160 lines
21 KiB
XML
160 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.96.104957" ID-Pensoft-Pub="1314-2607-96-599" ID-Pensoft-UUID="A32C8DF9AEC35B35AC4D32F6EEB83552" ID-ZooBank="16A4A16551854C89960D614A74E6D394" ModsDocID="1314-2607-96-599" checkinTime="1690372396501" checkinUser="pensoft" docAuthor="Bogusch, Petr" docDate="2023" docId="13A43AC00DB05E09B31D51DE9341AD6C" docLanguage="en" docName="JourHymenoptRes 96: 599-628" docOrigin="Journal of Hymenoptera Research 96" docPubDate="2023-07-25" docSource="http://dx.doi.org/10.3897/jhr.96.104957" docTitle="Aglaoapis tridentata" docType="treatment" docVersion="1" id="A32C8DF9AEC35B35AC4D32F6EEB83552" lastPageNumber="599" masterDocId="A32C8DF9AEC35B35AC4D32F6EEB83552" masterDocTitle="European cuckoo bees of the tribe Dioxyini (Hymenoptera, Megachilidae): distribution, annotated checklist and identification key" masterLastPageNumber="628" masterPageNumber="599" pageNumber="599" updateTime="1690372396501" updateUser="pensoft">
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<mods:title id="3A03994749D1A97E706EF8DC700551C5">European cuckoo bees of the tribe Dioxyini (Hymenoptera, Megachilidae): distribution, annotated checklist and identification key</mods:title>
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<mods:name id="B546A7B4C38D4C95C8EBE7BAC41BC5BC" type="personal">
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<mods:namePart id="86F74340EDDFC5A24AAFF26A477D1F22">Bogusch, Petr</mods:namePart>
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<mods:affiliation id="3DDFA56E06A31D7A2F174992E26103D3">Department of Biology, Faculty of Science, University of Hradec Kralove, Rokitanskeho 62, CZ- 500 03 Hradec Kralove, Czech Republic</mods:affiliation>
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<mods:nameIdentifier id="E9DBF80AECA9293EA7D5FFDFA2539D91" type="email">bogusch.petr@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="84E748579C5BF8A17D8C7BC890E012AA">text</mods:typeOfResource>
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<mods:title id="AC416ABCB9A15B2B4D631076883220D6">Journal of Hymenoptera Research</mods:title>
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<mods:part id="124CC4BC4AEC16DF63944521E4F8B6E9">
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<mods:date id="6A1785D0CBAE11C33DC4877B937196FF">2023</mods:date>
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<mods:number id="3747956DCB164A53DAFCC19D501C4D07">2023-07-25</mods:number>
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<mods:number id="55E1DE54455781B665801D16B9514EA3">96</mods:number>
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<mods:start id="EADA723BE751BFA2C691F7C3CEDB5104">599</mods:start>
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<mods:end id="0BF5A3B16A502F794F582525FC777569">628</mods:end>
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<mods:url id="62B0B72136812142BC42A6B0017038AB">http://dx.doi.org/10.3897/jhr.96.104957</mods:url>
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<mods:classification id="A8AE25CE5758D1AC00977ECDDAE6758E">journal article</mods:classification>
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<mods:identifier id="238CBE05FAD17CC2DBD6311A074EBF24" type="DOI">http://dx.doi.org/10.3897/jhr.96.104957</mods:identifier>
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<mods:identifier id="3EC00EBC1ECC52568748AAEF76C884DE" type="Pensoft-Pub">1314-2607-96-599</mods:identifier>
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<treatment id="13A43AC00DB05E09B31D51DE9341AD6C" LSID="urn:lsid:plazi:treatment:13A43AC00DB05E09B31D51DE9341AD6C" httpUri="http://treatment.plazi.org/id/13A43AC00DB05E09B31D51DE9341AD6C" lastPageNumber="599" pageId="0" pageNumber="599">
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<subSubSection id="B1E73F78FC6BAB276524D10DBCEC341B" pageId="0" pageNumber="599" type="nomenclature">
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<paragraph id="C34770CC49CF5CF9AD7B95737F74DF52" pageId="0" pageNumber="599">
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<taxonomicName id="D2D99496A8B0E4BBBBE1D7878FA0F438" ID-CoL="65RBR" ID-ENA="549119" LSID="13A43AC0-0DB0-5E09-B31D-51DE9341AD6C" authority="(Nylander)" baseAuthorityName="Nylander" baseAuthorityYear="1848" class="Insecta" family="Megachilidae" genus="Aglaoapis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglaoapis tridentata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="tridentata">Aglaoapis tridentata (Nylander)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0D2A0D3C43CDBD965411AE53B773F51B" pageId="0" pageNumber="599" type="reference_group">
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<paragraph id="A220FAF9D915AA5388D2D6EE7DAA6937" pageId="0" pageNumber="599">
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<taxonomicName id="8E039E6684B4B5CF9BDB5735F9CEBB44" baseAuthorityName="Fabricius" baseAuthorityYear="1775" class="Insecta" family="Megachilidae" genus="Coelioxys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelioxys tridentata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="tridentata">Coelioxys tridentata</taxonomicName>
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Nylander, 1848 (nec
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<taxonomicName id="C84F2B85E0F098270D7330BA07A0CE70" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Megachilidae" genus="Apis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apis tridentata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="tridentata">Apis tridentata</taxonomicName>
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Fabricius, 1775): 254.
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</paragraph>
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<paragraph id="B15BD457F175184E773D47DD0235B941" pageId="0" pageNumber="599">
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<taxonomicName id="A2E8FE15097D88093B6D0A22A8224FBF" authorityName="Schenck" authorityYear="1861" class="Insecta" family="Megachilidae" genus="Dioxys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioxys fasciata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="fasciata">Dioxys fasciata</taxonomicName>
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Schenck, 1861: 383.
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</paragraph>
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<paragraph id="B8E905D3C389FDE84A44355E5CAFDB3A" pageId="0" pageNumber="599">
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<taxonomicName id="5C134AD5F00A67A7EBCAC64E270D6653" authorityName="Noskiewicz" authorityYear="1940" class="Insecta" family="Megachilidae" genus="Dioxys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioxys kuntzei" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="kuntzei">Dioxys kuntzei</taxonomicName>
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Noskiewicz, 1940: 99.
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</paragraph>
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<paragraph id="16D219B19DE0DD89DBC1B599D0DDC9C4" pageId="0" pageNumber="599">
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<taxonomicName id="CE2DDD5E1F48AEF48BAC888553DD160B" authorityName="Mavromoustakis" authorityYear="1949" class="Insecta" family="Megachilidae" genus="Dioxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioxoides tridentata subsp. limassolica" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="limassolica">Dioxoides tridentata ssp. limassolica</taxonomicName>
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Mavromoustakis, 1949: 587.
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</paragraph>
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</subSubSection>
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<subSubSection id="76834321AAE59F57439893A507CC75BC" pageId="0" pageNumber="599" type="diagnosis">
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<paragraph id="CE09A2D6629EE1008F3E1FC803977B8C" pageId="0" pageNumber="599">Diagnosis.</paragraph>
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<paragraph id="B147701D910BBE236A8F48896FF5644C" pageId="0" pageNumber="599">
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Larger species (9-12 mm), both sexes are black with well-developed white bands of short hair on metasomal terga (Fig.
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<figureCitation id="DB4F14ABD86BA4F27443E90481F4ABD5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aglaoapis tridentata A female, dorsal view B male, dorsal view C female, mesosoma dorsal view D female, metasoma, dorsal view E female, fore coxa F male, T 7, dorsal view. Red scale bars represent the length of 1 mm." figureDoi="10.3897/jhr.96.104957.figure1" httpUri="https://binary.pensoft.net/fig/883548" pageId="0" pageNumber="599">1A, B</figureCitation>
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). Both sexes have the fore coxa with a carina anteriorly (Fig.
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<figureCitation id="5798C33FDD7A555367C71DAF845415B5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aglaoapis tridentata A female, dorsal view B male, dorsal view C female, mesosoma dorsal view D female, metasoma, dorsal view E female, fore coxa F male, T 7, dorsal view. Red scale bars represent the length of 1 mm." figureDoi="10.3897/jhr.96.104957.figure1" httpUri="https://binary.pensoft.net/fig/883548" pageId="0" pageNumber="599">1E</figureCitation>
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) and a short projection and the scutellum with a medial toothlike projection (Fig.
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<figureCitation id="1E6A03436087FA106B50551B19E90113" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aglaoapis tridentata A female, dorsal view B male, dorsal view C female, mesosoma dorsal view D female, metasoma, dorsal view E female, fore coxa F male, T 7, dorsal view. Red scale bars represent the length of 1 mm." figureDoi="10.3897/jhr.96.104957.figure1" httpUri="https://binary.pensoft.net/fig/883548" pageId="0" pageNumber="599">1C</figureCitation>
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). Females have a longer last tergum than females of the genus
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<taxonomicName id="5B1097598C871A3DFF298C3C9FE4F7A3" class="Insecta" family="Megachilidae" genus="Dioxys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioxys" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="genus">
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<emphasis id="F4454B0138AC77D61CA92043D7DC48BE" italics="true" pageId="0" pageNumber="599">Dioxys</emphasis>
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</taxonomicName>
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and an emarginated last sternum (Fig.
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<figureCitation id="63D6172232EA1B4B581549D98A4DB347" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aglaoapis tridentata A female, dorsal view B male, dorsal view C female, mesosoma dorsal view D female, metasoma, dorsal view E female, fore coxa F male, T 7, dorsal view. Red scale bars represent the length of 1 mm." figureDoi="10.3897/jhr.96.104957.figure1" httpUri="https://binary.pensoft.net/fig/883548" pageId="0" pageNumber="599">1D</figureCitation>
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). The last tergum is emarginated for males (Fig.
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<figureCitation id="7B369D9937FF4C46AEBB9FA6CE854A30" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aglaoapis tridentata A female, dorsal view B male, dorsal view C female, mesosoma dorsal view D female, metasoma, dorsal view E female, fore coxa F male, T 7, dorsal view. Red scale bars represent the length of 1 mm." figureDoi="10.3897/jhr.96.104957.figure1" httpUri="https://binary.pensoft.net/fig/883548" pageId="0" pageNumber="599">1F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="2062F0CAF611582F406AC2C66EA4740F" pageId="0" pageNumber="599" type="distribution">
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<paragraph id="E5F6B4F715A9FF3DCA7018FE16853F1D" pageId="0" pageNumber="599">Distribution.</paragraph>
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<paragraph id="C5F4AABFB7AABF92ACB589533E4D62F4" pageId="0" pageNumber="599">
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<taxonomicName id="7F3DB2BBFD8C28BFD471F876981CE703" baseAuthorityName="Nylander" baseAuthorityYear="1848" class="Insecta" family="Megachilidae" genus="Aglaoapis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglaoapis tridentata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="tridentata">
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<emphasis id="C85478D31D3BA4FDEE5E2356F64AA819" italics="true" pageId="0" pageNumber="599">Aglaoapis tridentata</emphasis>
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</taxonomicName>
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is a Palaearctic species that occurs in Europe, from Spain in the west to Russia in the east (Fig.
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<figureCitation id="012AE5974211EFA354B2561E2F55B8B6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aglaoapis tridentata, distribution in Europe." figureDoi="10.3897/jhr.96.104957.figure2" httpUri="https://binary.pensoft.net/fig/883549" pageId="0" pageNumber="599">2</figureCitation>
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), and in Asia from the Caucasus, Kyrgyzstan and China, Kazakhstan, Siberia and Russian Far East. This species reaches the farthest north of any species of the tribe, with records from Finland and Sweden (
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<bibRefCitation id="C56AE1C3C1D55B6EB31BE36E2A930522" DOI="https://doi.org/10.3989/graellsia.2008.v64.i1.55" author="Ornosa, C" journalOrPublisher="Graellsia" pageId="0" pageNumber="599" pagination="61 - 86" refId="B24" refString="Ornosa, C, Ortiz-Sanchez, FJ, Torres, F, 2008. Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Graellsia 64: 61 - 86, DOI: https://doi.org/10.3989/graellsia.2008.v64.i1.55" title="Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini." url="https://doi.org/10.3989/graellsia.2008.v64.i1.55" volume="64" year="2008">Ornosa et al. 2008</bibRefCitation>
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;
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<bibRefCitation id="569671F2F04B5FA3C698AF5F96BB89C5" author="Madsen, HB" journalOrPublisher="Entomologiske Meddelelser" pageId="0" pageNumber="599" pagination="73 - 99" refId="B20" refString="Madsen, HB, Calabuig, I, 2010. Kommenteret checkliste over Danmarks bier - Del 3: Melittidae & Megachilidae (Hymenoptera, Apoidea). Entomologiske Meddelelser 78 (2): 73 - 99" title="Kommenteret checkliste over Danmarks bier - Del 3: Melittidae & Megachilidae (Hymenoptera, Apoidea)." volume="78" year="2010">Madsen and Calabuig 2010</bibRefCitation>
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;
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<bibRefCitation id="F43B12D364F808E2B22D9AD26778AFB0" author="Ascher, JS" pageId="0" pageNumber="599" publicationUrl="http://www.discoverlife.org/" refId="B1" refString="Ascher, JS, Pickering, J, 2023. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). http://www.discoverlife.org/" title="Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila)." url="http://www.discoverlife.org/" year="2023">Ascher and Pickering 2023</bibRefCitation>
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).
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</paragraph>
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<caption id="259F6B6209CDE6AE7270094C3AF2074E" doi="10.3897/jhr.96.104957.figure2" httpUri="https://binary.pensoft.net/fig/883549" pageId="0" pageNumber="599" start="Figure 2" startId="F2">
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<paragraph id="3874329D9A13BD20D3DD49E1A2A89846" pageId="0" pageNumber="599">
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<emphasis id="819A45034C659FCEC9D7F89F18302A0C" bold="true" pageId="0" pageNumber="599">Figure 2.</emphasis>
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<taxonomicName id="ACF49338FE2019B9E980F1911A2CA151" baseAuthorityName="Nylander" baseAuthorityYear="1848" class="Insecta" family="Megachilidae" genus="Aglaoapis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aglaoapis tridentata" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="tridentata">
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<emphasis id="8BD70A0F34F6B767BF0127A000493C24" italics="true" pageId="0" pageNumber="599">Aglaoapis tridentata</emphasis>
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</taxonomicName>
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, distribution in Europe.
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</paragraph>
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</caption>
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<paragraph id="7742B4B6C38210B4DB1EA120E293C1DE" pageId="0" pageNumber="599">
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Biology and hosts: Species recorded especially in steppic formations, sunny slopes, forest steppes and other open or semiopen habitats. Occurs also in abandoned sandpits, spoil heaps and other habitats of anthropogenic origin. This species attacks nests of bees of the family
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<taxonomicName id="7D518F2F74D09E3976377C1825B52DE8" class="Insecta" family="Megachilidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthopoda" rank="family">Megachilidae</taxonomicName>
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, especially those nesting underground or making their own nests near the ground surface.
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<taxonomicName id="3AE433AF1223E1ACD9B2BBF68430B682" baseAuthorityName="Schenck" baseAuthorityYear="1853" class="Insecta" family="Apidae" genus="Hoplitis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hoplitis anthocopoides" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="anthocopoides">
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<emphasis id="BAF6282D3E689CDEE702C73155799122" italics="true" pageId="0" pageNumber="599">Hoplitis anthocopoides</emphasis>
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</taxonomicName>
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(Schenck),
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<taxonomicName id="144FEFE9249857D92901CE534BFFD219" baseAuthorityName="Perez" baseAuthorityYear="1902" class="Insecta" family="Apidae" genus="Hoplitis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hoplitis ravouxi" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="ravouxi">
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<emphasis id="A58924CD6E59F7661F66AF7806A93279" italics="true" pageId="0" pageNumber="599">Hoplitis ravouxi</emphasis>
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</taxonomicName>
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(
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<normalizedToken id="5F62F8D62CB2AAA019933712B85A0FEF" originalValue="Pérez">Perez</normalizedToken>
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), probably also
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<taxonomicName id="33D57CEDA502A34FCB9F09A50EB0AD11" baseAuthorityName="Panzer" baseAuthorityYear="1798" class="Insecta" family="Apidae" genus="Hoplitis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hoplitis adunca" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="adunca">
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<emphasis id="0B9D54E0450831CE408A50F501C4A4F0" italics="true" pageId="0" pageNumber="599">Hoplitis adunca</emphasis>
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</taxonomicName>
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(Panzer) and
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<taxonomicName id="CB272C73E50F318976A7754CB8928743" authorityName="Alfken" authorityYear="1924" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megachile pilidens" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthopoda" rank="species" species="pilidens">
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<emphasis id="8BF414BCBEFF0185C17E4C2D55C00BD1" italics="true" pageId="0" pageNumber="599">Megachile pilidens</emphasis>
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</taxonomicName>
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Alfken, in southern parts of Europe, and also
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<taxonomicName id="70E9D2EEBC22014F54FA2A4CA455B7F0" baseAuthorityName="Geoffroy" baseAuthorityYear="1785" class="Insecta" family="Apidae" genus="Chalicodoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chalicodoma parietina" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthropoda" rank="species" species="parietina">
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<emphasis id="FE6CAEBBAF384EF413CD7F442F0B7EF3" italics="true" pageId="0" pageNumber="599">Chalicodoma parietina</emphasis>
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</taxonomicName>
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(Geoffroy) were confirmed as hosts of this species (
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<bibRefCitation id="DB50388AA70D3024E67A266D141ACD84" author="Westrich, P" journalOrPublisher="Eugen Ulmer, Stuttgart" pageId="0" pageNumber="599" refId="B39" refString="Westrich, P, 2018. Die Wildbienen Deutschlands. Eugen Ulmer, Stuttgart" title="Die Wildbienen Deutschlands." year="2018">Westrich 2018</bibRefCitation>
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).
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<bibRefCitation id="59513D24C27F8F9AEF754FFB1EDB46C5" author="Scheuchl, E" journalOrPublisher="Quelle & Meyer, Leipzig" pageId="0" pageNumber="599" refId="B30" refString="Scheuchl, E, Willner, M, 2016. Taschenlexikon der Wildbienen Mitteleuropas: Alle Arten im Portraet. Quelle & Meyer, Leipzig" title="Taschenlexikon der Wildbienen Mitteleuropas: Alle Arten im Portraet." year="2016">Scheuchl and Willner (2016)</bibRefCitation>
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also listed
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<taxonomicName id="CC35C275A8D6459DAE48619488CDA9DA" authorityName="Curtis" authorityYear="1828" class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megachile leachella" order="Hymenoptera" pageId="0" pageNumber="599" phylum="Arthopoda" rank="species" species="leachella">
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<emphasis id="FF15B93C93524F0900FFB5AFDCF1B7BA" italics="true" pageId="0" pageNumber="599">Megachile leachella</emphasis>
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</taxonomicName>
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Curtis as a likely host.
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</paragraph>
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</subSubSection>
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<subSubSection id="D1D66BBF41C3A7AA218CEDA5FA39D5F2" pageId="0" pageNumber="599" type="conservation status">
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<paragraph id="37EABD4065C65382ECBFE51A436B6128" pageId="0" pageNumber="599">Conservation status.</paragraph>
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<paragraph id="B6E691E1454618C63122C1C223AD9DD3" pageId="0" pageNumber="599">
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<bibRefCitation id="FD5701D570830EC7C6A405970062C4CD" author="Nieto, A" journalOrPublisher="Publication Office of the European Union, Brussels, Belgium" pageId="0" pageNumber="599" refId="B23" refString="Nieto, A, Roberts, SPM, Kemp, J, Rasmont, P, Kuhlmann, M, Garcia Criado, M, Biesmeijer, JC, Bogusch, P, Dathe, HH, De la Rua, P, De Meulemeester, T, Dehon, M, Dewulf, A, Ortiz-Sanchez, FJ, Lhomme, P, Pauly, A, Potts, SG, Praz, C, Quaranta, M, Radchenko, VG, Scheuchl, E, Smit, J, Straka, J, Terzo, M, Tomozei, B, Window, J, Michez, D, 2014. European Red List of Bees. Publication Office of the European Union, Brussels, Belgium" title="European Red List of Bees." year="2014">Nieto et al. (2014)</bibRefCitation>
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classified this species as LC - data deficient. This species is relatively rare throughout its range. Its distribution in Europe is the largest of any species, and it is still being recorded in most countries - it was reported to be regionally extinct only in Belgium and Finland (
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<bibRefCitation id="91B2AC23B8E832CE9C0A9AE89E57E867" author="Ghisbain, G" journalOrPublisher="Memorias de la Real Sociedad espanola de Historia Natural, Tomo del Cincuentenario" pageId="0" pageNumber="599" refId="B8" refString="Ghisbain, G, Rosa, P, Bogusch, P, Flaminio, S, LeDivelec, R, Dorchin, A, Kasparek, M, Litman, J, Mignot, M, Mueller, A, Praz, C, Radchenko, V, Rasmont, P, Smit, J, Wood, TJ, Michez, D, Reverte, S, 2023. The new annotated checklist of the wild bees of Europe (Hymenoptera: Anthophila). Zootaxa, accepted." title="The new annotated checklist of the wild bees of Europe (Hymenoptera: Anthophila). Zootaxa, accepted." year="2023">Ghisbain et al. 2023</bibRefCitation>
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). It should therefore be classified as LC.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |