395 lines
45 KiB
XML
395 lines
45 KiB
XML
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<mods:title id="40F790C4DEDA171FE750D9A5681BCEB3">A taxonomic revision of the Malagasy genus Emphania Erichson, 1847 (Insecta, Coleoptera, Scarabaeidae, Sericini)</mods:title>
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<mods:affiliation id="9A6105A4124FA534A672E7B83BC4DAA4">The Natural History Museum, Department of Entomology, Cromwell Road, London SW 7 5 BD (United Kingdom) d. ahrens @ nhm. ac. uk</mods:affiliation>
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–
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. —
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<emphasis id="A989191FFFFCFFBFFF9357C2A44FFE7D" box="[113,267,421,445]" italics="true" pageId="3" pageNumber="920">Emphania chloris</emphasis>
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. —
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; synonymized by
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. —
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: “
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Cat. Mus.
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[blue label]/
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(
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“
|
||
<location id="9E2293D6FFFCFFBFFE1E5443A7C2FDFC" LSID="urn:lsid:plazi:treatment:1354741BFFFCFFB9FF4A56B0A409FE92:9E2293D6FFFCFFBFFE1E5443A7C2FDFC" box="[508,646,548,572]" country="Madagascar" municipality="Goudot" name="Museum Paris" pageId="3" pageNumber="920">Muséum Paris</location>
|
||
<collectingCountry id="E3EA859DFFFCFFBFFF925458A59BFD97" box="[112,223,575,599]" name="Madagascar" pageId="3" pageNumber="920">Madagascar</collectingCountry>
|
||
<collectingMunicipality id="7B265F77FFFCFFBFFF015458A46AFD97" box="[227,302,575,599]" pageId="3" pageNumber="920">Goudot</collectingMunicipality>
|
||
[blue label]/477 34”,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE155458A5C4FDB1" pageId="3" pageNumber="920" type="female" typeStatus="paralectotype">1 paralectotype ♀</specimenCount>
|
||
(
|
||
<collectionCode id="FDEC5DC8FFFCFFBFFF73543EA5ADFDB1" box="[145,233,601,625]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="920">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3E79686FFFCFFBFFF935414A745FA7A" pageId="3" pageNumber="920" type="reference_group">
|
||
<paragraph id="9B42C50DFFFCFFBFFF935414A704FD35" blockId="3.[111,647,495,757]" pageId="3" pageNumber="920">
|
||
<taxonomicName id="5CFDBE8EFFFCFFBFFF935414A488FD4B" authority="Burmeister, 1855" authorityName="Burmeister" authorityYear="1855" box="[113,460,627,651]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="920" phylum="Arthropoda" rank="species" species="chloris">
|
||
<emphasis id="A989191FFFFCFFBFFF935414A452FD4B" box="[113,278,627,651]" italics="true" pageId="3" pageNumber="920">Emphania chloris</emphasis>
|
||
<materialsCitation id="2B95CF50FFFCFFBFFEFC5414A488FD4B" box="[286,460,627,651]" collectionCode="ZMHB" pageId="3" pageNumber="920" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
|
||
<bibRefCitation id="FF6CB8FCFFFCFFBFFEFC5414A488FD4B" author="BURMEISTER H." box="[286,460,627,651]" pageId="3" pageNumber="920" refId="ref6446" refString="BURMEISTER H. 1855. - Handbuch der Entomologie. 4. Band. Besondere Entomologie, Fortsetzung. 2. Abteilung, Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Fr. Enslin, Berlin, 467 p." type="book" year="1855">Burmeister, 1855</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
: “24966/
|
||
<taxonomicName id="5CFDBE8EFFFCFFBFFDCB5414A44FFD66" authority="N. Madag. Goud." authorityName="N. Madag. Goud." class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="920" phylum="Arthropoda" rank="species" species="chloris">
|
||
<emphasis id="A989191FFFFCFFBFFDCB5414A720FD4B" box="[553,612,627,651]" italics="true" pageId="3" pageNumber="920">chloris</emphasis>
|
||
N. Madag. Goud.
|
||
</taxonomicName>
|
||
[blue label]”,
|
||
<typeStatus id="44467BAFFFFCFFBFFE4054E9A4BBFD66" box="[418,511,654,678]" pageId="3" pageNumber="920" type="lectotype">lectotype</typeStatus>
|
||
<specimenCount id="8DFB0E84FFFCFFBFFDE454E8A75CFD66" box="[518,536,655,678]" pageId="3" pageNumber="920" type="female">♀</specimenCount>
|
||
(ZMHB);
|
||
<materialsCitation id="2B95CF50FFFCFFBFFF9154CFA49CFD00" box="[115,472,680,705]" collectionCode="ZMHB" isTailEnumeration="true" pageId="3" pageNumber="920" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
|
||
“24966”,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF3354CFA432FD01" box="[209,374,680,705]" pageId="3" pageNumber="920" type="female" typeStatus="paralectotype">1 paralectotype ♀</specimenCount>
|
||
(
|
||
<collectionCode id="FDEC5DC8FFFCFFBFFE6054CFA494FD00" box="[386,464,680,704]" pageId="3" pageNumber="920">ZMHB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2B95CF50FFFCFFBFFE0254CFA778FD35" collectionCode="WB, MLUH" pageId="3" pageNumber="920" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
|
||
“MLU Halle
|
||
<collectionCode id="FDEC5DC8FFFCFFBFFDBD54CFA7C2FD00" box="[607,646,680,704]" country="USA" name="Department of Bacteriology, University of Wisconsin" pageId="3" pageNumber="920">WB</collectionCode>
|
||
Zoologie S.-Nr. 8/3/12 [white label]/
|
||
<taxonomicName id="5CFDBE8EFFFCFFBFFE3B54A4A5DBFD35" authority="Kl. Madag. MB." authorityName="Kl. Madag. MB." class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="920" phylum="Arthropoda" rank="species" species="chloris">
|
||
<emphasis id="A989191FFFFCFFBFFE3B54A4A750FD1B" box="[473,532,707,731]" italics="true" pageId="3" pageNumber="920">chloris</emphasis>
|
||
Kl. Madag. MB.
|
||
</taxonomicName>
|
||
[blue label]”,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFECB54BAA492FD35" box="[297,470,733,757]" pageId="3" pageNumber="920" type="female" typeStatus="paralectotype">1 paralectotype ♀</specimenCount>
|
||
(
|
||
<collectionCode id="FDEC5DC8FFFCFFBFFE0654BAA771FD35" box="[484,565,733,757]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="3" pageNumber="920">MLUH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFF935569A745FA7A" blockId="3.[111,649,781,1466]" pageId="3" pageNumber="920">
|
||
ADDITIONAL MATERIAL EXAMINED (list not all sexed). —
|
||
<emphasis id="A989191FFFFCFFBFFF935540A5A8FCFF" bold="true" box="[113,236,807,831]" pageId="3" pageNumber="920">
|
||
<collectingCountry id="E3EA859DFFFCFFBFFF935540A5A3FCFF" box="[113,231,807,831]" name="Madagascar" pageId="3" pageNumber="920">Madagascar</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
Forêt de la Mandraka,
|
||
<date id="EF43E3CDFFFCFFBFFE2B5540A762FCFF" box="[457,550,807,831]" pageId="3" pageNumber="920" value="1946-12">XII.1946</date>
|
||
, J. Millot,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF935525A5E4FC9A" box="[113,160,834,858]" pageId="3" pageNumber="920" type="generic">1 ex.</specimenCount>
|
||
(MNHN). — La Mandraka,
|
||
<date id="EF43E3CDFFFCFFBFFE5D5525A769FC9A" box="[447,557,834,858]" pageId="3" pageNumber="920" value="1946-12-04">4.XII.1946</date>
|
||
, RA,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFD815525A7C2FC9A" box="[611,646,834,858]" pageId="3" pageNumber="920" type="female">1 ♀</specimenCount>
|
||
(MNHN). —
|
||
<collectingRegion id="59390BEFFFFCFFBFFEE4553BA4C2FCB4" box="[262,390,860,884]" country="Madagascar" name="Fianarantsoa" pageId="3" pageNumber="920">Fianarantsoa</collectingRegion>
|
||
, 2e semestre 1892, Perrot Frères,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF5C5510A5B4FC4F" box="[190,240,887,911]" pageId="3" pageNumber="920" type="generic">2 ex.</specimenCount>
|
||
(MNHN). — Antisianaka, 2
|
||
<superScript id="6C886845FFFCFFBFFDC35510A76CFC48" attach="left" box="[545,552,887,904]" fontSize="7" pageId="3" pageNumber="920">e</superScript>
|
||
semestre 1893, Perrot Frères,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFEDD55F6A438FC69" box="[319,380,913,937]" pageId="3" pageNumber="920" type="generic">13 ex.</specimenCount>
|
||
(MNHN). — Antisianaka et lac Alaotra, 2
|
||
<superScript id="6C886845FFFCFFBFFEF255CBA453FC7D" attach="left" box="[272,279,940,957]" fontSize="7" pageId="3" pageNumber="920">e</superScript>
|
||
trimestre 1889, Perrot Frères,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFDA855CBA7C3FC04" box="[586,647,940,964]" pageId="3" pageNumber="920" type="generic">11 ex.</specimenCount>
|
||
(MNHN). — H. Perrot,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE8E55A1A4DFFC1E" box="[364,411,966,990]" pageId="3" pageNumber="920" type="generic">2 ex.</specimenCount>
|
||
(MNHN). — Feneriva, H. Perrot,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF035586A457FC39" box="[225,275,993,1017]" pageId="3" pageNumber="920" type="generic">6 ex.</specimenCount>
|
||
(MNHN). — Ambohitsritondroina Inst. Scient., Maroantsetra,
|
||
<date id="EF43E3CDFFFCFFBFFE71559CA740FBD3" box="[403,516,1019,1043]" pageId="3" pageNumber="920" value="1949-02-15">15.II.1949</date>
|
||
, Vadon,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFD83559CA7C3FBD3" box="[609,647,1019,1043]" pageId="3" pageNumber="920" type="female">1 ♀</specimenCount>
|
||
(MNHN). — Est Marojejy Rés. nat. int. XII, Beondraka,
|
||
<quantity id="5C0568E8FFFCFFBFFF925257A5FEFB88" box="[112,186,1072,1096]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="920" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="EF43E3CDFFFCFFBFFF205257A45EFB88" box="[194,282,1072,1096]" pageId="3" pageNumber="920" value="1960-12">XII.1960</date>
|
||
, P. Soga,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE895257A4CAFB88" box="[363,398,1072,1096]" pageId="3" pageNumber="920" type="female">1 ♀</specimenCount>
|
||
(MNHN). — Madagascar E,
|
||
<quantity id="5C0568E8FFFCFFBFFF6C522DA596FBA2" box="[142,210,1098,1122]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="920" unit="km" value="30.0">30 km</quantity>
|
||
E of Moramanga, Maromizaha,
|
||
<quantity id="5C0568E8FFFCFFBFFDCE522DA7C4FBA2" box="[556,640,1098,1122]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="920" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="EF43E3CDFFFCFFBFFF915202A5BAFBBD" box="[115,254,1125,1149]" pageId="3" pageNumber="920" value="1996-12-07" valueMax="1996-12-10" valueMin="1996-12-07">7-10.XII.1996</date>
|
||
, Bednarik leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE685202A4F2FBBD" box="[394,438,1125,1149]" pageId="3" pageNumber="920" type="generic">3 ex.</specimenCount>
|
||
(coll. PP). — Madagascar C., Moramanga env.,
|
||
<date id="EF43E3CDFFFCFFBFFE845218A745FB57" box="[358,513,1151,1175]" pageId="3" pageNumber="920" value="1997-12-10" valueMax="1997-12-18" valueMin="1997-12-10">10-18.XII.1997</date>
|
||
, P. Pacholátko leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF7952FDA583FB72" box="[155,199,1178,1202]" pageId="3" pageNumber="920" type="generic">7 ex.</specimenCount>
|
||
(coll. PP). — Tamatave distr., Moramanga env.,
|
||
<date id="EF43E3CDFFFCFFBFFF9252D3A440FB0C" box="[112,260,1204,1228]" pageId="3" pageNumber="920" value="1995-12-14" valueMax="1995-12-18" valueMin="1995-12-14">14-18.XII.1995</date>
|
||
, Ivo Jeniš leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE6752D3A4F4FB0C" box="[389,432,1204,1228]" pageId="3" pageNumber="920" type="generic">1 ex.</specimenCount>
|
||
(coll. PP). —Tamatave distr., Moramanga env.,
|
||
<date id="EF43E3CDFFFCFFBFFEBE52A8A4BCFB27" box="[348,504,1231,1255]" pageId="3" pageNumber="920" value="1995-12-21" valueMax="1995-12-24" valueMin="1995-12-21">21-24.XII.1995</date>
|
||
, Ivo Jeniš leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF90528EA5E7FAC1" box="[114,163,1257,1281]" pageId="3" pageNumber="920" type="generic">8 ex.</specimenCount>
|
||
(coll. PP). — Tamatave distr., Moramanga env.,
|
||
<date id="EF43E3CDFFFCFFBFFF915363A443FADC" box="[115,263,1284,1308]" pageId="3" pageNumber="920" value="1995-12-27" valueMax="1995-12-30" valueMin="1995-12-27">27-30.XII.1995</date>
|
||
, Ivo Jeniš leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE6A5363A4F7FADC" box="[392,435,1284,1308]" pageId="3" pageNumber="920" type="generic">3 ex.</specimenCount>
|
||
(coll. PP). — Madagascar Est, Moramanga env.,
|
||
<date id="EF43E3CDFFFCFFBFFEB45379A4ADFAF6" box="[342,489,1310,1334]" pageId="3" pageNumber="920" value="1998-12-17" valueMax="1998-12-24" valueMin="1998-12-17">17-24.XII.1998</date>
|
||
, leg.Jiri Moravec,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFF8D535FA5DAFA90" box="[111,158,1336,1360]" pageId="3" pageNumber="920" type="generic">1 ex.</specimenCount>
|
||
(coll. PP). —
|
||
<collectingRegion id="59390BEFFFFCFFBFFECA535FA4E1FA90" box="[296,421,1336,1360]" country="Madagascar" name="Fianarantsoa" pageId="3" pageNumber="920">Fianarantsoa</collectingRegion>
|
||
, Ranomafana National Park,Talatakely,Trail FF 4, 915-
|
||
<quantity id="5C0568E8FFFCFFBFFE745334A4A5FAAB" box="[406,481,1363,1387]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="920" unit="m" value="1000.0">1000 m</quantity>
|
||
,
|
||
<date id="EF43E3CDFFFCFFBFFE0B5334A713FAAB" box="[489,599,1363,1387]" pageId="3" pageNumber="920" value="1998-11-20">20.XI.1998</date>
|
||
, V. F. Lee, K. J. Ribardo leg.,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFEA8530AA4C1FA46" box="[330,389,1389,1414]" pageId="3" pageNumber="920" type="male">2 ♂♂</specimenCount>
|
||
(CASC). — East Andasibe (Perinet),
|
||
<geoCoordinate id="FEC9A3CAFFFCFFBFFF2D53EFA45AFA60" box="[207,286,1416,1440]" degrees="18.56" direction="south" orientation="latitude" pageId="3" pageNumber="920" precision="555" value="-18.56">18.56°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FEC9A3CAFFFCFFBFFEC553EFA43DFA60" box="[295,377,1416,1440]" degrees="48.25" direction="east" orientation="longitude" pageId="3" pageNumber="920" precision="555" value="48.25">48.25°E</geoCoordinate>
|
||
,
|
||
<date id="EF43E3CDFFFCFFBFFE6053EFA4B1FA60" box="[386,501,1416,1440]" pageId="3" pageNumber="920" value="1998-11-14">14.XI.1998</date>
|
||
, E-Y: 3374, on vegetation, leg. R. Müller,
|
||
<specimenCount id="8DFB0E84FFFCFFBFFE9053C5A4E5FA7A" box="[370,417,1442,1466]" pageId="3" pageNumber="920" type="generic">1 ex.</specimenCount>
|
||
(TMSA).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3E79686FFFCFFB9FF93538CA409FE92" lastPageId="5" lastPageNumber="922" pageId="3" pageNumber="920" type="description">
|
||
<paragraph id="9B42C50DFFFCFFBFFF93538CA4FEF9C3" blockId="3.[111,647,1515,1700]" box="[113,442,1515,1541]" pageId="3" pageNumber="920">
|
||
REDESCRIPTION OF
|
||
<typeStatus id="44467BAFFFFCFFBFFEA75397A4FEF9C3" box="[325,442,1520,1539]" pageId="3" pageNumber="920" type="lectotype">LECTOTYPE</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFF93506DA4D7F964" blockId="3.[111,647,1515,1700]" pageId="3" pageNumber="920">
|
||
Length: 8.0 mm; length of elytra:
|
||
<quantity id="5C0568E8FFFCFFBFFE3C506DA772F9E5" box="[478,566,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="920" unit="mm" value="5.9">5.9 mm</quantity>
|
||
; width:
|
||
<quantity id="5C0568E8FFFCFFBFFF93504DA581F985" box="[113,197,1578,1605]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="920" unit="mm" value="4.8">4.8 mm</quantity>
|
||
. Body oval, dorsal surface brown with greenish shine, ventral surface metallic-green, completely shiny and glabrous, except for a few small setae on the head (habitus
|
||
<figureCitation id="03C6D988FFFCFFBFFED150EEA4C1F964" box="[307,389,1673,1700]" captionStart="FIG" captionStartId="4.[113,124,1277,1294]" captionTargetBox="[490,671,1083,1223]" captionTargetPageId="4" captionText="FIG. 1. — A, Distribution of the species of Emphania (Emphania erichsoni n. sp.could not be localized due to insufficient label information “Madagascar”),the habitus of the type species E.metallica (Blanchard,1850) is shown (not scaled) in upper left corner;B-D, E. metallica (Blanchard, 1850) (lectotype); E-G, E. lacroixi n. sp. (holotype); B, E, aedeagus, left side lateral views; C, F, aedeagus, dorsal views; D, G, aedeagus, right side lateral views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4525863" httpUri="https://zenodo.org/record/4525863/files/figure.png" pageId="3" pageNumber="920">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFD5956BFA6F9FC8D" blockId="3.[674,1212,216,1700]" pageId="3" pageNumber="920">Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins unevenly convex and convergent to moderately rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anteriorly shallowly sinuate medially; entire surface weakly convex, shiny, finely and densely punctate, with a few short, erect setae anteriorly; frontoclypeal suture feebly impressed and weakly curved medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and sharply pointed, finely and sparsely punctate, with a short single terminal hair. Frons with fine, moderately dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.6. Antenna brown, with nine antennomeres, antennomeres 3-5 fused; club with three antennomeres, a little longer than the remaining antennomeres combined.</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFD595536A60DFB69" blockId="3.[674,1212,216,1700]" pageId="3" pageNumber="920">Pronotum subtrapezoidal, widest at base, lateral margins very weakly and evenly convex and strongly convergent anteriorly, anterior angles moderately produced but sharp, posterior angles blunt, anterior marginal line complete, margin weakly convexly produced medially; basal marginal line fine and widely interrupted medially; surface sparsely and finely punctate, glabrous; anterior and lateral borders sparsely setose. Scutellum wide, triangular, apex sharp, with fine and moderately dense punctures, glabrous.</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFD5952C9A107FA26" blockId="3.[674,1212,216,1700]" pageId="3" pageNumber="920">
|
||
Elytra oblong, widest at middle, with convex semicircular declivity distinctly separated from apical portion of elytra, apical interior angle rightangled, apical external angle blunt but moderately rounded in the tip, apical border straight; striae not impressed and impunctate, almost invisible, intervals completely flat, with fine, moderately dense punctures, glabrous; epipleural edge robust, ending at the external apical angle of elytra, epipleura very sparsely setose (setae in
|
||
<typeStatus id="44467BAFFFFCFFBFFC4653ACA142FA26" box="[932,1030,1483,1510]" pageId="3" pageNumber="920" type="lectotype">lectotype</typeStatus>
|
||
lost).
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFCFFBFFD59538CA1FCF964" blockId="3.[674,1212,216,1700]" pageId="3" pageNumber="920">External angle of metacoxa posteriorly produced forming a sharp tooth, lateral marginal line in posterior half slightly reflexed; mesal process of metacoxa slender and longer than remainder metacoxa and its ventral surface longitudinally convex.Ratio of length of metepisternum/metacoxa:1/2.0. Pygidium weakly</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF0B5685A5DEFE9E" pageId="4" pageNumber="921">
|
||
<geoCoordinate id="FEC9A3CAFFFBFFB8FF0B5685A455FF33" box="[233,273,226,243]" degrees="44" direction="east" orientation="longitude" pageId="4" pageNumber="921" precision="55555" value="44.0">44°E</geoCoordinate>
|
||
46° 48° 50° A B C D
|
||
<geoCoordinate id="FEC9A3CAFFFBFFB8FF93572AA5DEFE9E" box="[113,154,333,350]" degrees="12" direction="south" orientation="latitude" pageId="4" pageNumber="921" precision="55555" value="-12.0">12°S</geoCoordinate>
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9B57ADA5D2FE1B" blockId="4.[121,150,458,475]" box="[121,150,458,475]" pageId="4" pageNumber="921">14°</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9B542FA5D2FD99" blockId="4.[121,150,584,601]" box="[121,150,584,601]" pageId="4" pageNumber="921">16°</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9B54A2A5D2FD16" blockId="4.[121,150,709,726]" box="[121,150,709,726]" pageId="4" pageNumber="921">18°</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FD0A5569A5D1FC94" pageId="4" pageNumber="921">E F G 20°</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9B55A7A5D1FC11" blockId="4.[121,149,960,977]" box="[121,149,960,977]" pageId="4" pageNumber="921">22°</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9B525AA5D1FB0C" pageId="4" pageNumber="921">24° 26°</paragraph>
|
||
<caption id="CF829585FFFBFFB8FF93529AA71EFA92" ID-DOI="http://doi.org/10.5281/zenodo.4525863" ID-Zenodo-Dep="4525863" httpUri="https://zenodo.org/record/4525863/files/figure.png" pageId="4" pageNumber="921" startId="4.[113,124,1277,1294]" targetBox="[490,671,1083,1223]" targetPageId="4">
|
||
<paragraph id="9B42C50DFFFBFFB8FF93529AA71EFA92" blockId="4.[112,1211,1277,1362]" pageId="4" pageNumber="921">
|
||
FIG. 1. —
|
||
<emphasis id="A989191FFFFBFFB8FF5E529AA58DFACE" bold="true" box="[188,201,1277,1294]" pageId="4" pageNumber="921">A</emphasis>
|
||
, Distribution of the species of
|
||
<taxonomicName id="5CFDBE8EFFFBFFB8FE5E529AA74AFACE" authorityName="Erichson" authorityYear="1847" box="[444,526,1277,1294]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="921" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A989191FFFFBFFB8FE5E529AA74AFACE" box="[444,526,1277,1294]" italics="true" pageId="4" pageNumber="921">Emphania</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="5CFDBE8EFFFBFFB8FDF4529AA7F2FACE" authority="Ahrens & Fabrizi, 2008" authorityName="Ahrens & Fabrizi" authorityYear="2008" box="[534,694,1277,1294]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="921" phylum="Arthropoda" rank="species" species="erichsoni" status="sp. nov.">
|
||
<emphasis id="A989191FFFFBFFB8FDF4529AA7F2FACE" box="[534,694,1277,1294]" italics="true" pageId="4" pageNumber="921">Emphania erichsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2BAA464FFFBFFB8FD58529AA7ACFACE" box="[698,744,1277,1294]" pageId="4" pageNumber="921" rank="species">n. sp.</taxonomicNameLabel>
|
||
could not be localized due to insufficient label information “Madagascar”),the habitus of the type species
|
||
<taxonomicName id="5CFDBE8EFFFBFFB8FE125374A79BFAE5" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[496,735,1299,1317]" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="921" phylum="Arthropoda" rank="species" species="metallica">
|
||
<emphasis id="A989191FFFFBFFB8FE125374A709FAE4" box="[496,589,1299,1316]" italics="true" pageId="4" pageNumber="921">E.metallica</emphasis>
|
||
(
|
||
<bibRefCitation id="FF6CB8FCFFFBFFB8FDB75373A79FFAE5" author="BLANCHARD M. E." box="[597,731,1300,1317]" pageId="4" pageNumber="921" refId="ref6312" refString="BLANCHARD M. E. 1850. - Ordre des Coleoptera, in MILNE- EDWARDS H., BLANCHARD C. E. & LUCUS H. (eds), Museum d'Histoire naturelle de Paris. Catalogue de la collection entomologique. Classe des Insectes. Volume 1, part 1. Gide & Baudry, Paris, 128 p." type="book" year="1850">Blanchard,1850</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is shown (not scaled) in upper left corner;
|
||
<emphasis id="A989191FFFFBFFB8FBD05374A110FAE4" bold="true" box="[1074,1108,1299,1317]" pageId="4" pageNumber="921">B -D</emphasis>
|
||
,
|
||
<taxonomicName id="5CFDBE8EFFFBFFB8FBBE5374A440FAFC" authority="(Blanchard, 1850)" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="921" phylum="Arthropoda" rank="species" species="metallica">
|
||
<emphasis id="A989191FFFFBFFB8FBBE5374A1FEFAE4" box="[1116,1210,1299,1316]" italics="true" pageId="4" pageNumber="921">E. metallica</emphasis>
|
||
(
|
||
<bibRefCitation id="FF6CB8FCFFFBFFB8FF94534CA444FAFC" author="BLANCHARD M. E." box="[118,256,1323,1340]" pageId="4" pageNumber="921" refId="ref6312" refString="BLANCHARD M. E. 1850. - Ordre des Coleoptera, in MILNE- EDWARDS H., BLANCHARD C. E. & LUCUS H. (eds), Museum d'Histoire naturelle de Paris. Catalogue de la collection entomologique. Classe des Insectes. Volume 1, part 1. Gide & Baudry, Paris, 128 p." type="book" year="1850">Blanchard, 1850</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(lectotype);
|
||
<emphasis id="A989191FFFFBFFB8FE89534DA4C8FAFB" bold="true" box="[363,396,1322,1340]" pageId="4" pageNumber="921">E -G</emphasis>
|
||
,
|
||
<taxonomicName id="5CFDBE8EFFFBFFB8FE75534DA4A3FAFB" authority="Ahrens & Fabrizi, 2008" authorityName="Ahrens & Fabrizi" authorityYear="2008" box="[407,487,1322,1339]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="921" phylum="Arthropoda" rank="species" species="lacroixi" status="sp. nov.">
|
||
<emphasis id="A989191FFFFBFFB8FE75534DA4A3FAFB" box="[407,487,1322,1339]" italics="true" pageId="4" pageNumber="921">E. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2BAA464FFFBFFB8FE0F534CA759FAFC" box="[493,541,1323,1340]" pageId="4" pageNumber="921" rank="species">n. sp.</taxonomicNameLabel>
|
||
(holotype);
|
||
<emphasis id="A989191FFFFBFFB8FD9C534DA7CFFAFB" bold="true" box="[638,651,1322,1339]" pageId="4" pageNumber="921">B</emphasis>
|
||
,
|
||
<emphasis id="A989191FFFFBFFB8FD77534DA7E5FAFB" bold="true" box="[661,673,1322,1339]" pageId="4" pageNumber="921">E</emphasis>
|
||
, aedeagus, left side lateral views;
|
||
<emphasis id="A989191FFFFBFFB8FC20534DA694FAFB" bold="true" box="[962,976,1322,1339]" pageId="4" pageNumber="921">C</emphasis>
|
||
,
|
||
<emphasis id="A989191FFFFBFFB8FC38534DA6A1FAFB" bold="true" box="[986,997,1322,1339]" pageId="4" pageNumber="921">F</emphasis>
|
||
, aedeagus, dorsal views;
|
||
<emphasis id="A989191FFFFBFFB8FF935326A53BFA92" bold="true" box="[113,127,1345,1362]" pageId="4" pageNumber="921">D</emphasis>
|
||
,
|
||
<emphasis id="A989191FFFFBFFB8FF685326A5DCFA92" bold="true" box="[138,152,1345,1362]" pageId="4" pageNumber="921">G</emphasis>
|
||
, aedeagus, right side lateral views. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B42C50DFFFBFFB8FF9353EBA75FFA26" blockId="4.[113,648,1420,1700]" pageId="4" pageNumber="921">convex, very finely and sparsely punctate, without smooth midline, punctures without microscopic setae and with a few long setae apically.</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FF6A538CA106F944" blockId="4.[113,648,1420,1700]" lastBlockId="4.[674,1210,1420,1700]" pageId="4" pageNumber="921">Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.4, dorsally longitudinally convex, with two groups of spines, basal one at one third, apical one at two thirds of metatibial length; lateral face longitudinally convex, with dense and fine longitudinally impressed punctures, glabrous; ventral edge not serrate, with five strong spines equidistant from each other, medial face not punctate.Tarsomeres dorsally smooth, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina, first metatarsomere a little shorter than the two following segments combined and a little longer than the upper tibial spur.</paragraph>
|
||
<paragraph id="9B42C50DFFFBFFB8FD5950EEA6FAF964" blockId="4.[674,1210,1420,1700]" box="[699,958,1673,1700]" pageId="4" pageNumber="921">
|
||
Aedeagus:
|
||
<figureCitation id="03C6D988FFFBFFB8FCCC50EEA6C7F964" box="[814,899,1673,1700]" captionStart="FIG" captionStartId="4.[113,124,1277,1294]" captionTargetBox="[490,671,1083,1223]" captionTargetPageId="4" captionText="FIG. 1. — A, Distribution of the species of Emphania (Emphania erichsoni n. sp.could not be localized due to insufficient label information “Madagascar”),the habitus of the type species E.metallica (Blanchard,1850) is shown (not scaled) in upper left corner;B-D, E. metallica (Blanchard, 1850) (lectotype); E-G, E. lacroixi n. sp. (holotype); B, E, aedeagus, left side lateral views; C, F, aedeagus, dorsal views; D, G, aedeagus, right side lateral views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4525863" httpUri="https://zenodo.org/record/4525863/files/figure.png" pageId="4" pageNumber="921">Figure 1</figureCitation>
|
||
B-D.
|
||
</paragraph>
|
||
<paragraph id="9B42C50DFFFAFFB9FF9356BFA5A3FF30" blockId="5.[112,646,216,338]" box="[113,231,216,242]" pageId="5" pageNumber="922">VARIATION</paragraph>
|
||
<paragraph id="9B42C50DFFFAFFB9FF93569FA409FE92" blockId="5.[112,646,216,338]" pageId="5" pageNumber="922">
|
||
Length:
|
||
<quantity id="5C0568E8FFFAFFB9FF28569FA408FED3" box="[202,332,248,275]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="8.5" metricValueMin="7.9" pageId="5" pageNumber="922" unit="mm" value="8.2" valueMax="8.5" valueMin="7.9">7.9-8.5 mm</quantity>
|
||
; length of elytra:
|
||
<quantity id="5C0568E8FFFAFFB9FDE2569FA7C6FED3" box="[512,642,248,275]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.1" metricValueMin="5.8" pageId="5" pageNumber="922" unit="mm" value="5.949999999999999" valueMax="6.1" valueMin="5.8">5.8-6.1 mm</quantity>
|
||
; width:
|
||
<quantity id="5C0568E8FFFAFFB9FF5C5770A401FEF2" box="[190,325,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="5" pageNumber="922" unit="mm" value="4.9" valueMax="5.3" valueMin="4.5">4.5-5.3 mm</quantity>
|
||
. The female has the antennal club slightly shorter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |