789,1119,1143]" colspan="3" colspanRight="2" gridcol="3" gridrow="7" pageId="7" pageNumber="342">Ovi_L PH</td>
<td box="[868,906,1119,1143]" gridcol="7" gridrow="7" pageId="7" pageNumber="342">PL</td>
<td box="[923,1032,1119,1143]" gridcol="8" gridrow="7" pageId="7" pageNumber="342">
<emphasis bold="true" box="[977,1032,1119,1143]" pageId="7" pageNumber="342">FI_L</emphasis>
</td>
<td box="[1106,1281,1119,1143]" colspan="2" colspanRight="1" gridcol="10" gridrow="7" pageId="7" pageNumber="342">
<emphasis bold="true" box="[1106,1281,1119,1143]" pageId="7" pageNumber="342">TI_L FIII_L</emphasis>
</td>
<td box="[1311,1399,1119,1143]" gridcol="12" gridrow="7" pageId="7" pageNumber="342">
<emphasis bold="true" box="[1323,1399,1119,1143]" pageId="7" pageNumber="342">TIII_L</emphasis>
</td>
</tr>
<tr box="[151,1399,1159,1183]" gridrow="8" pageId="7" pageNumber="342" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th box="[151,350,1159,1183]" gridcol="0" gridrow="8" pageId="7" pageNumber="342">
<emphasis bold="true" box="[166,257,1159,1183]" pageId="7" pageNumber="342">Allotype</emphasis>
</th>
<td box="[402,475,1159,1183]" gridcol="1" gridrow="8" pageId="7" pageNumber="342">25.3</td>
<td box="[504,576,1159,1183]" gridcol="2" gridrow="8" pageId="7" pageNumber="342">19.4</td>
<td box="[625,681,1159,1183]" gridcol="3" gridrow="8" pageId="7" pageNumber="342">14</td>
<td box="[757,789,1159,1183]" gridcol="5" gridrow="8" pageId="7" pageNumber="342">3.5</td>
<td box="[868,906,1159,1183]" gridcol="7" gridrow="8" pageId="7" pageNumber="342">5.4</td>
<td box="[923,1032,1159,1183]" gridcol="8" gridrow="8" pageId="7" pageNumber="342">6.9</td>
<td box="[1106,1145,1159,1183]" gridcol="10" gridrow="8" pageId="7" pageNumber="342">7.6</td>
<td box="[1171,1281,1159,1183]" gridcol="11" gridrow="8" pageId="7" pageNumber="342">17.3</td>
<td box="[1311,1399,1159,1183]" gridcol="12" gridrow="8" pageId="7" pageNumber="342">18.5</td>
</tr>
<tr box="[151,1399,1199,1224]" gridrow="9" pageId="7" pageNumber="342" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th box="[151,350,1199,1224]" gridcol="0" gridrow="9" pageId="7" pageNumber="342">F_1 paratype dry</th>
<td box="[402,475,1199,1224]" gridcol="1" gridrow="9" pageId="7" pageNumber="342">23.5</td>
<td box="[504,576,1199,1224]" gridcol="2" gridrow="9" pageId="7" pageNumber="342">20</td>
<td box="[625,681,1199,1224]" gridcol="3" gridrow="9" pageId="7" pageNumber="342">11.32</td>
<td box="[757,789,1199,1224]" gridcol="5" gridrow="9" pageId="7" pageNumber="342">4.4</td>
<td box="[868,906,1199,1224]" gridcol="7" gridrow="9" pageId="7" pageNumber="342">5.2</td>
<td box="[923,1032,1199,1224]" gridcol="8" gridrow="9" pageId="7" pageNumber="342">6.8</td>
<td box="[1106,1145,1199,1224]" gridcol="10" gridrow="9" pageId="7" pageNumber="342">8.7</td>
<td box="[1171,1281,1199,1224]" gridcol="11" gridrow="9" pageId="7" pageNumber="342">18</td>
<td box="[1311,1399,1199,1224]" gridcol="12" gridrow="9" pageId="7" pageNumber="342">18.3</td>
</tr>
<tr box="[151,1399,1240,1264]" gridrow="10" pageId="7" pageNumber="342" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th box="[151,350,1240,1264]" gridcol="0" gridrow="10" pageId="7" pageNumber="342">F_2 paratype wet</th>
<td box="[402,475,1240,1264]" gridcol="1" gridrow="10" pageId="7" pageNumber="342">26.5</td>
<td box="[504,576,1240,1264]" gridcol="2" gridrow="10" pageId="7" pageNumber="342">20.2</td>
<td box="[625,681,1240,1264]" gridcol="3" gridrow="10" pageId="7" pageNumber="342">14.3</td>
<td box="[757,789,1240,1264]" gridcol="5" gridrow="10" pageId="7" pageNumber="342">3.2</td>
<td box="[868,906,1240,1264]" gridcol="7" gridrow="10" pageId="7" pageNumber="342">5.3</td>
<td box="[923,1032,1240,1264]" gridcol="8" gridrow="10" pageId="7" pageNumber="342">6.4</td>
<td box="[1106,1145,1240,1264]" gridcol="10" gridrow="10" pageId="7" pageNumber="342">7.1</td>
<td box="[1171,1281,1240,1264]" gridcol="11" gridrow="10" pageId="7" pageNumber="342">17</td>
<td box="[1311,1399,1240,1264]" gridcol="12" gridrow="10" pageId="7" pageNumber="342">17.4</td>
</tr>
<tr box="[151,1399,1280,1305]" gridrow="11" pageId="7" pageNumber="342" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th box="[151,350,1280,1305]" gridcol="0" gridrow="11" pageId="7" pageNumber="342">F_3 paratype wet</th>
<td box="[402,475,1280,1305]" gridcol="1" gridrow="11" pageId="7" pageNumber="342">22.8</td>
<td box="[504,576,1280,1305]" gridcol="2" gridrow="11" pageId="7" pageNumber="342">19.5</td>
<td box="[625,681,1280,1305]" gridcol="3" gridrow="11" pageId="7" pageNumber="342">13.6</td>
<td box="[757,789,1280,1305]" gridcol="5" gridrow="11" pageId="7" pageNumber="342">3.9</td>
<td box="[868,906,1280,1305]" gridcol="7" gridrow="11" pageId="7" pageNumber="342">5.5</td>
<td box="[923,1032,1280,1305]" gridcol="8" gridrow="11" pageId="7" pageNumber="342">4.2</td>
<td box="[1106,1145,1280,1305]" gridcol="10" gridrow="11" pageId="7" pageNumber="342">4.8</td>
<td box="[1171,1281,1280,1305]" gridcol="11" gridrow="11" pageId="7" pageNumber="342">9.9</td>
<td box="[1311,1399,1280,1305]" gridcol="12" gridrow="11" pageId="7" pageNumber="342">11</td>
</tr>
</table>
</paragraph>
<paragraph blockId="7.[151,1436,1351,1522]" pageId="7" pageNumber="342">
<emphasis bold="true" box="[199,486,1351,1378]" pageId="7" pageNumber="342">Acoustic measurements.</emphasis>
The continuous call of
<taxonomicName authorityName="Ghosh, Jaiswara &amp; Rajaraman" authorityYear="2023" box="[741,1003,1351,1377]" class="Insecta" family="Tettigoniidae" genus="Hexacentrus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="342" phylum="Arthropoda" rank="species" species="khasiensis">
<emphasis box="[741,1003,1351,1377]" italics="true" pageId="7" pageNumber="342">Hexacentrus khasiensis</emphasis>
</taxonomicName>
consists of a broadband trill alternating with an amplitude modulated trill (
<figureCitation box="[544,635,1387,1413]" captionStart="FIGURE 6" captionStartId="9.[152,255,1904,1928]" captionTargetBox="[207,1381,182,1875]" captionTargetId="figure-19@9.[203,1385,181,1880]" captionTargetPageId="9" captionText="FIGURE 6. Hexacentrus khasiensis sp. nov. (A) Waveform of calling song, (BD) Oscillogram of the first part of call at various time scales, (F) Spectrogram and (E) Power spectra of the first part of the call (GI) Oscillogram of the second part of call at various time scales, (J) Spectrogram of the second part of call and Power spectra of the high amplitude syllables (K) and low amplitude syllables (L)." figureDoi="http://doi.org/10.5281/zenodo.7691013" httpUri="https://zenodo.org/record/7691013/files/figure.png" pageId="7" pageNumber="342">Fig. 6A</figureCitation>
). All measurements were made for at least 10 syllables per call portion per animal, for multiple animals. In the field, it sounds as if two different individuals are calling because of the continuous droning sound produced by opening and closing of the wing, but recordings in the lab verified that the call was produced by a singular individual.
</paragraph>
</subSubSection>
</treatment>
</document>