427 lines
46 KiB
XML
427 lines
46 KiB
XML
<document id="25C1704BDA73E0C3A971320C3DAB5FAD" ID-CLB-Dataset="8987" ID-DOI="10.11646/zootaxa.4903.4.2" ID-GBIF-Dataset="b3fb2585-d858-4d3d-acde-bf5597a58107" ID-ISSN="1175-5326" ID-Zenodo-Dep="4431300" ID-ZooBank="3A34F874-7DC7-42AF-B87C-57371FB842D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610349161732" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto & Reynoso-Velasco, Daniel" docDate="2021" docId="14038780FFB0FF99FF0E46B4FBAAF9C0" docLanguage="en" docName="zootaxa.4903.4.2.pdf" docOrigin="Zootaxa 4903 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrodactylus miguelangeli Arce-Perez 2021, new species" docType="treatment" docVersion="7" lastPageNumber="497" masterDocId="E83AFFF8FFB1FF9DFF99405CFFBAFF99" masterDocTitle="Two new species of Macrodactylus (Coleoptera: Scarabaeidae: Melolonthinae Macrodactylini) from Mexico and Guatemala" masterLastPageNumber="500" masterPageNumber="493" pageNumber="494" updateTime="1698924125442" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0606C6B03696E8527016FDED1313F883">Two new species of Macrodactylus (Coleoptera: Scarabaeidae: Melolonthinae Macrodactylini) from Mexico and Guatemala</mods:title>
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<mods:name id="8F04FE3D5590B94C42D55326F6F0E05F" type="personal">
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<mods:namePart id="7141EE6B5E20E226B51FDA2EBCF5DE6C">Arce-Pérez, Roberto</mods:namePart>
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<mods:namePart id="777A1ABE5FCD3B0CEA2CF663EFF98D29">Reynoso-Velasco, Daniel</mods:namePart>
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<mods:title id="C135623DC343C84703C600B685E00D57">Zootaxa</mods:title>
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<mods:part id="B61A1BC5FE8C84B3D2E833B3D9417F37">
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<mods:date id="3A4DF1C38C2ED1F156D670E9EF131126">2021</mods:date>
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<mods:number id="326FE2C6A3E4C9BC5CFA1DE304435E23">2021-01-11</mods:number>
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<treatment id="14038780FFB0FF99FF0E46B4FBAAF9C0" ID-DOI="http://doi.org/10.5281/zenodo.4431290" ID-GBIF-Taxon="176569441" ID-Zenodo-Dep="4431290" LSID="urn:lsid:plazi:treatment:14038780FFB0FF99FF0E46B4FBAAF9C0" httpUri="http://treatment.plazi.org/id/14038780FFB0FF99FF0E46B4FBAAF9C0" lastPageId="4" lastPageNumber="497" pageId="1" pageNumber="494">
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<subSubSection id="D4B0651DFFB0FF9CFF0E46B4FECEF8BE" pageId="1" pageNumber="494" type="nomenclature">
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<paragraph id="9C153696FFB0FF9CFF0E46B4FC9BF89B" blockId="1.[151,801,1768,1831]" box="[151,801,1768,1795]" pageId="1" pageNumber="494">
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<heading id="C75D81FAFFB0FF9CFF0E46B4FC9BF89B" bold="true" box="[151,801,1768,1795]" fontSize="11" level="1" pageId="1" pageNumber="494" reason="1">
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<taxonomicName id="5BAA4D15FFB0FF9CFF0E46B4FD3EF89A" authority="Arce-Perez" authorityName="Arce-Perez" authorityYear="2021" box="[151,644,1768,1795]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="494" phylum="Arthropoda" rank="species" species="miguelangeli" status="sp. nov.">
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<emphasis id="AEDEEA84FFB0FF9CFF0E46B4FE4AF89A" bold="true" box="[151,496,1768,1795]" italics="true" pageId="1" pageNumber="494">Macrodactylus miguelangeli</emphasis>
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Arce-Pérez
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</taxonomicName>
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,
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<taxonomicNameLabel id="B5ED57FFFFB0FF9CFD0846B5FC9BF89B" box="[657,801,1768,1795]" pageId="1" pageNumber="494" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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<paragraph id="9C153696FFB0FF9CFF0E4750FECEF8BE" blockId="1.[151,801,1768,1831]" box="[151,372,1804,1831]" pageId="1" pageNumber="494">(Figs. 1–2, 5–8, 17)</paragraph>
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<subSubSection id="D4B0651DFFB0FF9CFF0E4713FC04F860" pageId="1" pageNumber="494" type="materials_examined">
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<paragraph id="9C153696FFB0FF9CFF0E4713FC04F860" blockId="1.[151,1436,1871,2044]" pageId="1" pageNumber="494">
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<emphasis id="AEDEEA84FFB0FF9CFF0E4713FE9BF8F0" bold="true" box="[151,289,1871,1897]" pageId="1" pageNumber="494">Type series.</emphasis>
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<materialsCitation id="2CC23CCBFFB0FF9CFEBE4713FE33F814" ID-GBIF-Occurrence="3014874309" collectingDate="1994-04-16" collectionCode="IEXA" collectorName="M. Goyzueta" country="Guatemala" elevation="1000" latitude="14.65" location="El Palmar" longLatPrecision="1287" longitude="-91.583336" pageId="1" pageNumber="494" specimenCount="1" stateProvince="Quetzaltenango" typeStatus="holotype">
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³.
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<collectingCountry id="E4BD7606FFB0FF9CFE7B4713FD23F8F0" box="[482,665,1871,1897]" name="Guatemala" pageId="1" pageNumber="494">GUATEMALA</collectingCountry>
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:
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<collectingRegion id="5E6EF874FFB0FF9CFD3F4713FCE3F8F0" box="[678,857,1871,1897]" country="Guatemala" name="Quetzaltenango" pageId="1" pageNumber="494">Quetzaltenango</collectingRegion>
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,
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,
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,
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,
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</quantity>
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,
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<date id="E8141056FFB0FF9CFA804713FA22F8F0" box="[1305,1432,1871,1897]" pageId="1" pageNumber="494" value="1994-04-16">
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<collectingDate id="F850E9BEFFB0FF9CFA804713FA22F8F0" box="[1305,1432,1871,1897]" pageId="1" pageNumber="494" value="1994-04-16">16.IV.1994</collectingDate>
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</date>
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,
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<collectorName id="315F5340FFB0FF9CFF0E4728FE93F814" box="[151,297,1907,1934]" pageId="1" pageNumber="494">M. Goyzueta</collectorName>
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(
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<collectionCode id="FABBAE53FFB0FF9CFEAE472FFE3AF814" box="[311,384,1907,1933]" pageId="1" pageNumber="494">IEXA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="2CC23CCBFFB0FF9CFE08472FFC14F814" ID-GBIF-Occurrence="3014874303" box="[401,942,1907,1934]" collectingDate="1994-04-16" collectionCode="IEXA" collectorName="M. Goyzueta" country="Guatemala" elevation="1000" latitude="14.65" location="El Palmar" longLatPrecision="1287" longitude="-91.583336" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Quetzaltenango" typeStatus="allotype">
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: same data as holotype (
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<collectionCode id="FABBAE53FFB0FF9CFCC6472FFC1DF814" box="[863,935,1907,1933]" pageId="1" pageNumber="494">IEXA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="2CC23CCBFFB0FF9CFC21472FFE3FF828" ID-GBIF-Occurrence="3014874305" collectingDate="1994-04-16" collectionCode="IEXA" collectorName="M. Goyzueta" country="Guatemala" elevation="1000" latitude="14.65" location="El Palmar" longLatPrecision="1287" longitude="-91.583336" pageId="1" pageNumber="494" specimenCount="9" specimenCount-female="9" stateProvince="Quetzaltenango" typeStatus="paratype">
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||
<typeStatus id="43118834FFB0FF9CFC21472FFBE9F814" box="[952,1107,1907,1933]" pageId="1" pageNumber="494" type="paratype">PARATYPES</typeStatus>
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(4³,
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<specimenCount id="8AACFD1FFFB0FF9CFB0A4728FB02F817" box="[1171,1208,1908,1934]" pageId="1" pageNumber="494" type="female">7♀</specimenCount>
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||
): same data as holotype (1³,
|
||
<specimenCount id="8AACFD1FFFB0FF9CFE9247CBFE97F828" box="[267,301,1943,1969]" pageId="1" pageNumber="494" type="female">2♀</specimenCount>
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<collectionCode id="FABBAE53FFB0FF9CFEAC47CBFEC7F828" box="[309,381,1943,1969]" pageId="1" pageNumber="494">IEXA</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="2CC23CCBFFB0FF9CFE0847CBFB5CF828" ID-GBIF-Occurrence="3014874308" box="[401,1254,1943,1970]" collectingDate="1956-04-18" collectionCode="EMEC" collectorName="Janzen" country="Guatemala" elevation="500" location="Coatepeque" pageId="1" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Quetzaltenango" typeStatus="paratype">
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||
<collectingRegion id="5E6EF874FFB0FF9CFE0847CBFDFEF828" box="[401,580,1943,1969]" country="Guatemala" name="Quetzaltenango" pageId="1" pageNumber="494">Quetzaltenango</collectingRegion>
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||
,
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<location id="9975604DFFB0FF9CFDC947CBFD6DF828" LSID="urn:lsid:plazi:treatment:14038780FFB0FF99FF0E46B4FBAAF9C0:9975604DFFB0FF9CFDC947CBFD6DF828" box="[592,727,1943,1969]" country="Guatemala" name="Coatepeque" pageId="1" pageNumber="494" stateProvince="Quetzaltenango">Coatepeque</location>
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,
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<quantity id="5B529B73FFB0FF9CFD7A47CBFC97F82B" box="[739,813,1943,1970]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="494" unit="m" value="500.0">
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<elevation id="1787D1A5FFB0FF9CFD7A47CBFC97F82B" box="[739,813,1943,1970]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="494" unit="m" value="500.0">500 m</elevation>
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</quantity>
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,
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<date id="E8141056FFB0FF9CFCA347CBFC02F82B" box="[826,952,1943,1970]" pageId="1" pageNumber="494" value="1956-04-18">
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<collectingDate id="F850E9BEFFB0FF9CFCA347CBFC02F82B" box="[826,952,1943,1970]" pageId="1" pageNumber="494" value="1956-04-18">18.IV.1956</collectingDate>
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</date>
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, D.H.
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||
<collectorName id="315F5340FFB0FF9CFB9A47C4FBF5F82B" box="[1027,1103,1944,1970]" pageId="1" pageNumber="494">Janzen</collectorName>
|
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(
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||
<specimenCount id="8AACFD1FFFB0FF9CFBFA47CBFB3EF828" box="[1123,1156,1943,1969]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
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||
<collectionCode id="FABBAE53FFB0FF9CFB1247CBFB64F828" box="[1163,1246,1943,1969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="1" pageNumber="494" type="University or college">EMEC</collectionCode>
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||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2CC23CCBFFB0FF9CFB6B47CBFD3EF84C" ID-GBIF-Occurrence="3014874307" collectingDate="1956-04-16" collectionCode="CMNC" country="Guatemala" location="Mazatenango" pageId="1" pageNumber="494" specimenCount="2" specimenCount-female="2" stateProvince="Suchitepequez" typeStatus="paratype">
|
||
<collectingRegion id="5E6EF874FFB0FF9CFB6B47CBFA22F828" box="[1266,1432,1943,1969]" country="Guatemala" name="Suchitepequez" pageId="1" pageNumber="494">Suchitepequez</collectingRegion>
|
||
,
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||
<location id="9975604DFFB0FF9CFF0E47E0FE8BF84F" LSID="urn:lsid:plazi:treatment:14038780FFB0FF99FF0E46B4FBAAF9C0:9975604DFFB0FF9CFF0E47E0FE8BF84F" box="[151,305,1980,2006]" country="Guatemala" name="Mazatenango" pageId="1" pageNumber="494" stateProvince="Suchitepequez">Mazatenango</location>
|
||
,
|
||
<date id="E8141056FFB0FF9CFEA547E7FE02F84F" box="[316,440,1979,2006]" pageId="1" pageNumber="494" value="1956-04-16">
|
||
<collectingDate id="F850E9BEFFB0FF9CFEA547E7FE02F84F" box="[316,440,1979,2006]" pageId="1" pageNumber="494" value="1956-04-16">16.IV.1956</collectingDate>
|
||
</date>
|
||
(1³,
|
||
<specimenCount id="8AACFD1FFFB0FF9CFE6347E7FDA7F84C" box="[506,541,1979,2005]" pageId="1" pageNumber="494" type="female">2♀</specimenCount>
|
||
<collectionCode id="FABBAE53FFB0FF9CFDBC47E7FDC7F84C" box="[549,637,1979,2005]" pageId="1" pageNumber="494">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="2CC23CCBFFB0FF9CFD1647E7FC00F860" ID-GBIF-Occurrence="3014874306" collectingDate="2001-06-12" collectionCode="VI, IEXA, MXAL" collectorName="Galdamez" country="Mexico" county="Escuintla" elevation="2000" location="Nueva Morelia" municipality="Reserva Cordon Paxtal" pageId="1" pageNumber="494" specimenCount="2" specimenCount-female="2" stateProvince="Chiapas" typeStatus="paratype">
|
||
<emphasis id="AEDEEA84FFB0FF9CFD1647E7FCB4F84C" bold="true" box="[655,782,1979,2005]" pageId="1" pageNumber="494">
|
||
<collectingCountry id="E4BD7606FFB0FF9CFD1647E7FCB0F84C" box="[655,778,1979,2005]" name="Mexico" pageId="1" pageNumber="494">MEXICO</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="5E6EF874FFB0FF9CFC8F47E7FCC9F84C" box="[790,883,1979,2005]" country="Mexico" name="Chiapas" pageId="1" pageNumber="494">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCounty id="75744E1AFFB0FF9CFCE747E7FC53F84C" box="[894,1001,1979,2005]" pageId="1" pageNumber="494">Escuintla</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="7C71ACECFFB0FF9CFC6C47E0FB41F84C" box="[1013,1275,1979,2006]" pageId="1" pageNumber="494">Reserva Cordón Paxtal</collectingMunicipality>
|
||
,
|
||
<location id="9975604DFFB0FF9CFA9F47E0FF0CF860" LSID="urn:lsid:plazi:treatment:14038780FFB0FF99FF0E46B4FBAAF9C0:9975604DFFB0FF9CFA9F47E0FF0CF860" country="Mexico" county="Escuintla" municipality="Reserva Cordon Paxtal" name="Nueva Morelia" pageId="1" pageNumber="494" stateProvince="Chiapas">Nueva Morelia</location>
|
||
,
|
||
<quantity id="5B529B73FFB0FF9CFF584783FEA2F863" box="[193,280,2015,2042]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="494" unit="m" value="2000.0">
|
||
<elevation id="1787D1A5FFB0FF9CFF584783FEA2F863" box="[193,280,2015,2042]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="494" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="E8141056FFB0FF9CFEBD4783FE1CF860" box="[292,422,2015,2041]" pageId="1" pageNumber="494" value="2001-06-12">
|
||
<collectingDate id="F850E9BEFFB0FF9CFEBD4783FE1CF860" box="[292,422,2015,2041]" pageId="1" pageNumber="494" value="2001-06-12">
|
||
12.
|
||
<collectionCode id="FABBAE53FFB0FF9CFEDD4783FEDFF860" box="[324,357,2015,2041]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="494">VI</collectionCode>
|
||
.2001
|
||
</collectingDate>
|
||
</date>
|
||
, E.
|
||
<collectorName id="315F5340FFB0FF9CFE484783FDF9F860" box="[465,579,2015,2041]" pageId="1" pageNumber="494">Galdámez</collectorName>
|
||
(1³,
|
||
<specimenCount id="8AACFD1FFFB0FF9CFD1A4783FD1CF860" box="[643,678,2015,2041]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
|
||
<collectionCode id="FABBAE53FFB0FF9CFD344783FD49F860" box="[685,755,2015,2041]" pageId="1" pageNumber="494">IEXA</collectionCode>
|
||
; 1³,
|
||
<specimenCount id="8AACFD1FFFB0FF9CFCA94783FCE9F860" box="[816,851,2015,2041]" pageId="1" pageNumber="494" type="female">1♀</specimenCount>
|
||
<collectionCode id="FABBAE53FFB0FF9CFCC34783FC08F860" box="[858,946,2015,2041]" pageId="1" pageNumber="494">MXAL</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4B0651DFFB3FF9FFF5E40CAFC3CF850" pageId="2" pageNumber="495" type="diagnosis">
|
||
<paragraph id="9C153696FFB3FF9FFF5E40CAFDE9FE84" blockId="2.[151,1437,150,285]" pageId="2" pageNumber="495">
|
||
<emphasis id="AEDEEA84FFB3FF9FFF5E40CAFEF8FF29" bold="true" box="[199,322,150,176]" pageId="2" pageNumber="495">Diagnosis.</emphasis>
|
||
Integument of the body reddish yellow; pronotum, elytra, and pygidium covered with pale-yellow vestiture; elytra with three broad, longitudinal bands of setae; tarsi without rings of setae (Figs. 1–2); protibia with a well-developed keel basally (not visible in the figures); pygidial plate yellowish red (Fig. 2); parameres oval, glabrous, with lanceolate apex (Figs. 5–6).
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB3FF9FFF0E4736FC3CF850" blockId="2.[151,1437,1898,1994]" pageId="2" pageNumber="495">
|
||
<emphasis id="AEDEEA84FFB3FF9FFF0E4736FEACF81B" bold="true" box="[151,278,1898,1922]" pageId="2" pageNumber="495">Figures 5–8</emphasis>
|
||
. Genital structures of the male and female of
|
||
<taxonomicName id="5BAA4D15FFB3FF9FFD694736FBA8F81B" authority="Arce-Pérez & Reynoso-Velasco, 2021" authorityName="Arce-Pérez & Reynoso-Velasco" authorityYear="2021" box="[752,1042,1898,1922]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="miguelangeli" status="sp. nov.">
|
||
<emphasis id="AEDEEA84FFB3FF9FFD694736FBA8F81B" box="[752,1042,1898,1922]" italics="true" pageId="2" pageNumber="495">Macrodactylus miguelangeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AEDEEA84FFB3FF9FFB834736FB22F81B" bold="true" box="[1050,1176,1898,1922]" pageId="2" pageNumber="495">
|
||
<taxonomicNameLabel id="B5ED57FFFFB3FF9FFB834736FB22F81B" box="[1050,1176,1898,1922]" pageId="2" pageNumber="495" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="AEDEEA84FFB3FF9FFB3C4736FB08F81B" bold="true" box="[1189,1202,1898,1922]" pageId="2" pageNumber="495">5</emphasis>
|
||
, Male parameres, posterior view (arrow indicates keel);
|
||
<emphasis id="AEDEEA84FFB3FF9FFE6647D2FDB6F83F" bold="true" box="[511,524,1934,1958]" pageId="2" pageNumber="495">6</emphasis>
|
||
, male parameres, lateral view;
|
||
<emphasis id="AEDEEA84FFB3FF9FFCC847D2FCE4F83F" bold="true" box="[849,862,1934,1958]" pageId="2" pageNumber="495">7</emphasis>
|
||
, male phallobase, dorsal view;
|
||
<emphasis id="AEDEEA84FFB3FF9FFB3F47D2FB09F83F" bold="true" box="[1190,1203,1934,1958]" pageId="2" pageNumber="495">8</emphasis>
|
||
, female genital plates, posterior view. Scale bars in 5–6 =
|
||
<quantity id="5B529B73FFB3FF9FFE6447EEFDE3F850" box="[509,601,1970,1994]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="495" unit="mm" value="0.44">0.44 mm</quantity>
|
||
, 7 =
|
||
<quantity id="5B529B73FFB3FF9FFD1247EEFD72F850" box="[651,712,1970,1994]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="495" unit="mm" value="1.0">1 mm</quantity>
|
||
, and 8 =
|
||
<quantity id="5B529B73FFB3FF9FFCBC47EEFC38F850" box="[805,898,1970,1994]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="495" unit="mm" value="0.43">0.43 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4B0651DFFB2FF9EFF5E40CAFED7FB5C" pageId="3" pageNumber="496" type="description">
|
||
<paragraph id="9C153696FFB2FF9EFF5E40CAFE28FD68" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E40CAFEE3FF29" bold="true" box="[199,345,150,176]" pageId="3" pageNumber="496">Description.</emphasis>
|
||
<typeStatus id="43118834FFB2FF9EFEF840CBFE77FF28" box="[353,461,151,177]" pageId="3" pageNumber="496" type="holotype">Holotype</typeStatus>
|
||
. Male: Body length 11.00 mm; humeral width
|
||
<quantity id="5B529B73FFB2FF9EFC7C40CAFBF4FF28" box="[997,1102,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="496" unit="mm" value="3.5">3.50 mm</quantity>
|
||
.
|
||
<emphasis id="AEDEEA84FFB2FF9EFBC340CBFB1FFF28" box="[1114,1189,151,177]" italics="true" pageId="3" pageNumber="496">Color:</emphasis>
|
||
body reddish yellow; clypeus, antennae, elytra, legs, and pygidium darker; head and pronotum dark red; pterothorax, abdomen, apex of tibiae, and tarsi dark (Figs. 1–2).
|
||
<emphasis id="AEDEEA84FFB2FF9EFD974083FDD7FF60" box="[526,621,223,249]" italics="true" pageId="3" pageNumber="496">Surface:</emphasis>
|
||
head and pronotum with scattered yellow setae not completely covering the integument; elytra with three broad, longitudinal bands of setae; ventrally with long yellow vestiture; pygidium with scattered vestiture, similar to that of the venter (Fig. 2).
|
||
<emphasis id="AEDEEA84FFB2FF9EFCAA417BFCC2FED8" box="[819,888,295,321]" italics="true" pageId="3" pageNumber="496">Head:</emphasis>
|
||
elongate; clypeus trapezoidal, dark red, punctate, with anterior border straight.
|
||
<emphasis id="AEDEEA84FFB2FF9EFE704117FDF9FEFC" box="[489,579,331,357]" italics="true" pageId="3" pageNumber="496">Thorax:</emphasis>
|
||
pronotum hexagonal, convex, length
|
||
<quantity id="5B529B73FFB2FF9EFC6F4117FBE5FEFC" box="[1014,1119,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="496" unit="mm" value="2.8">2.80 mm</quantity>
|
||
, width
|
||
<quantity id="5B529B73FFB2FF9EFB2C4117FAA5FEFC" box="[1205,1311,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="496" unit="mm" value="2.7">2.70 mm</quantity>
|
||
; prosternal process long, flattened, apex rounded (easily visible in frontal view); scutellum subtriangular; elytra with three well-defined longitudinal bands of pale-yellow setae on interstriae 1, 3, and 5; bands 1 and 5 slightly converging posteriorly, band 3 short and broad, and one faint sublateral row of setae (Fig. 1); interstriae 2 and 4 not raised (not costate). Metasternum broad, with two longitudinal rows of spine-like setae between the mesocoxae and metacoxae.
|
||
<emphasis id="AEDEEA84FFB2FF9EFF0E41A3FF6CFD81" box="[151,214,511,536]" italics="true" pageId="3" pageNumber="496">Legs:</emphasis>
|
||
profemur with a subrectangular projection near the base; ventral surface of protibia with a well-developed, convex keel in basal half; mesotibiae widened toward the apex, with two long, straight, acute spurs; tarsi without rings of white setae; all tarsal claws cleft.
|
||
<emphasis id="AEDEEA84FFB2FF9EFDED421BFD5DFDF8" box="[628,743,583,609]" italics="true" pageId="3" pageNumber="496">Abdomen:</emphasis>
|
||
robust, sternites 2–5 moderately concave (lateral view), with two or three spine-like, slender setae on each side of midline; pygidial plate elongate, oval, convex, with long setae apically.
|
||
<emphasis id="AEDEEA84FFB2FF9EFF6542D3FED4FD30" box="[252,366,655,681]" italics="true" pageId="3" pageNumber="496">Genitalia:</emphasis>
|
||
total length
|
||
<quantity id="5B529B73FFB2FF9EFE6342D2FDE4FD30" box="[506,606,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="3" pageNumber="496" unit="mm" value="3.05">3.05 mm</quantity>
|
||
(Fig. 7); parameres length
|
||
<quantity id="5B529B73FFB2FF9EFC1542D3FC48FD30" box="[908,1010,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="3" pageNumber="496" unit="mm" value="1.32">1.32 mm</quantity>
|
||
, width
|
||
<quantity id="5B529B73FFB2FF9EFBDD42D3FB13FD30" box="[1092,1193,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="496" unit="mm" value="0.96">0.96 mm</quantity>
|
||
, stout, oval (posterior view), glabrous, completely fused dorsobasally, with a short, transversal keel above junction, each apex strongly lanceolate (Figs. 5–6).
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB2FF9EFF5E42A6FD48FC18" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E42A6FE38FC8D" bold="true" box="[199,386,762,788]" pageId="3" pageNumber="496">
|
||
<typeStatus id="43118834FFB2FF9EFF5E42A6FE96FC8D" box="[199,300,762,788]" pageId="3" pageNumber="496" type="allotype">Allotype</typeStatus>
|
||
female
|
||
</emphasis>
|
||
: Similar to male except for: body length
|
||
<quantity id="5B529B73FFB2FF9EFCD342A6FC05FC8C" box="[842,959,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="496" unit="mm" value="10.5">10.50 mm</quantity>
|
||
; humeral width
|
||
<quantity id="5B529B73FFB2FF9EFBE842A7FB62FC8C" box="[1137,1240,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.48" pageId="3" pageNumber="496" unit="mm" value="3.48">3.48 mm</quantity>
|
||
. Pronotum length
|
||
<quantity id="5B529B73FFB2FF9EFF0E4343FF44FCA0" box="[151,254,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="496" unit="mm" value="2.6">2.60 mm</quantity>
|
||
, width
|
||
<quantity id="5B529B73FFB2FF9EFECA4343FE01FCA0" box="[339,443,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="496" unit="mm" value="2.8">2.80 mm</quantity>
|
||
; abdominal sternites convex (lateral view), with slender, scattered, reddish-yellow setae along midline; pygidium short, subtriangular. Genital plates large, nearly subtriangular, length
|
||
<quantity id="5B529B73FFB2FF9EFB2C431FFAA0FCC4" box="[1205,1306,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="496" unit="mm" value="1.3">1.30 mm</quantity>
|
||
, width
|
||
<quantity id="5B529B73FFB2FF9EFAF2431EFF7FFC18" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="496" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, lateral margin slightly expanded basally (Fig. 8).
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB2FF9EFF5E43D6FCB2FC50" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E43D6FEFBFC3D" bold="true" box="[199,321,906,932]" pageId="3" pageNumber="496">Variation:</emphasis>
|
||
Body length 10.50–11.00 mm; humeral width 3.00–3.50; ventrally intense dark red; elytra with fine vestiture; prosternal process small and not conspicuous.
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB2FF9EFF5E438EFDE2FBE4" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E438EFDBEFC75" bold="true" box="[199,516,978,1004]" pageId="3" pageNumber="496">Geographical distribution.</emphasis>
|
||
This species is distributed in the southern Mexican state of
|
||
<collectingRegion id="5E6EF874FFB2FF9EFB07438FFB42FC74" box="[1182,1272,979,1005]" country="Mexico" name="Chiapas" pageId="3" pageNumber="496">Chiapas</collectingRegion>
|
||
and the southwestern Guatemalan departments of
|
||
<collectingRegion id="5E6EF874FFB2FF9EFDAE43ABFD5DFB88" box="[567,743,1015,1041]" country="Guatemala" name="Quetzaltenango" pageId="3" pageNumber="496">Quetzaltenango</collectingRegion>
|
||
and
|
||
<collectingRegion id="5E6EF874FFB2FF9EFC8643ABFC78FB88" box="[799,962,1015,1041]" country="Guatemala" name="Suchitepequez" pageId="3" pageNumber="496">Suchitepequez</collectingRegion>
|
||
(Fig. 17). The localities in both countries are in the
|
||
<collectingRegion id="5E6EF874FFB2FF9EFE9D4447FEE4FBAC" box="[260,350,1051,1077]" country="Mexico" name="Chiapas" pageId="3" pageNumber="496">Chiapas</collectingRegion>
|
||
Highlands biogeographic province. This new species is part of the group of species of
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFAB84447FF76FBC0" authorityName="Dejean" authorityYear="1821" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFAB84447FF76FBC0" italics="true" pageId="3" pageNumber="496">Macrodactylus</emphasis>
|
||
</taxonomicName>
|
||
distributed in the Central American Nucleus (see
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFC9A4463FB82FBC0" author="Arce-Perez, R. & Moron, M. A." box="[771,1080,1087,1113]" pageId="3" pageNumber="496" pagination="567 - 584" refId="ref3299" refString="Arce-Perez, R. & Moron, M. A. (2020) Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae) from the Central American Nucleus. Zootaxa, 4772 (3), 567 - 584. https: // doi. org / 10.11646 / zootaxa. 4772.3.7" type="journal article" year="2020">Arce-Pérez & Morón 2020</bibRefCitation>
|
||
). The number of species of this genus distributed in the area reaches 21.
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB2FF9EFF5E44DBFED7FB5C" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E44DBFE3FFB39" bold="true" box="[199,389,1158,1185]" pageId="3" pageNumber="496">Natural history.</emphasis>
|
||
This species inhabits a tropical rain forest at elevations of
|
||
<quantity id="5B529B73FFB2FF9EFB9644DAFB26FB38" box="[1039,1180,1158,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="3" pageNumber="496" unit="m" value="1250.0" valueMax="2000.0" valueMin="500.0">500–2000 m</quantity>
|
||
and was collected during April and June.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4B0651DFFB2FF9EFF5E4493FB82FA94" pageId="3" pageNumber="496" type="etymology">
|
||
<paragraph id="9C153696FFB2FF9EFF5E4493FB82FA94" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E4493FEF4FB70" bold="true" box="[199,334,1231,1257]" pageId="3" pageNumber="496">Etymology.</emphasis>
|
||
This species is dedicated to the memory of Dr. Miguel Ángel Morón Ríos, who was a mentor of R.A.P. and contributed extensively to the knowledge of Neotropical scarab beetles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4B0651DFFB2FF99FF5E454AFBAAF9C0" lastPageId="4" lastPageNumber="497" pageId="3" pageNumber="496" type="discussion">
|
||
<paragraph id="9C153696FFB2FF9EFF5E454AFD98F8E8" blockId="3.[151,1437,150,1905]" pageId="3" pageNumber="496">
|
||
<emphasis id="AEDEEA84FFB2FF9EFF5E454AFEF2FAA9" bold="true" box="[199,328,1302,1328]" pageId="3" pageNumber="496">Comments</emphasis>
|
||
:
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFEC3454BFD27FAA8" authorityName="Arce-Pérez & Reynoso-Velasco" authorityYear="2021" box="[346,669,1303,1329]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="miguelangeli">
|
||
<emphasis id="AEDEEA84FFB2FF9EFEC3454BFD27FAA8" box="[346,669,1303,1329]" italics="true" pageId="3" pageNumber="496">Macrodactylus miguelangeli</emphasis>
|
||
</taxonomicName>
|
||
is included in the
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFCE5454BFC47FAA8" authorityName="Chevrolat" authorityYear="1834" box="[892,1021,1303,1329]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="lineatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFCE5454BFC29FAA9" box="[892,915,1303,1328]" italics="true" pageId="3" pageNumber="496">M</emphasis>
|
||
.
|
||
<emphasis id="AEDEEA84FFB2FF9EFC3D454BFC47FAA8" box="[932,1021,1303,1329]" italics="true" pageId="3" pageNumber="496">lineatus</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<emphasis id="AEDEEA84FFB2FF9EFB2E454BFB4FFAA9" box="[1207,1269,1303,1328]" italics="true" pageId="3" pageNumber="496">sensu</emphasis>
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFB67454BFE9CFACC" author="Arce-Perez, R. & Moron, M. A." pageId="3" pageNumber="496" pagination="123 - 239" refId="ref3097" refString="Arce-Perez, R. & Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, N. S., 97, 123 - 239." type="journal article" year="2000">Arce-Pérez & Morón 2000</bibRefCitation>
|
||
) because it has the clypeus with the anterior margin nearly straight; the metasternum with irregular, longitudinal rows with few short setae between the mesocoxae and metacoxae; the mesotibia has two long, narrow, straight, and acute spurs; the tarsi do not have rings of setae; and the parameres are oval and glabrous. In the work of
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFF6A45FBFD89FA58" author="Arce-Perez, R. & Moron, M. A." box="[243,563,1447,1473]" pageId="3" pageNumber="496" pagination="123 - 239" refId="ref3097" refString="Arce-Perez, R. & Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, N. S., 97, 123 - 239." type="journal article" year="2000">Arce-Pérez & Morón (2000)</bibRefCitation>
|
||
,
|
||
<specimenCount id="8AACFD1FFFB2FF9EFDA745FBFD6FFA58" box="[574,725,1447,1473]" pageId="3" pageNumber="496" type="generic">13 specimens</specimenCount>
|
||
from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFC8145FBFCD4FA58" box="[792,878,1447,1473]" name="Mexico" pageId="3" pageNumber="496">Mexico</collectingCountry>
|
||
and
|
||
<collectingCountry id="E4BD7606FFB2FF9EFC3B45FBFBA1FA58" box="[930,1051,1447,1473]" name="Guatemala" pageId="3" pageNumber="496">Guatemala</collectingCountry>
|
||
were identified (only based on the original description) and reported as
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFDA84597FCEFFA7C" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[561,853,1483,1509]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFDA84597FD72FA7C" box="[561,712,1483,1509]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
Bates, 1887
|
||
</taxonomicName>
|
||
. Of those, the six specimens from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFB4A4597FAF6FA7C" box="[1235,1356,1483,1509]" name="Guatemala" pageId="3" pageNumber="496">Guatemala</collectingCountry>
|
||
(
|
||
<collectingRegion id="5E6EF874FFB2FF9EFAC24597FEA8F993" country="Guatemala" name="Quetzaltenango" pageId="3" pageNumber="496">Quetzaltenango</collectingRegion>
|
||
) exhibited some superficial differences that at the time were considered as geographical variation. After reviewing approximately
|
||
<specimenCount id="8AACFD1FFFB2FF9EFE2F464EFDE6F9B4" box="[438,604,1554,1581]" pageId="3" pageNumber="496" type="generic">500 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFD19464FFCADF9B4" authorityName="Bates" authorityYear="1887" box="[640,791,1555,1581]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFD19464FFCADF9B4" box="[640,791,1555,1581]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFCC2464FFC6DF9B4" box="[859,983,1555,1581]" name="Costa Rica" pageId="3" pageNumber="496">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="E4BD7606FFB2FF9EFC7B464FFBDDF9B4" box="[994,1127,1555,1581]" name="El Salvador" pageId="3" pageNumber="496">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="E4BD7606FFB2FF9EFBEB464FFB54F9B4" box="[1138,1262,1555,1581]" name="Guatemala" pageId="3" pageNumber="496">Guatemala</collectingCountry>
|
||
,
|
||
<collectingCountry id="E4BD7606FFB2FF9EFB60464FFAD3F9B4" box="[1273,1385,1555,1581]" name="Honduras" pageId="3" pageNumber="496">Honduras</collectingCountry>
|
||
, and
|
||
<collectingCountry id="E4BD7606FFB2FF9EFF0E466BFF57F9C8" box="[151,237,1591,1617]" name="Mexico" pageId="3" pageNumber="496">Mexico</collectingCountry>
|
||
reported in previous works (
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFDB7466BFCE5F9C8" author="Arce-Perez, R. & Moron, M. A." box="[558,863,1590,1617]" pageId="3" pageNumber="496" pagination="23 - 27" refId="ref3148" refString="Arce-Perez, R. & Moron, M. A. (2005) New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Honduras and Nicaragua. Zootaxa, 1012 (1), 23 - 27. https: // doi. org / 10.11646 / zootaxa. 1012.1.3" type="journal article" year="2005">Arce-Pérez & Morón 2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFCF3466BFC1FF9C8" author="Arce-Perez, R. & Moron, M. A." box="[874,933,1591,1617]" pageId="3" pageNumber="496" pagination="379 - 403" refId="ref3245" refString="Arce-Perez, R. & Moron, M. A. (2018) Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica and Panama. Zootaxa, 4462, 379 - 403. https: // doi. org / 10.11646 / zootaxa. 4462.3.5" type="journal article" year="2018">2018</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFC36466BFC51F9C8" author="Arce-Perez, R. & Moron, M. A." box="[943,1003,1591,1617]" pageId="3" pageNumber="496" pagination="567 - 584" refId="ref3299" refString="Arce-Perez, R. & Moron, M. A. (2020) Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae) from the Central American Nucleus. Zootaxa, 4772 (3), 567 - 584. https: // doi. org / 10.11646 / zootaxa. 4772.3.7" type="journal article" year="2020">2020</bibRefCitation>
|
||
), R.A.P found that the specimens from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFF0E4607FEAAF9EC" box="[151,272,1627,1653]" name="Guatemala" pageId="3" pageNumber="496">Guatemala</collectingCountry>
|
||
reported by
|
||
<bibRefCitation id="F83B4B67FFB2FF9EFE394607FD59F9EC" author="Arce-Perez, R. & Moron, M. A." box="[416,739,1627,1653]" pageId="3" pageNumber="496" pagination="123 - 239" refId="ref3097" refString="Arce-Perez, R. & Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, N. S., 97, 123 - 239." type="journal article" year="2000">Arce-Pérez & Morón (2000)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFC934607FC18F9EC" authorityName="Bates" authorityYear="1887" box="[778,930,1627,1653]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFC934607FC18F9EC" box="[778,930,1627,1653]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
, as well as three specimens from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFABA4607FA26F9EC" box="[1315,1436,1627,1653]" name="Guatemala" pageId="3" pageNumber="496">Guatemala</collectingCountry>
|
||
(
|
||
<collectingRegion id="5E6EF874FFB2FF9EFF064623FEFCF903" box="[159,326,1663,1690]" country="Guatemala" name="Suchitepequez" pageId="3" pageNumber="496">Suchitepequez</collectingRegion>
|
||
) and four specimens from
|
||
<collectingCountry id="E4BD7606FFB2FF9EFDEC4623FD71F900" box="[629,715,1663,1689]" name="Mexico" pageId="3" pageNumber="496">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion id="5E6EF874FFB2FF9EFD434623FC8DF903" box="[730,823,1663,1690]" country="Mexico" name="Chiapas" pageId="3" pageNumber="496">Chiapas</collectingRegion>
|
||
) not previously reported, presented specific characteristics that distinguished them as a distinct species, herein described as
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFC3446FFFBDEF924" authorityName="Arce-Pérez & Reynoso-Velasco" authorityYear="2021" box="[941,1124,1699,1725]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="miguelangeli">
|
||
<emphasis id="AEDEEA84FFB2FF9EFC3446FFFBDEF924" box="[941,1124,1699,1725]" italics="true" pageId="3" pageNumber="496">M. miguelangeli</emphasis>
|
||
</taxonomicName>
|
||
. At least five characters are useful in order to distinguish this new species from its closest congener: elytra with three well-defined longitudinal bands of setae (Fig. 1) [only two in
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFDB446B7FD7CF89C" authorityName="Bates" authorityYear="1887" box="[557,710,1771,1797]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFDB446B7FD7CF89C" box="[557,710,1771,1797]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 3)], interstriae 2 and 4 not raised (raised in
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFB6D46B7FA36F89C" authorityName="Bates" authorityYear="1887" box="[1268,1420,1771,1797]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFB6D46B7FA36F89C" box="[1268,1420,1771,1797]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
), pygidial plate yellowish red (Fig. 2) [generally black in
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFC884753FC13F8B0" authorityName="Bates" authorityYear="1887" box="[785,937,1807,1833]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFC884753FC13F8B0" box="[785,937,1807,1833]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 4)], and parameres with a short, transversal keel above junction (Fig. 5) (without keel in
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFD4A476FFCD0F8D4" authorityName="Bates" authorityYear="1887" box="[723,874,1843,1869]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFD4A476FFCD0F8D4" box="[723,874,1843,1869]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
) and apical half of lateral margins straight (Fig. 5) (slightly concave in
|
||
<taxonomicName id="5BAA4D15FFB2FF9EFEE2470BFDA8F8E8" authorityName="Bates" authorityYear="1887" box="[379,530,1879,1905]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="costulatus">
|
||
<emphasis id="AEDEEA84FFB2FF9EFEE2470BFDA8F8E8" box="[379,530,1879,1905]" italics="true" pageId="3" pageNumber="496">M. costulatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9C153696FFB5FF99FF0E4641FBAAF9C0" blockId="4.[151,1437,1565,1625]" pageId="4" pageNumber="497">
|
||
<emphasis id="AEDEEA84FFB5FF99FF0E4641FE9BF9AC" bold="true" box="[151,289,1565,1589]" pageId="4" pageNumber="497">Figures 9–12</emphasis>
|
||
. Dorsal and lateral habitus of species of
|
||
<taxonomicName id="5BAA4D15FFB5FF99FD224641FCE8F9AC" authorityName="Dejean" authorityYear="1821" box="[699,850,1565,1589]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AEDEEA84FFB5FF99FD224641FCE8F9AC" box="[699,850,1565,1589]" italics="true" pageId="4" pageNumber="497">Macrodactylus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="AEDEEA84FFB5FF99FCC64641FC28F9AC" bold="true" box="[863,914,1565,1589]" pageId="4" pageNumber="497">9–10</emphasis>
|
||
, Male
|
||
<typeStatus id="43118834FFB5FF99FC414641FB8BF9AC" box="[984,1073,1565,1589]" pageId="4" pageNumber="497" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="5BAA4D15FFB5FF99FBCB4641FAD1F9AC" authority="Arce-Pérez & Reynoso-Velasco, 2021" authorityName="Arce-Pérez & Reynoso-Velasco" authorityYear="2021" box="[1106,1387,1565,1589]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="oaxacaensis" status="sp. nov.">
|
||
<emphasis id="AEDEEA84FFB5FF99FBCB4641FAD1F9AC" box="[1106,1387,1565,1589]" italics="true" pageId="4" pageNumber="497">Macrodactylus oaxacaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AEDEEA84FFB5FF99FAE84641FF58F9C0" bold="true" pageId="4" pageNumber="497">
|
||
<taxonomicNameLabel id="B5ED57FFFFB5FF99FAE84641FF58F9C0" pageId="4" pageNumber="497" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="AEDEEA84FFB5FF99FF76461DFE97F9C0" bold="true" box="[239,301,1601,1625]" pageId="4" pageNumber="497">11–12</emphasis>
|
||
,
|
||
<taxonomicName id="5BAA4D15FFB5FF99FEA3461EFD67F9C0" authority="Arce-Perez & Moron, 2000" authorityName="Arce-Perez & Moron" authorityYear="2000" box="[314,733,1601,1625]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="AEDEEA84FFB5FF99FEA3461EFE05F9C0" box="[314,447,1602,1625]" italics="true" pageId="4" pageNumber="497">M. montanus</emphasis>
|
||
<bibRefCitation id="F83B4B67FFB5FF99FE5A461DFD67F9C0" author="Arce-Perez, R. & Moron, M. A." box="[451,733,1601,1625]" pageId="4" pageNumber="497" pagination="123 - 239" refId="ref3097" refString="Arce-Perez, R. & Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) en Mexico y Estados Unidos de America. Acta Zoologica Mexicana, N. S., 97, 123 - 239." type="journal article" year="2000">Arce-Pérez & Morón, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
. Scale bars in 9–10 = 5.0 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |