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<document id="6EB96B5D0D42C4DCD00CFA38A6270DB1" ID-CLB-Dataset="29714" ID-DOI="10.11646/zootaxa.4444.2.2" ID-GBIF-Dataset="f3de278f-f94c-4170-aa4d-addda0a22766" ID-ISSN="1175-5326" ID-Zenodo-Dep="1309536" ID-ZooBank="1A4D6398-EC3A-4279-9846-54EC268CB3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1531313605748" checkinUser="plazi" docAuthor="Oliveira, Verônica Maria De, Jacobsen, Danny Eibye &amp; Lana, Paulo" docDate="2018" docId="137A0A12FFF8FFF20193D362A34CC32B" docLanguage="en" docName="zootaxa.4444.2.2.pdf" docOrigin="Zootaxa 4444 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paranaitis cirriformata Oliveira &amp; Jacobsen &amp; Lana 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="127" masterDocId="EF43726AFFFFFFFA0104D207A62BC26B" masterDocTitle="Four new species of Paranaitis Southern, 1914 (Phyllodocidae, Annelida) from southern and southeastern Brazil" masterLastPageNumber="136" masterPageNumber="120" pageNumber="127" updateTime="1698552899352" updateUser="ExternalLinkService">
<mods:mods id="379903FF0D227F9DE8AB4BEF582564C7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="05305F64DFF4B59DBAA66B9311B13954">Four new species of Paranaitis Southern, 1914 (Phyllodocidae, Annelida) from southern and southeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="BB6A4F9CA6B6AB7666CABE9238A12BF9" type="personal">
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<mods:namePart id="0F4E774C9B8709E910214B4483E545C9">Oliveira, Verônica Maria De</mods:namePart>
</mods:name>
<mods:name id="1F56A3B9C8DCBD4629913FBC9AE5C2FC" type="personal">
<mods:role id="4F83CFC2D4B5E86E438A776CF038FAE4">
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<mods:namePart id="D0A1B71D7B7F1BB747824014F5BD558D">Jacobsen, Danny Eibye</mods:namePart>
</mods:name>
<mods:name id="93E55A7391B980455FFBD3F7D2F16057" type="personal">
<mods:role id="4835F3789D169F358579E809A79F34B1">
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<mods:namePart id="1E74824B36CBCA2931FFFD1E7B28B891">Lana, Paulo</mods:namePart>
</mods:name>
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<mods:title id="5803297CFF1ECD610C92AED7AC9E91F5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4D11C96B690BA996BF946A35B7AD6FD6">
<mods:date id="C9DB7948063D1E523102016F09E34005">2018</mods:date>
<mods:detail id="60CF0124BEBDE9852A60E0BE93526A0B" type="pubDate">
<mods:number id="D0E475AC4A641B71E709D6514E405A1A">2018-07-09</mods:number>
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<mods:number id="69D65C64FC7C32493D3AB36C12058D14">4444</mods:number>
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<mods:classification id="FE36CC7DB28D2AC5FE32546D4A52E5E2">journal article</mods:classification>
<mods:identifier id="AC4105270541D4D36730A5734B14A02C" type="CLB-Dataset">29714</mods:identifier>
<mods:identifier id="BD6E16F2763977B6217580AC17158A39" type="DOI">10.11646/zootaxa.4444.2.2</mods:identifier>
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<mods:identifier id="B354C4FCEBC0E8664DF307960D240531" type="ISSN">1175-5326</mods:identifier>
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<treatment id="137A0A12FFF8FFF20193D362A34CC32B" ID-DOI="http://doi.org/10.5281/zenodo.5951292" ID-GBIF-Taxon="146045046" ID-Zenodo-Dep="5951292" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B" httpUri="http://treatment.plazi.org/id/137A0A12FFF8FFF20193D362A34CC32B" lastPageId="8" lastPageNumber="127" pageId="7" pageNumber="127">
<subSubSection id="D3C9E88FFFF8FFFD0193D362A6D2C3CB" pageId="7" pageNumber="127" type="nomenclature">
<paragraph id="9B6CBB04FFF8FFFD0193D362A43FC314" blockId="7.[151,532,357,416]" box="[151,532,357,383]" pageId="7" pageNumber="127">
<heading id="C0240C68FFF8FFFD0193D362A43FC314" bold="true" box="[151,532,357,383]" fontSize="11" level="1" pageId="7" pageNumber="127" reason="1">
<emphasis id="A9A76716FFF8FFFD0193D362A43FC314" bold="true" box="[151,532,357,383]" pageId="7" pageNumber="127">
<taxonomicName id="5CD3C087FFF8FFFD0193D362A79BC314" ID-CoL="4D4FL" authority="Oliveira &amp; Jacobsen &amp; Lana, 2018" authorityName="Oliveira &amp; Jacobsen &amp; Lana" authorityYear="2018" box="[151,432,357,383]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="cirriformata" status="sp. nov.">
<emphasis id="A9A76716FFF8FFFD0193D362A79BC314" bold="true" box="[151,432,357,383]" italics="true" pageId="7" pageNumber="127">Paranaitis cirriformata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B294DA6DFFF8FFFD00B2D362A43FC314" box="[438,532,357,383]" pageId="7" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B6CBB04FFF8FFFD0193D380A6D2C3CB" blockId="7.[151,532,357,416]" box="[151,249,391,416]" pageId="7" pageNumber="127">
<figureCitation id="03E8A781FFF8FFFD0193D380A6D2C3CB" box="[151,249,391,416]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="8.[151,250,978,1000]" captionStartId-1="8.[151,250,1975,1997]" captionTargetBox-0="[227,1368,369,967]" captionTargetBox-1="[303,1291,1105,1969]" captionTargetId-0="figure@8.[227,1368,369,967]" captionTargetId-1="figure@8.[303,1291,1105,1969]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionText-0="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." captionText-1="FIGURE 9. Paranaitis cirriformata sp. nov. Holotype. AC—Compound spinigerous Chaetae; D—Pygidium with two pairs of anal Cirri. SCale bars: A, 0.01 mm; B, 0.02 mm; C, 0.01 mm; D, 0.25 mm." httpUri-0="https://zenodo.org/record/1309552/files/figure.png" httpUri-1="https://zenodo.org/record/1309554/files/figure.png" pageId="7" pageNumber="127">Figs 89</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFFD0193D3C8A5BBC0D4" pageId="7" pageNumber="127" type="materials_examined">
<paragraph id="9B6CBB04FFF8FFFD0193D3C8A417C066" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<materialsCitation id="2BBBB159FFF8FFFD0193D3C8A417C066" ID-GBIF-Occurrence="1880439276" collectingDate="2009-07-22" country="Brazil" county="Foz" elevation="20" latitude="-21.1715" location="Margin" longLatPrecision="1" longitude="-40.765835" municipality="Hab" pageId="7" pageNumber="127" specimenCount="1" typeStatus="holotype">
<emphasis id="A9A76716FFF8FFFD0193D3C8A720C383" bold="true" box="[151,267,463,488]" pageId="7" pageNumber="127">
<typeStatus id="446805A6FFF8FFFD0193D3C8A72DC383" box="[151,262,463,488]" pageId="7" pageNumber="127" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="9E0CEDDFFFF8FFFD0016D3C8A74FC383" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B:9E0CEDDFFFF8FFFD0016D3C8A74FC383" box="[274,356,463,488]" country="Brazil" county="Foz" latitude="-21.1715" longLatPrecision="1" longitude="-40.765835" municipality="Hab" name="Margin" pageId="7" pageNumber="127">Margin</location>
of continental shelf,
<location id="9E0CEDDFFFF8FFFD0350D3D7A4D1C383" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B:9E0CEDDFFFF8FFFD0350D3D7A4D1C383" box="[596,762,463,488]" country="Brazil" county="Foz" latitude="-21.1715" longLatPrecision="1" longitude="-40.765835" municipality="Hab" name="Campos Basin" pageId="7" pageNumber="127">Campos Basin</location>
,
<collectingCountry id="E3C4FB94FFF8FFFD0203D3C8A565C383" box="[775,846,463,488]" name="Brazil" pageId="7" pageNumber="127">Brazil</collectingCountry>
,
<collectingMunicipality id="7B08217EFFF8FFFD025DD3C8A5A1C383" box="[857,906,463,488]" pageId="7" pageNumber="127">Hab</collectingMunicipality>
17
<collectingCounty id="720DC388FFF8FFFD02A8D3C8A5F3C383" box="[940,984,463,488]" pageId="7" pageNumber="127">Foz</collectingCounty>
15 R01,
<geoCoordinate id="FEE7DDC3FFF8FFFD0533D3D7A2E0C383" box="[1079,1227,464,488]" direction="south" orientation="latitude" pageId="7" pageNumber="127" precision="1" value="-21.1715">21°10'17.4&quot;S</geoCoordinate>
,
<geoCoordinate id="FEE7DDC3FFF8FFFD05D3D3C8A35FC383" box="[1239,1396,463,488]" direction="west" orientation="longitude" pageId="7" pageNumber="127" precision="1" value="-40.765835">40°45'57.0&quot;W</geoCoordinate>
,
<elevation id="10FE5C37FFF8FFFD0484D3D7A684C067" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="127" unit="m" value="20.0">
<quantity id="5C2B16E1FFF8FFFD0484D3D7A684C067" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="127" unit="m" value="20.0">20 m</quantity>
</elevation>
,
<collectingDate id="FF29642CFFF8FFFD01BFD3F3A76FC067" box="[187,324,500,525]" pageId="7" pageNumber="127" value="2009-07-22">
<date id="EF6D9DC4FFF8FFFD01BFD3F3A76FC067" box="[187,324,500,525]" pageId="7" pageNumber="127" value="2009-07-22">22 Jul. 2009</date>
</collectingDate>
(ZUEC-POL-16339).
</materialsCitation>
</paragraph>
<paragraph id="9B6CBB04FFF8FFFD01C3D010A5BBC0D4" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<materialsCitation id="2BBBB159FFF8FFFD01C3D010A790C03F" ID-GBIF-Occurrence="1880439280" country="Brazil" county="Port Paranagua" location="Paranagua Bay" municipality="In" pageId="7" pageNumber="127" specimenCount="2" stateProvince="Parana" typeStatus="paratype">
<emphasis id="A9A76716FFF8FFFD01C3D010A76DC05B" bold="true" box="[199,326,535,560]" pageId="7" pageNumber="127">
<typeStatus id="446805A6FFF8FFFD01C3D010A76AC05B" box="[199,321,535,560]" pageId="7" pageNumber="127" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="8DD5708DFFF8FFFD0048D01FA7E5C05B" box="[332,462,536,560]" pageId="7" pageNumber="127" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
, length ranging from
<quantity id="5C2B16E1FFF8FFFD03BAD01FA503C044" box="[702,808,536,560]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="7" pageNumber="127" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">411 mm</quantity>
for 36104 segments.
<location id="9E0CEDDFFFF8FFFD0521D01FA2E5C05B" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B:9E0CEDDFFFF8FFFD0521D01FA2E5C05B" box="[1061,1230,536,560]" country="Brazil" county="Port Paranagua" municipality="In" name="Paranagua Bay" pageId="7" pageNumber="127" stateProvince="Parana">Paranaguá Bay</location>
,
<collectingRegion id="591775E6FFF8FFFD05DDD01FA303C05B" box="[1241,1320,536,560]" country="Brazil" name="Parana" pageId="7" pageNumber="127">Paraná</collectingRegion>
,
<collectingCountry id="E3C4FB94FFF8FFFD0436D010A350C05B" box="[1330,1403,535,560]" name="Brazil" pageId="7" pageNumber="127">Brazil</collectingCountry>
:
<collectingMunicipality id="7B08217EFFF8FFFD0481D01FA3B7C044" box="[1413,1436,536,559]" pageId="7" pageNumber="127">In</collectingMunicipality>
front of
<collectingCounty id="720DC388FFF8FFFD0001D03AA790C03F" box="[261,443,572,596]" pageId="7" pageNumber="127">Port Paranaguá</collectingCounty>
</materialsCitation>
,
<materialsCitation id="2BBBB159FFF8FFFD00D4D03BA4C5C013" ID-GBIF-Occurrence="1880439285" collectingDate="2011-08" country="Brazil" county="Foz" latitude="-25.511555" location="Continental" longLatPrecision="1" longitude="-48.497307" municipality="Hab" pageId="7" pageNumber="127" specimenCount="1" stateProvince="Parana" typeStatus="paratype">
<geoCoordinate id="FEE7DDC3FFF8FFFD00D4D03BA44CC03E" box="[464,615,572,597]" direction="south" orientation="latitude" pageId="7" pageNumber="127" precision="1" value="-25.511555">25°30'41.6&quot;S</geoCoordinate>
,
<geoCoordinate id="FEE7DDC3FFF8FFFD037FD03BA531C03E" box="[635,794,572,597]" direction="west" orientation="longitude" pageId="7" pageNumber="127" precision="1" value="-48.497307">48°29'50.3&quot;W</geoCoordinate>
, intertidal,
<collectingDate id="FF29642CFFF8FFFD02AFD03BA201C03F" box="[939,1066,572,596]" pageId="7" pageNumber="127" value="2011-08">
<date id="EF6D9DC4FFF8FFFD02AFD03BA201C03F" box="[939,1066,572,596]" pageId="7" pageNumber="127" value="2011-08">Aug. 2011</date>
</collectingDate>
(
<specimenCount id="8DD5708DFFF8FFFD0541D03BA2EFC03E" box="[1093,1220,572,597]" pageId="7" pageNumber="127" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, NHMD-207863).
<location id="9E0CEDDFFFF8FFFD0193D058A731C013" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B:9E0CEDDFFFF8FFFD0193D058A731C013" box="[151,282,607,632]" country="Brazil" county="Foz" latitude="-25.511555" longLatPrecision="1" longitude="-48.497307" municipality="Hab" name="Continental" pageId="7" pageNumber="127" stateProvince="Parana">Continental</location>
shelf of
<location id="9E0CEDDFFFF8FFFD0078D067A409C013" LSID="urn:lsid:plazi:treatment:137A0A12FFF8FFF20193D362A34CC32B:9E0CEDDFFFF8FFFD0078D067A409C013" box="[380,546,607,632]" country="Brazil" county="Foz" latitude="-25.511555" longLatPrecision="1" longitude="-48.497307" municipality="Hab" name="Campos Basin" pageId="7" pageNumber="127" stateProvince="Parana">Campos Basin</location>
:
<collectingMunicipality id="7B08217EFFF8FFFD0329D058A475C013" box="[557,606,607,632]" pageId="7" pageNumber="127">Hab</collectingMunicipality>
17
<collectingCounty id="720DC388FFF8FFFD037AD058A480C013" box="[638,683,607,632]" pageId="7" pageNumber="127">Foz</collectingCounty>
15 R1
</materialsCitation>
, 21°10'17.4&quot;S, 40°45'57.0&quot;W,
<quantity id="5C2B16E1FFF8FFFD0544D067A250C01C" box="[1088,1147,608,632]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="127" unit="m" value="20.0">20 m</quantity>
,
<date id="EF6D9DC4FFF8FFFD0583D067A33BC013" box="[1159,1296,607,632]" pageId="7" pageNumber="127" value="2009-07-22">22 Jul. 2009</date>
(1 paratype, ZUEC-POL-16340). Collected in silt-clay, poorly sorted bottoms on the slope and in coarse sand, sorted bottoms on the continental shelf from the intertidal down to a depth of
<quantity id="5C2B16E1FFF8FFFD0254D0A0A5A7C0D4" box="[848,908,679,704]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="7" pageNumber="127" unit="m" value="52.0">52 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFFD01C3D0CBA7B7C13B" pageId="7" pageNumber="127" type="diagnosis">
<paragraph id="9B6CBB04FFF8FFFD01C3D0CBA7B7C13B" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D0CBA76AC08E" bold="true" box="[199,321,716,741]" pageId="7" pageNumber="127">Diagnosis.</emphasis>
Presence of a pair of cavities or fissures between prostomium and peristomium. Proximal part of proboscis with lateral papillae and dorsal chitinous papillae; 4 longitudinal rows of tubercles, 2 pairs on each side, gradually forming rows of tubercles, terminal part with proboscideal ring with oval papillae. One pair of long, cirriform pygidial cirri.
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFFD01C3D15BA4E2C4DB" pageId="7" pageNumber="127" type="description">
<paragraph id="9B6CBB04FFF8FFFD01C3D15BA5E5C4E7" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D15BA772C11E" bold="true" box="[199,345,860,885]" pageId="7" pageNumber="127">Description.</emphasis>
<typeStatus id="446805A6FFF8FFFD005BD15BA7ECC11E" box="[351,455,860,885]" pageId="7" pageNumber="127" type="holotype">Holotype</typeStatus>
complete specimen,
<quantity id="5C2B16E1FFF8FFFD03B7D15BA529C11F" box="[691,770,860,884]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="127" unit="mm" value="10.0">10 mm</quantity>
long for 61 segments,
<quantity id="5C2B16E1FFF8FFFD0505D15BA27DC11F" box="[1025,1110,860,884]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="127" unit="mm" value="1.4">1.4 mm</quantity>
wide at median part of body, including parapodia and excluding chaetae. Body long, dorso-ventrally flattened, of uniform width, tapering at posterior end. Prostomium anteriorly rounded, wider than long, with ligule creating distinct depression in posterior part, where nuchal papilla inseres, resembling a rounded button (
<figureCitation id="03E8A781FFF8FFFD026CD1C0A5C8C18B" box="[872,995,967,992]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8AB</figureCitation>
). Nuchal organs present on each side of prostomium (
<figureCitation id="03E8A781FFF8FFFD0037D1EBA7BAC66F" box="[307,401,1004,1029]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8B</figureCitation>
). Paired frontal, conical and robust antennae and palps, of equal size, postero-laterally positioned on prostomium. Antennae and palps 1/4 as long as prostomium (
<figureCitation id="03E8A781FFF8FFFD02E0D608A274C643" box="[996,1119,1039,1064]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8AB</figureCitation>
). A pair of brown eyes, subepidermal, with lenses, located at posterior margin of prostomium. Proximal part of proboscis with lateral papillae and dorsal chitinous papillae; four longitudinal rows of tubercles, two pairs of rows on each side, terminal part of proboscideal ring with oval papillae (
<figureCitation id="03E8A781FFF8FFFD034FD67BA48DC6FF" box="[587,678,1148,1173]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8C</figureCitation>
). Terminal ring of proboscis with oval papillae. First and second segments dorso-ventrally fused as a collar, laterally with pair of cavities at border of prostomium (
<figureCitation id="03E8A781FFF8FFFD05FCD698A37AC6D3" box="[1272,1361,1183,1208]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8B</figureCitation>
). Four pairs of cylindrical tentacular cirri, biarticulate, with short cirrophores and long cirrostyles, arranged on first three segments. Cirri of segment 1 reaching segment 6. Dorsal and ventral cirri of segment 2 reaching segments 13 and 5, respectively. Dorsal cirri of segment 3 extending to segment 10. Aciculae not observed in segment 2. Neuropodia and ventral cirri from segment 3. Dorsal cirri symmetrical, with asymmetrical, well-developed cirrophores without external extension, rounded, from segment 4. Dorsal cirri longer than wide in anterior segments, clearly wider than long in median segments, and almost as wide as long in posterior segments. Parapodial lobes shorter than dorsal cirri and longer than ventral cirri, with aciculae and neurochaetae. Prechaetal lobes bilobate, asymmetrical and rounded, supraacicular lobe twice as wide as subacicular lobe. Ventral cirri horizontally oriented in relation to parapodial lobes, asymmetrical, on anterior segments oval with rounded edges, on median and posterior segments longer (
<figureCitation id="03E8A781FFF8FFFD0375D400A4C2C44B" box="[625,745,1543,1568]" captionStart="FIGURE 8" captionStartId="8.[151,250,978,1000]" captionTargetBox="[227,1368,369,967]" captionTargetId="figure@8.[227,1368,369,967]" captionTargetPageId="8" captionText="FIGURE 8. Paranaitis cirriformata sp. nov. AB—Prostomium, dorsal view; C—Everted probosCis, dorsal view; D— Parapodium from segment 12; E—Parapodium from segment 37; F—Parapodium from segment 57.. SCale bars: A, 0.8 mm; B, 0.5 mm; C, 0.4 mm; DE 0.1 mm, C, 0.4 mm.." httpUri="https://zenodo.org/record/1309552/files/figure.png" pageId="7" pageNumber="127">Fig. 8DF</figureCitation>
). Compound spinigerous chaetae from segment 3. Rostrum of chaetal shaft asymmetrical, with large, conical main fang, base of fang surrounded by small cylindrical teeth of unequal length; chaetal articles with serrated outer edges (
<figureCitation id="03E8A781FFF8FFFD0220D448A58BC403" box="[804,928,1615,1640]" captionStart="FIGURE 9" captionStartId="8.[151,250,1975,1997]" captionTargetBox="[303,1291,1105,1969]" captionTargetId="figure@8.[303,1291,1105,1969]" captionTargetPageId="8" captionText="FIGURE 9. Paranaitis cirriformata sp. nov. Holotype. AC—Compound spinigerous Chaetae; D—Pygidium with two pairs of anal Cirri. SCale bars: A, 0.01 mm; B, 0.02 mm; C, 0.01 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/1309554/files/figure.png" pageId="7" pageNumber="127">Fig. 9AC</figureCitation>
). Pygidium with two pairs of cylindrical anal cirri, cirriform and slender. Cylindrical pygidial papilla present (
<figureCitation id="03E8A781FFF8FFFD0263D473A5E9C4E7" box="[871,962,1652,1677]" captionStart="FIGURE 9" captionStartId="8.[151,250,1975,1997]" captionTargetBox="[303,1291,1105,1969]" captionTargetId="figure@8.[303,1291,1105,1969]" captionTargetPageId="8" captionText="FIGURE 9. Paranaitis cirriformata sp. nov. Holotype. AC—Compound spinigerous Chaetae; D—Pygidium with two pairs of anal Cirri. SCale bars: A, 0.01 mm; B, 0.02 mm; C, 0.01 mm; D, 0.25 mm." httpUri="https://zenodo.org/record/1309554/files/figure.png" pageId="7" pageNumber="127">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="9B6CBB04FFF8FFFD01C3D490A4E2C4DB" blockId="7.[151,1437,463,2037]" box="[199,713,1687,1712]" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D490A764C4C4" bold="true" box="[199,335,1687,1712]" pageId="7" pageNumber="127">Coloration.</emphasis>
Preserved specimens are opaque.
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFFD01C3D4BBA499C493" pageId="7" pageNumber="127" type="distribution">
<paragraph id="9B6CBB04FFF8FFFD01C3D4BBA499C493" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D4BBA74BC4BE" bold="true" box="[199,352,1724,1749]" pageId="7" pageNumber="127">Distribution.</emphasis>
Atlantic Ocean, Brazilian continental margin; areas of the continental shelf in the Campos Basin. Itiberê River in Paranaguá Bay, Paraná—Brazil.
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFFD01C3D503A44CC52B" pageId="7" pageNumber="127" type="etymology">
<paragraph id="9B6CBB04FFF8FFFD01C3D503A44CC52B" blockId="7.[151,1437,463,2037]" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D503A765C576" bold="true" box="[199,334,1796,1821]" pageId="7" pageNumber="127">Etymology.</emphasis>
The name of this species is derived from the Latin word
<emphasis id="A9A76716FFF8FFFD02E8D502A204C577" box="[1004,1071,1797,1820]" italics="true" pageId="7" pageNumber="127">cirrus</emphasis>
(cirrus) and
<emphasis id="A9A76716FFF8FFFD05C2D503A325C577" box="[1222,1294,1796,1820]" italics="true" pageId="7" pageNumber="127">formis</emphasis>
(shape) and refers to the morphology of the anal cirri.
</paragraph>
</subSubSection>
<subSubSection id="D3C9E88FFFF8FFF201C3D54BA34CC32B" lastPageId="8" lastPageNumber="128" pageId="7" pageNumber="127" type="discussion">
<paragraph id="9B6CBB04FFF8FFF201C3D54BA34CC32B" blockId="7.[151,1437,463,2037]" lastBlockId="8.[151,1436,151,320]" lastPageId="8" lastPageNumber="128" pageId="7" pageNumber="127">
<emphasis id="A9A76716FFF8FFFD01C3D54BA710C50E" bold="true" box="[199,315,1868,1893]" pageId="7" pageNumber="127">Remarks.</emphasis>
The small number of specimens, almost all fragmented, hindered the careful observation of the pygidial region. We observed two pairs of pygidial cirri in the
<typeStatus id="446805A6FFF8FFFD0262D568A5E3C5E3" box="[870,968,1903,1928]" pageId="7" pageNumber="127" type="holotype">holotype</typeStatus>
(ZUEC-POL-16339), but we do not rule out the possibility that this is a morphological anomaly.
<taxonomicName id="5CD3C087FFF8FFFD0214D592A20AC5C7" authority="Oliveira &amp; Jacobsen &amp; Lana, 2018" authorityName="Oliveira &amp; Jacobsen &amp; Lana" authorityYear="2018" box="[784,1057,1940,1964]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="cordiformata" status="sp. nov.">
<emphasis id="A9A76716FFF8FFFD0214D592A20AC5C7" box="[784,1057,1940,1964]" italics="true" pageId="7" pageNumber="127">Paranaitis cordiformata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B294DA6DFFF8FFFD052DD593A2A8C5C0" box="[1065,1155,1940,1965]" pageId="7" pageNumber="127" rank="species">
<emphasis id="A9A76716FFF8FFFD052DD593A2A8C5C0" bold="true" box="[1065,1155,1940,1965]" pageId="7" pageNumber="127">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="5CD3C087FFF8FFFD041DD592A7FAC5BA" authority="(Hartmann-Schroder, 1975)" authorityName="Hartmann-Schroder" authorityYear="1975" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1975" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="abyssalis">
<emphasis id="A9A76716FFF8FFFD041DD592A3B7C5C7" box="[1305,1436,1940,1964]" italics="true" pageId="7" pageNumber="127">P. abyssalis</emphasis>
(
<bibRefCitation id="FF42C6F5FFF8FFFD019AD5BFA7E3C5BA" box="[158,456,1976,2001]" pageId="7" pageNumber="127" refId="ref6907" refString="Hartmann-SChroder, G. (1975) PolyChaeten der IberisChen Tiefsee, gesammelt auf der 3. Reise der Meteor im Jahre 1966. Mitteilungen aus dem Hamburgischen zoologischen Museum und Institut, 72, 47 - 73." type="journal article">Hartmann-Schröder, 1975</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="5CD3C087FFF8FFFD00DBD5BEA458C5BB" authorityName="Knox" authorityYear="1960" box="[479,627,1976,2000]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="benthicola">
<emphasis id="A9A76716FFF8FFFD00DBD5BEA458C5BB" box="[479,627,1976,2000]" italics="true" pageId="7" pageNumber="127">P. benthicola</emphasis>
</taxonomicName>
,
<taxonomicName id="5CD3C087FFF8FFFD0387D5BEA4D2C5BB" baseAuthorityName="Benham" baseAuthorityYear="1927" box="[643,761,1976,2000]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="bowersi">
<emphasis id="A9A76716FFF8FFFD0387D5BEA4D2C5BB" box="[643,761,1976,2000]" italics="true" pageId="7" pageNumber="127">P. bowersi</emphasis>
</taxonomicName>
,
<taxonomicName id="5CD3C087FFF8FFFD020ED5BEA543C5BB" baseAuthorityName="Moore" baseAuthorityYear="1903" box="[778,872,1977,2000]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="caeca">
<emphasis id="A9A76716FFF8FFFD020ED5BEA543C5BB" box="[778,872,1977,2000]" italics="true" pageId="7" pageNumber="127">P. caeca</emphasis>
</taxonomicName>
, and
<taxonomicName id="5CD3C087FFF8FFFD02AED5BEA213C5BB" authorityName="Eibye-Jacobsen" authorityYear="1991" box="[938,1080,1976,2000]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="uschakovi">
<emphasis id="A9A76716FFF8FFFD02AED5BEA213C5BB" box="[938,1080,1976,2000]" italics="true" pageId="7" pageNumber="127">P. uschakovi</emphasis>
</taxonomicName>
in having eyes, but also in the morphology of the dorsal cirri, which are almost circular in
<taxonomicName id="5CD3C087FFF8FFFD0234D5DAA599C59F" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1975" box="[816,946,2012,2036]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="abyssalis">
<emphasis id="A9A76716FFF8FFFD0234D5DAA599C59F" box="[816,946,2012,2036]" italics="true" pageId="7" pageNumber="127">P. abyssalis</emphasis>
</taxonomicName>
and
<taxonomicName id="5CD3C087FFF8FFFD02E3D5DAA253C59F" authorityName="Knox" authorityYear="1960" box="[999,1144,2012,2036]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="benthicola">
<emphasis id="A9A76716FFF8FFFD02E3D5DAA253C59F" box="[999,1144,2012,2036]" italics="true" pageId="7" pageNumber="127">P. benthicola</emphasis>
</taxonomicName>
, and reniform in
<emphasis id="A9A76716FFF8FFF2043DD5DAA722C2DB" italics="true" lastPageId="8" lastPageNumber="128" pageId="7" pageNumber="127">
<taxonomicName id="5CD3C087FFF8FFFD043DD5DAA3B3C59F" baseAuthorityName="Moore" baseAuthorityYear="1903" box="[1337,1432,2013,2036]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="127" phylum="Annelida" rank="species" species="caeca">P. caeca</taxonomicName>
,
<taxonomicName id="5CD3C087FFF7FFF20193D29EA722C2DB" baseAuthorityName="Benham" baseAuthorityYear="1927" box="[151,265,152,176]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="bowersi">P. bowersi</taxonomicName>
</emphasis>
and
<taxonomicName id="5CD3C087FFF7FFF20041D29EA7CBC2DB" box="[325,480,152,176]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="usachakovi">
<emphasis id="A9A76716FFF7FFF20041D29EA7CBC2DB" box="[325,480,152,176]" italics="true" pageId="8" pageNumber="128">P. usachakovi</emphasis>
</taxonomicName>
.
<taxonomicName id="5CD3C087FFF7FFF200EBD29FA4D3C2DB" authority="Oliveira &amp; Jacobsen &amp; Lana, 2018" authorityName="Oliveira &amp; Jacobsen &amp; Lana" authorityYear="2018" box="[495,760,152,176]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="cirriformata" status="sp. nov.">
<emphasis id="A9A76716FFF7FFF200EBD29FA4D3C2DB" box="[495,760,152,176]" italics="true" pageId="8" pageNumber="128">Paranaitis cirriformata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B294DA6DFFF7FFF20204D290A571C2DB" box="[768,858,151,176]" pageId="8" pageNumber="128" rank="species">
<emphasis id="A9A76716FFF7FFF20204D290A571C2DB" bold="true" box="[768,858,151,176]" pageId="8" pageNumber="128">sp. nov.</emphasis>
</taxonomicNameLabel>
shares the rounded median dorsal cirri, wider than long, with
<emphasis id="A9A76716FFF7FFF20017D2BAA7BCC2BF" box="[275,407,189,212]" italics="true" pageId="8" pageNumber="128">
<taxonomicName id="5CD3C087FFF7FFF20017D2BAA7B8C2BF" baseAuthorityName="Webster" baseAuthorityYear="1879" box="[275,403,189,212]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="speciosa">P. speciosa</taxonomicName>
,
</emphasis>
but they differ in the presence of a rounded proboscideal papillae in, asymmetrical dorsal cirrophores and cylindrical anal cirri in the first.
<taxonomicName id="5CD3C087FFF7FFF203BCD2E6A574C293" authority="Oliveira &amp; Jacobsen &amp; Lana, 2018" authorityName="Oliveira &amp; Jacobsen &amp; Lana" authorityYear="2018" box="[696,863,224,248]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="cirriformata" status="sp. nov.">
<emphasis id="A9A76716FFF7FFF203BCD2E6A574C293" box="[696,863,224,248]" italics="true" pageId="8" pageNumber="128">P. cirriformata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B294DA6DFFF7FFF20261D2D8A594C293" box="[869,959,223,248]" pageId="8" pageNumber="128" rank="species">
<emphasis id="A9A76716FFF7FFF20261D2D8A594C293" bold="true" box="[869,959,223,248]" pageId="8" pageNumber="128">sp. nov.</emphasis>
</taxonomicNameLabel>
iffers from the other species of
<taxonomicName id="5CD3C087FFF7FFF20421D2E7A3B0C29C" authorityName="Southern" authorityYear="1914" box="[1317,1435,224,247]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="genus">
<emphasis id="A9A76716FFF7FFF20421D2E7A3B0C29C" box="[1317,1435,224,247]" italics="true" pageId="8" pageNumber="128">Paranaitis</emphasis>
</taxonomicName>
by the presence of nuchal organ on each side of the prostomium (
<bibRefCitation id="FF42C6F5FFF7FFF20283D303A25CC377" box="[903,1143,260,285]" pageId="8" pageNumber="128" refId="ref7007" refString="Kato, T. &amp; Pleijel, F. (2003) A revision of Paranaitis Southern, 1914 (PolyChaeta: PhyllodoCidae). Zoological Journal of the Linnean Society, 138, 379 - 429. https: // doi. org / 10.1046 / j. 1096 - 3642.2003.00069. x" type="journal article">Kato &amp; Pleijel, 2003</bibRefCitation>
) and cylindrical and long anal cirri. The anal cirri of
<taxonomicName id="5CD3C087FFF7FFF200C1D32EA44EC32B" baseAuthorityName="Malmgren" baseAuthorityYear="1867" box="[453,613,296,320]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="kosteriensis">
<emphasis id="A9A76716FFF7FFF200C1D32EA44EC32B" box="[453,613,296,320]" italics="true" pageId="8" pageNumber="128">P. kosteriensis</emphasis>
</taxonomicName>
are also cylindrical but much shorter than in
<taxonomicName id="5CD3C087FFF7FFF20565D32EA32CC32B" authority="Oliveira &amp; Jacobsen &amp; Lana, 2018" authorityName="Oliveira &amp; Jacobsen &amp; Lana" authorityYear="2018" box="[1121,1287,296,320]" class="Polychaeta" family="Phyllodocidae" genus="Paranaitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="128" phylum="Annelida" rank="species" species="cirriformata" status="sp. nov.">
<emphasis id="A9A76716FFF7FFF20565D32EA32CC32B" box="[1121,1287,296,320]" italics="true" pageId="8" pageNumber="128">P. cirriformata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B294DA6DFFF7FFF20409D320A34CC32B" box="[1293,1383,295,320]" pageId="8" pageNumber="128" rank="species">
<emphasis id="A9A76716FFF7FFF20409D320A34CC32B" bold="true" box="[1293,1383,295,320]" pageId="8" pageNumber="128">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>