676 lines
77 KiB
XML
676 lines
77 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1176.107829" ID-Pensoft-Pub="1313-2970-1176-221" ID-Pensoft-UUID="0E1B153259B55C91A69FFDFF0A954CF5" ID-ZooBank="D2EBCA72210D450BB2C6A06CBB485C4D" ModsDocID="1313-2970-1176-221" checkinTime="1693383710254" checkinUser="pensoft" docAuthor="El Alami, Majida, Vuataz, Laurent, El Yaagoubi, Sara & Gattolliat, Jean-Luc" docDate="2023" docId="1266EF475E895C658629080D4804BD3D" docLanguage="en" docName="ZooKeys 1176: 221-241" docOrigin="ZooKeys 1176" docPubDate="2023-08-29" docSource="http://dx.doi.org/10.3897/zookeys.1176.107829" docTitle="Alainites albai El Alami, Vuataz & Gattolliat 2023, sp. nov." docType="treatment" docUuid="0B24CF09-A193-42A2-9A9C-96266B5C0C4A" docUuidSource="ZooBank" docVersion="2" id="0E1B153259B55C91A69FFDFF0A954CF5" lastPageNumber="221" masterDocId="0E1B153259B55C91A69FFDFF0A954CF5" masterDocTitle="A new species of the genus Alainites Waltz & McCafferty, 1994 (Ephemeroptera, Baetidae) from the north of Morocco" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1693384015681" updateUser="ExternalLinkService">
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<mods:titleInfo id="E6B6E881472E2100F3C1B250193CBAD3">
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<mods:title id="414701AAA393A08CFA7F1E79A487DE79">A new species of the genus Alainites Waltz & McCafferty, 1994 (Ephemeroptera, Baetidae) from the north of Morocco</mods:title>
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<mods:name id="351387612A33E0A6DEACE41D72CFFA04" type="personal">
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<mods:roleTerm id="985A1AC0C9D2CC99E01B9988D337E223">Author</mods:roleTerm>
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<mods:namePart id="81038C26EEAB2474D310198E8DDBDC1B">El Alami, Majida</mods:namePart>
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<mods:nameIdentifier id="403CE88C3FE4190D314BE70097CF5110" type="ORCID">https://orcid.org/0000-0003-2664-646X</mods:nameIdentifier>
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<mods:affiliation id="D7A3592621E35931FAF04713E34B4399">Universite Abdelmalek Essaadi, Faculte des Sciences, Departement de Biologie, Laboratoire Ecologie, Systematique et Conservation de la Biodiversite (LESCB), Unite de Recherche Labellisee CNRST N ° 18. B. P. 2121. Tetouan 93002, Morocco</mods:affiliation>
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<mods:nameIdentifier id="0C3AB0DFB6C0D6BA68764F5E95FC069E" type="email">melalamielmoutaoukil@uae.ac.ma</mods:nameIdentifier>
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<mods:name id="DCB7B5DC5F395BB0FBDB21DA47E2FF16" type="personal">
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<mods:roleTerm id="E9F26033DF211900E724B5DD3674487B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="340905616F6507A141E6C57CDECDDB0A">Vuataz, Laurent</mods:namePart>
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<mods:nameIdentifier id="6F2FD16F7FEBD87F0A12E774B79E48AB" type="ORCID">https://orcid.org/0000-0001-9193-8683</mods:nameIdentifier>
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<mods:affiliation id="F372979DAD6AB7723C32FEC40C38C32D">Museum Cantonal des Sciences Naturelles, Departement de Zoologie, Palais de Rumine, Place Riponne 6, CH- 1005, Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name id="7A000B0FA2C91695CCCE9A85DF91FDD3" type="personal">
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<mods:role id="CD75057D5900D4636E99A79F52A6B264">
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<mods:roleTerm id="6D129FC89558CBAE5B6743079CA99D33">Author</mods:roleTerm>
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<mods:namePart id="D614EDFF72EC7C89A317DB96FF0E45D4">El Yaagoubi, Sara</mods:namePart>
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<mods:nameIdentifier id="A8802587B0AF945AEA0BDBDA8FBF3559" type="ORCID">https://orcid.org/0000-0003-1860-6433</mods:nameIdentifier>
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<mods:affiliation id="670C5157B157DAB26ED90574EC10AE4C">Universite Abdelmalek Essaadi, Faculte des Sciences, Departement de Biologie, Laboratoire Ecologie, Systematique et Conservation de la Biodiversite (LESCB), Unite de Recherche Labellisee CNRST N ° 18. B. P. 2121. Tetouan 93002, Morocco</mods:affiliation>
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<mods:namePart id="7E1C013AA46509AC1B0FF17179F56A6F">Gattolliat, Jean-Luc</mods:namePart>
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<mods:nameIdentifier id="32D1D5C488EDC0784D5241437F786F0B" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
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<mods:affiliation id="B1F039744723354549F6E0F15B5B724E">Museum Cantonal des Sciences Naturelles, Departement de Zoologie, Palais de Rumine, Place Riponne 6, CH- 1005, Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:title id="A3002DAEEB8C6C3EFA03A3E0613957C2">ZooKeys</mods:title>
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<mods:date id="25E5696BFAD2E4DE4CA637B9049763D6">2023</mods:date>
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<mods:number id="D683D30439360812B9B40DFC7C9FFEEF">2023-08-29</mods:number>
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<mods:number id="C5ECA01DAE7CEB792382C085A862176E">1176</mods:number>
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<mods:url id="56D0F9DF4A3474B8DB693B60867566DD">http://dx.doi.org/10.3897/zookeys.1176.107829</mods:url>
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<mods:classification id="98418F49849C53F7BAD7A29B7ADDC46F">journal article</mods:classification>
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<mods:identifier id="8D8C7FAB8BA2159957D471C28E778B42" type="DOI">http://dx.doi.org/10.3897/zookeys.1176.107829</mods:identifier>
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<mods:identifier id="ECC9E6CE9A5E47BF292E53CCCBBCEAE4" type="Pensoft-Pub">1313-2970-1176-221</mods:identifier>
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<mods:identifier id="E9352F99439A9CA06D3CFB1426051050" type="ZooBank">D2EBCA72210D450BB2C6A06CBB485C4D</mods:identifier>
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<treatment id="1266EF475E895C658629080D4804BD3D" LSID="urn:lsid:zoobank.org:act:0B24CF09-A193-42A2-9A9C-96266B5C0C4A" httpUri="http://treatment.plazi.org/id/1266EF475E895C658629080D4804BD3D" lastPageNumber="221" pageId="0" pageNumber="221">
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<subSubSection id="B539D76BF9070D4538FD6C969F764E2D" pageId="0" pageNumber="221" type="nomenclature">
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<paragraph id="38F99D349927E4EC1F57B6B557F415EE" pageId="0" pageNumber="221">
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<taxonomicName id="E9E34BBB65498D91A6920C9C7A6784B1" LSID="https://zoobank.org/0B24CF09-A193-42A2-9A9C-96266B5C0C4A" authority="El Alami, Vuataz & Gattolliat" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai" status="sp. nov.">Alainites albai El Alami, Vuataz & Gattolliat</taxonomicName>
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<taxonomicNameLabel id="FB41B15F97FC460EEF3011C60FFAA84A" pageId="0" pageNumber="221">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="3E60E4DD49646E088D365B4AD4B9AFE0" pageId="0" pageNumber="221" type="description">
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<paragraph id="DFABF8418AADCA19B046FD1051955CD3" pageId="0" pageNumber="221">
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||
<figureCitation id="B5AF8D79F1A25C430577749A6E108400" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The sampling site localization of Alainites albai sp. nov., in the Rif domain." figureDoi="10.3897/zookeys.1176.107829.figure1" httpUri="https://binary.pensoft.net/fig/898273" pageId="0" pageNumber="221">Figs 1</figureCitation>
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<figureCitation id="4EA514F68260BD81F53D6BE5A9D87E9A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alainites albai sp. nov., nymph habitus a female, dorsal view b female lateral view c male, dorsal view d lateral view of the first three gills." figureDoi="10.3897/zookeys.1176.107829.figure2" httpUri="https://binary.pensoft.net/fig/898274" pageId="0" pageNumber="221">, 2</figureCitation>
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||
<figureCitation id="EEFA43B166F705D2771BA87DA72265AB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">, 3</figureCitation>
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<figureCitation id="4741A33D53B3A0A58D1FA6C1C70489B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Alainites albai sp. nov., nymph morphology a right mandible b spines between mola and prostheca bifid c left mandible d spines between mola and prostheca." figureDoi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221">, 4</figureCitation>
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||
<figureCitation id="BCD2CF9EBDB7753645FA8DF7D6C64024" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Alainites albai sp. nov., nymph morphology a hypopharynx b maxilla c details of dentisetae and maxilla crown of setae d labium." figureDoi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221">, 5</figureCitation>
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||
<figureCitation id="279860C55AB70028495210B55B24E6DF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">, 6</figureCitation>
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<figureCitation id="59ACF60D02EDBD7C04BFACDE53BD7BD9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Alainites albai sp. nov., nymph morphology a distal margin of tergum IV b gill IV c gill V d left paraproct e right paraproct." figureDoi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221">, 7</figureCitation>
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||
<figureCitation id="87C395DB72F32044A82C4C52E4F43A9B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bayesian (BEAST) maximum clade credibility COI tree of the genus Alainites in the West Palearctic: branch ends labelled with GBIF codes in bold indicate newly sequenced specimens; the DTNHM 1449 - 21, TDAAT 323 - 19, and VMCOL 019 - 20 - VMCOL 021 - 20 codes are from BOLD; the other codes correspond to sequences obtained from GenBank. Colored vertical boxes indicate species delimitation hypothesis according to the ASAP, mPTP, and GMYC methods. For each MOTU, the corresponding species names (where available) and the country / region of origin is provided, with the newly described species and associated GBIF codes specified in bold. Circles on branches indicate Bayesian posterior probabilities> 0.9. Outgroup branches, labels, and species names are presented in grey." figureDoi="10.3897/zookeys.1176.107829.figure8" httpUri="https://binary.pensoft.net/fig/898280" pageId="0" pageNumber="221">, 8</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B2DA039FB66EDFA57D917BCF15CE9F34" pageId="0" pageNumber="221" type="type material">
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<paragraph id="24B2E6E277B2F32CED44E9B6369AA4BB" pageId="0" pageNumber="221">Type-material.</paragraph>
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<paragraph id="E9F59E1C0BCB3269AD5DA29A9462139A" pageId="0" pageNumber="221">
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<emphasis id="F624606FD58E2E642FDF8DDED47D3A1F" bold="true" pageId="0" pageNumber="221">
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<emphasis id="469EE19885AF856160AF00BDB2C5781C" italics="true" pageId="0" pageNumber="221">Holotype</emphasis>
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.
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</emphasis>
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Morocco. Nymph; Chefchaouen Province, S10 Oued
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<normalizedToken id="EDA7FEDAA983505E93A3058169689196" originalValue="Kelâa">Kelaa</normalizedToken>
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; Loc. Akchour;
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<geoCoordinate id="9961D8C439E789FB51A103D897122223" degrees="35" direction="north" minutes="14" orientation="latitude" precision="15" seconds="32" value="35.24222">35°14'32"N</geoCoordinate>
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,
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<geoCoordinate id="55798A1081C10A1F0E4A662EF8FDD549" degrees="05" direction="west" minutes="10" orientation="longitude" precision="15" seconds="10" value="-5.169444">05°10'10"W</geoCoordinate>
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; alt. 460 m; 3.iii.2021; El Yaagoubi leg.; GBIFCH00763782; MZL.
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</paragraph>
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<paragraph id="558B425170BEA095FF2780E18F2732C1" pageId="0" pageNumber="221">
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||
<emphasis id="AC096BE9A664A0A8F1653F7B20EF81D8" bold="true" pageId="0" pageNumber="221">
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<emphasis id="31F7EC5A02D8016B52790255494C317A" italics="true" pageId="0" pageNumber="221">Paratypes</emphasis>
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.
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</emphasis>
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Morocco. 1 nymph; same data as holotype; DNA; GBIFCH00980877; MZL • 2 nymphs; same data as holotype; 7.xi.2014; El Bazi leg.; in alcohol; LESCB • 1 nymph; Chefchaouen Province, S1 Oued Amazithen, Loc. El Ouesteyine;
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||
<geoCoordinate id="44457DFFA9D9A1AA46D7CB8C807E3E3A" degrees="35" direction="north" minutes="17" orientation="latitude" precision="1" seconds="57.264" value="35.29924">35°17'57.264"N</geoCoordinate>
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,
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<geoCoordinate id="DBF578CE527DD6E3D0B0F0BAD5E5ABAE" degrees="4" direction="west" minutes="54" orientation="longitude" precision="1" seconds="33.732" value="-4.90937">4°54'33.732"W</geoCoordinate>
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; alt. 166 m; 27.ii.2021; El Yaagoubi leg.; DNA; GBIFCH01144392; MZL • 2 nymphs; Chefchaouen Province, S2 Oued Sidi Yahia Aarab, Loc. Sidi Yahia Aarab;
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||
<geoCoordinate id="18FCBDF96BFD9F937ADBF2A98E7BFFB8" degrees="35" direction="north" minutes="17" orientation="latitude" precision="1" seconds="10.428" value="35.28623">35°17'10.428"N</geoCoordinate>
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,
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<geoCoordinate id="9A945DB53DBDB7E98831DDC3ED141311" degrees="4" direction="west" minutes="53" orientation="longitude" precision="1" seconds="37.644" value="-4.89379">4°53'37.644"W</geoCoordinate>
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||
; alt. 114 m; 28.v.2022; El Yaagoubi leg.; one in alcohol; GBIFCH00763784 and other DNA; GBIFCH01144394; MZL; 1 nymph; 18.vi.2014; Khadri leg.; in Alcohol; GBIFCH00763781; MZL; 4 nymphs; 28/v/2022; El Yaagoubi leg.; LESCB • 6 nymphs; Chefchaouen Province, S3 Oued Jenane Niche, Loc. Jenane Niche;
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||
<geoCoordinate id="62EC0FA592F4205D53318AE8A5423E53" degrees="35" direction="north" minutes="16" orientation="latitude" precision="1" seconds="44.904" value="35.27914">35°16'44.904"N</geoCoordinate>
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||
,
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<geoCoordinate id="A02602DC88B72603781F6167ED6200A7" degrees="4" direction="west" minutes="51" orientation="longitude" precision="1" seconds="40.788" value="-4.86133">4°51'40.788"W</geoCoordinate>
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||
; alt. 93 m; 9.ix.2021; El Yaagoubi leg.; LESCB • 11 nymphs; Chefchaouen, Province, S4 Oued
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||
<normalizedToken id="9E5CE05C96C07D2D69FA9C2C8C546B0E" originalValue="Aârkob">Aarkob</normalizedToken>
|
||
, Loc. Arherarose;
|
||
<geoCoordinate id="AC7E3B7B8454A2E8FF778948AB1E69AB" degrees="35" direction="north" minutes="15" orientation="latitude" precision="1" seconds="59.4" value="35.2665">35°15'59.4"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="252E0E7F0C06765987E055ADAD182288" degrees="4" direction="west" minutes="50" orientation="longitude" precision="1" seconds="33.216" value="-4.8425603">4°50'33.216"W</geoCoordinate>
|
||
; alt. 128 m; 9.xii.2021; El Yaagoubi leg.; in alcohol; LESCB • 22 nymphs; Chefchaouen Province, S5 Oued Assifane, Loc. Igourain;
|
||
<geoCoordinate id="90F30C8B97898F425C223333B3ECB085" degrees="35" direction="north" minutes="7" orientation="latitude" precision="1" seconds="6.7584" value="35.11854">35°7'6.7584"N</geoCoordinate>
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||
,
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||
<geoCoordinate id="904434CDF12544F2B6D6328983A4D194" degrees="4" direction="west" minutes="59" orientation="longitude" precision="1" seconds="3.9984" value="-4.984444">4°59'3.9984"W</geoCoordinate>
|
||
; alt. 1405 m; 9.ix.2021; El Yaagoubi leg; 1 on slide; LESCB • 5 nymphs; Chefchaouen Province, S6 Oued Beni Mhammed, Loc. Beni Mhammed;
|
||
<geoCoordinate id="FA9899446B24250132C85BA85656A19D" degrees="35" direction="north" minutes="09" orientation="latitude" precision="1" seconds="34.0812" value="35.15947">35°09'34.0812"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="9D56B5F800D9CF1F8C4D5268CF928D83" degrees="5" direction="west" minutes="07" orientation="longitude" precision="1" seconds="34.0212" value="-5.126117">5°07'34.0212"W</geoCoordinate>
|
||
; alt. 1330 m; 29.v.2008; El Alami leg.; in alcohol; GBIFCH00763777; MZL • 2 nymphs; Chefchaouen Province, S7 Oued Afeska, Loc. Afeska;
|
||
<geoCoordinate id="8A489D0975A4F20264FD7960FFBFF72E" degrees="35" direction="north" minutes="10" orientation="latitude" precision="1" seconds="11.0388" value="35.169735">35°10'11.0388"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="04ACC189A797FECBB08EAC3B1DADD413" degrees="5" direction="west" minutes="13" orientation="longitude" precision="1" seconds="6.2988" value="-5.218416">5°13'6.2988"W</geoCoordinate>
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||
; alt. 1293 m; 2.vi.2021; El Yaagoubi leg.; 1 nymph DNA; GBIFCH01144391; 2.vi.2020;2 in alcohol; GBIFCH00763777; GBIFCH00763783; MZL and 4 nymphs; 2.vi.2022; El Yaagoubi leg.; 3 in alcohol and 1 on slide; LESCB • 10 nymphs; Chefchaouen Province, S8 Oued Maggo, Loc. Maggo Nord village;
|
||
<geoCoordinate id="253E9B8C96B3B286CECDDF045BE07E0A" degrees="35" direction="north" minutes="6" orientation="latitude" precision="1" seconds="48.6" value="35.1135">35°6'48.6"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3ECE97F814D627AB1BF36C0AA660645C" degrees="5" direction="west" minutes="11" orientation="longitude" precision="1" seconds="26.7" value="-5.19075">5°11'26.7"W</geoCoordinate>
|
||
; alt. 905 m; 24.ii.2022; El Yaagoubi leg.; LESCB; 5 nymphs; 3.vi.2016; El Alami, leg.; in alcohol; GBIFCH00763780; MZL • 4 nymphs; Chefchaouen Province, S9 Oued Ouara, Loc. Khizana; 35°02'614"N,
|
||
<geoCoordinate id="4236838B1A57B75989201309163D3F49" degrees="5" direction="west" minutes="14" orientation="longitude" precision="15" seconds="016" value="-5.2377777">5°14'016"W</geoCoordinate>
|
||
; alt. 930 m; 23.vi.2022; El Yaagoubi leg.; 2 on slide; LESCB; 2 for DNA; GBIFCH01144393; GBIFCH01144390; MZL • 3 nymphs; Chefchaouen Province, S11 Oued Laou, Loc. Abiyati;
|
||
<geoCoordinate id="EF3AE1617B83519C996DDF8DF29EA44B" degrees="35" direction="north" minutes="17" orientation="latitude" precision="1" seconds="55.14" value="35.29865">35°17'55.14"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="57DD0174B069594C17470833D6C7F633" degrees="5" direction="west" minutes="13" orientation="longitude" precision="1" seconds="59.99" value="-5.2333307">5°13'59.99"W</geoCoordinate>
|
||
; alt. 140 m; 11.iii.2001; El Alami leg.; in alcohol; GBIFCH00763779; MZL • 1 nymph; Chefchaouen Province, S12 Oued Harakat, Loc. Mezine village;
|
||
<geoCoordinate id="D7D2944E51D589525AC3B549D8E6718F" degrees="35" direction="north" minutes="6" orientation="latitude" precision="15" seconds="8" value="35.102222">35°6'8"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6392B43844104FB94981C2644C6018A2" degrees="5" direction="west" minutes="20" orientation="longitude" precision="15" seconds="46" value="-5.3461113">5°20'46"W</geoCoordinate>
|
||
; alt. 740 m; 31.iii.2021; El Yaagoubi leg.; DNA, GBIFCH00980924; MZL • 20 nymphs; Chefchaouen Province, S13 Oued Mansoura, Loc. Tanaqoub;
|
||
<geoCoordinate id="B6CDF435597A98CE2EB1B71CF269EB9C" degrees="35" direction="north" minutes="5" orientation="latitude" precision="15" seconds="16" value="35.087776">35°5'16"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3D30F85434350255D03BF500BE7AD230" degrees="5" direction="west" minutes="30" orientation="longitude" precision="15" seconds="37" value="-5.5102777">5°30'37"W</geoCoordinate>
|
||
; alt. 124 m; 01.vi.2021; El Yaagoubi leg.; 2 on slide; LESCB • 2 nymphs; Ouezzane Province, S14 Oued Loukkos, Loc. Souk El Had;
|
||
<geoCoordinate id="026E2C1B7041B4ECC18DEAA7249DF4BE" degrees="35" direction="north" minutes="01" orientation="latitude" precision="15" seconds="21" value="35.0225">35°01'21"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F79E342D22B703FDC9BA6814DEF10994" degrees="5" direction="west" minutes="25" orientation="longitude" precision="15" seconds="14" value="-5.4205556">5°25'14"W</geoCoordinate>
|
||
; alt. 140 m; 11.iv.2021; El Yaagoubi leg.; in alcohol; GBIFCH00763776; MZL • 1 nymph; Tetouan Province, S15 Oued Tisgris, Loc. Hammadesh;
|
||
<geoCoordinate id="C545886EE60BDEE86FFBC4E8B6D6691A" degrees="35" direction="north" minutes="22" orientation="latitude" precision="15" seconds="02" value="35.36722">35°22'02"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="32AD3C41FE7E103F66B0650025A291C9" degrees="5" direction="west" minutes="32" orientation="longitude" precision="15" seconds="02" value="-5.533889">5°32'02"W</geoCoordinate>
|
||
; alt. 505 m; 20.iv.2021; El Yaagoubi leg.; DNA; GBIFCH01144254; MZL • 10 nymphs; Tetouan Province, S16 Oued Rmel, Loc. Ain Dchicha;
|
||
<geoCoordinate id="E4595AB29D31423B9A714C898FD52353" degrees="35" direction="north" minutes="52" orientation="latitude" precision="15" seconds="40" value="35.877777">35°52'40"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D4C16497EB79C1CB1FE4E0A8752B3E63" degrees="5" direction="west" minutes="28" orientation="longitude" precision="15" seconds="24" value="-5.4733334">5°28'24"W</geoCoordinate>
|
||
; alt. 49 m; 20.x.1997; El Alami leg.; in alcohol; LESCB and 2 nymphs in alcohol; GBIFCH00763778; MZL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C77E022AF1D89C026E14A9C5903F616A" pageId="0" pageNumber="221" type="differential diagnosis">
|
||
<paragraph id="C4779558E6C753C479A48D5DFFC7339A" pageId="0" pageNumber="221">Differential diagnosis.</paragraph>
|
||
<paragraph id="C7C9940C1EF173DEDD4B67A33EE691E2" pageId="0" pageNumber="221">
|
||
<taxonomicName id="5A371DAF3953164E0C71B8FD6CB755F9" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="DDD715470E88EA1FB72C8813D4E2AFD0" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from other West Palaearctic species of
|
||
<taxonomicName id="EEA85DA15C9BA17C938F39AA36FDF297" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FDD8C732EF4F69644B1EB7F0BAAEB39" italics="true" pageId="0" pageNumber="221">Alainites</emphasis>
|
||
</taxonomicName>
|
||
based on the combination of nymphal characters, summarized in Table
|
||
<tableCitation id="9E3587A05555AE67CBF4D9191E90173B" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Distinctive taxonomic criteria and distribution of West Palaearctic Alainites species." httpUri="http://table.plazi.org/id/B8A0F7B845D1C5A74FC4B5A69CE233EC" pageId="0" pageNumber="221" tableUuid="B8A0F7B845D1C5A74FC4B5A69CE233EC">2</tableCitation>
|
||
: (a) seven pairs of abdominal gills, (b) paraproct prolongation covered with small spines on its entire surface with broad, triangular spines laterally, (c) small teeth between prostheca and mola on both mandibles, (d) low number of dorsal setae on its fore-femora (10-15).
|
||
</paragraph>
|
||
<caption id="B8A0F7B845D1C5A74FC4B5A69CE233EC" ID-Table-UUID="B8A0F7B845D1C5A74FC4B5A69CE233EC" httpUri="http://table.plazi.org/id/B8A0F7B845D1C5A74FC4B5A69CE233EC" pageId="0" pageNumber="221" start="Table 2" startId="T2">
|
||
<paragraph id="C78CEE1E30E2D9720AF768BA13872E5A" pageId="0" pageNumber="221">
|
||
<emphasis id="390B0E5340A122BD41E007899883A230" bold="true" pageId="0" pageNumber="221">Table 2.</emphasis>
|
||
Distinctive taxonomic criteria and distribution of West Palaearctic
|
||
<taxonomicName id="025D2C81FFA6882C5FA9358465F926D5" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ADF3DA02521D1BDA96D386EE0618DE5A" italics="true" pageId="0" pageNumber="221">Alainites</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA86A86407D32C5C1F164CAA5B005FF6" pageId="0" pageNumber="221">
|
||
<table id="CF2B2C01304278E61B8BD0B5B573D5DA" pageId="0" pageNumber="221">
|
||
<tr id="1F1F5A20828E7680FE875D4AB5396E1E" pageId="0" pageNumber="221">
|
||
<th id="6FE150F0E1B18A3072F05FDB6038B1E3" colspan="1" pageId="0" pageNumber="221" rowspan="1">Species</th>
|
||
<th id="880DC4E441EEC2E16ED3795A8A8CE81A" colspan="1" pageId="0" pageNumber="221" rowspan="1">Distribution</th>
|
||
<th id="AD769E1E763D5BD62141AB4A071EB6B8" colspan="1" pageId="0" pageNumber="221" rowspan="1">Number of gill pairs</th>
|
||
<th id="59D0198105D7093172302B18EB17D416" colspan="1" pageId="0" pageNumber="221" rowspan="1">Right mandible: margin between prostheca and mola</th>
|
||
<th id="ADD5323A7FA15A015963E88B0DBD6BDE" colspan="1" pageId="0" pageNumber="221" rowspan="1">Mandible lateral side</th>
|
||
<th id="690064829BDA99F97551F57024FDD61C" colspan="1" pageId="0" pageNumber="221" rowspan="1">Fore-femur dorsal margin: setae number</th>
|
||
<th id="0DE6B89422BAF55EB1AD5C9FF5F5E6FD" colspan="1" pageId="0" pageNumber="221" rowspan="1">Fore-tibia dorsal margin: setae number</th>
|
||
<th id="0490154AF9E5CBAA4A96CAE3EFFEABFD" colspan="1" pageId="0" pageNumber="221" rowspan="1">Cuticle abdominal terga and sterna</th>
|
||
<th id="9D6CB1CD4DE04FDE5EA7885A100A817C" colspan="1" pageId="0" pageNumber="221" rowspan="1">Tergite IV: spines on distal margin</th>
|
||
<th id="D0BB1E5487F7CF918F5BA55F19550A08" colspan="1" pageId="0" pageNumber="221" rowspan="1">Prolongation of paraproct</th>
|
||
</tr>
|
||
<tr id="777EC3DAA4333C37E486EA1907BDB6DD" pageId="0" pageNumber="221">
|
||
<td id="8455EB3D8652DC2244DB0C231F463F52" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="C55C258C9515CDADA9AB109D1164D2B3" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="73DA544C1FE85B435AB7EA0E23949B44" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</td>
|
||
<td id="1F58F24359DC853891C631872ACB74EF" colspan="1" pageId="0" pageNumber="221" rowspan="1">Morocco</td>
|
||
<td id="62860A20C3905749331CA1D9D985881A" colspan="1" pageId="0" pageNumber="221" rowspan="1">7</td>
|
||
<td id="08A5B941199C578D02B36FFC858C1E3D" colspan="1" pageId="0" pageNumber="221" rowspan="1">10-16 small teeth</td>
|
||
<td id="B297AADDBFD25928F8ECDEA9AD768032" colspan="1" pageId="0" pageNumber="221" rowspan="1">shagreened with scattered fine setae and deep scale bases</td>
|
||
<td id="28FEBDFF16F4877299BB4CAA112F84AB" colspan="1" pageId="0" pageNumber="221" rowspan="1">10-15</td>
|
||
<td id="5C76E6E14EF921C377636430DAE518EA" colspan="1" pageId="0" pageNumber="221" rowspan="1">6-10</td>
|
||
<td id="3E182BADA2D451808A759EB02A8E9C2C" colspan="1" pageId="0" pageNumber="221" rowspan="1">strongly shagreened</td>
|
||
<td id="1598FFF053B1EB0AC01003B58E8D06B5" colspan="1" pageId="0" pageNumber="221" rowspan="1">long triangular, pointed</td>
|
||
<td id="8B9A9A5ACCE19C7984ABA55AFB4E6B5F" colspan="1" pageId="0" pageNumber="221" rowspan="1">covered by spines</td>
|
||
</tr>
|
||
<tr id="1824008D94C2C73ADECA415E82D380FD" pageId="0" pageNumber="221">
|
||
<td id="E6326D21126803F1E2719A02648E8712" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="FF90428EEC3E08C7D053B46843D7ADE7" baseAuthorityName="Sartori & Thomas" baseAuthorityYear="1989" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albinatii" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albinatii">
|
||
<emphasis id="86DC641FBADF1A54E3398E290CF90799" italics="true" pageId="0" pageNumber="221">Alainites albinatii</emphasis>
|
||
</taxonomicName>
|
||
(Sartori & Thomas, 1989)
|
||
</td>
|
||
<td id="C68DBF7AFC8AFC6F10FB177F9B503966" colspan="1" pageId="0" pageNumber="221" rowspan="1">Corsica</td>
|
||
<td id="3BAEC309FDF4DCE99A92F582C5A2375C" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="7FBFA09BA18CA0A80860488176E02E54" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 10 small teeth</td>
|
||
<td id="4DCA7BC9E3A0C3964714E27D50553712" colspan="1" pageId="0" pageNumber="221" rowspan="1">scale bases shagreened</td>
|
||
<td id="15AC6809E947F6D925B16FAFA75D0B74" colspan="1" pageId="0" pageNumber="221" rowspan="1">15</td>
|
||
<td id="AB0BC33EBDA970A59AE3BC27F2B62455" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="681747B27EC5E414ED92DCD249CB95CE" colspan="1" pageId="0" pageNumber="221" rowspan="1">slightly shagreened</td>
|
||
<td id="14294C97D868AB5CB84FD223D5313867" colspan="1" pageId="0" pageNumber="221" rowspan="1">long triangular, pointed</td>
|
||
<td id="07ADBC5F1740D9083CB30EDEEEF03BF6" colspan="1" pageId="0" pageNumber="221" rowspan="1">apically covered by spines</td>
|
||
</tr>
|
||
<tr id="17DD2079B126171860D6699D38595727" pageId="0" pageNumber="221">
|
||
<td id="CE33C240DB1F752AFE5D64131A7164FA" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="29106F2B3B5A8F4BE695481401A396F3" authorityName="Yanai & Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites bengunn" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="bengunn">
|
||
<emphasis id="50A64947A8AB8909DFE05F46FD4467B1" italics="true" pageId="0" pageNumber="221">Alainites bengunn</emphasis>
|
||
</taxonomicName>
|
||
Yanai & Gattolliat, 2022
|
||
</td>
|
||
<td id="7371E65FEDBF52301C8D5C2389B1088F" colspan="1" pageId="0" pageNumber="221" rowspan="1">Sardinia</td>
|
||
<td id="FAFA03CB5F5EEEB1045EBCEA6FA55E51" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="A2774C4234D4991F0E2C2D801DC2ECEB" colspan="1" pageId="0" pageNumber="221" rowspan="1">serrated</td>
|
||
<td id="124BC8AD031E22E80F065F5F6EB6553C" colspan="1" pageId="0" pageNumber="221" rowspan="1">scale bases, slightly shagreened</td>
|
||
<td id="2EDC5BA9308A3B7117437FF8A9CDCE63" colspan="1" pageId="0" pageNumber="221" rowspan="1">14-20</td>
|
||
<td id="C3D2FBDC752919E2C7BF923D2585DACE" colspan="1" pageId="0" pageNumber="221" rowspan="1">9-17</td>
|
||
<td id="97F8C1FAEC0976DC45677EC83A972595" colspan="1" pageId="0" pageNumber="221" rowspan="1">shagreened</td>
|
||
<td id="CC793D6B4484225DC198689263A43454" colspan="1" pageId="0" pageNumber="221" rowspan="1">slightly lanceolate</td>
|
||
<td id="2BB34A876668241B2310E0771CAE2016" colspan="1" pageId="0" pageNumber="221" rowspan="1">covered by spines</td>
|
||
</tr>
|
||
<tr id="B4D835D0C6D0FF0A5855E5E474FB9609" pageId="0" pageNumber="221">
|
||
<td id="43144FAEBEE562E541E83F754D2CF706" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="F299F5402B197AD0221C0C34DFB064D8" authorityName="Yanai & Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites gasithi" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="gasithi">
|
||
<emphasis id="A805348770E250DCEFA49A6DEA828B23" italics="true" pageId="0" pageNumber="221">Alainites gasithi</emphasis>
|
||
</taxonomicName>
|
||
Yanai & Gattolliat, 2022
|
||
</td>
|
||
<td id="05E5B72ED362CB864F5A4614FEB14E09" colspan="1" pageId="0" pageNumber="221" rowspan="1">Israel</td>
|
||
<td id="8A61875637A61B236FA4461AEA06D9E4" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="9F4947AAAB264BB7AF6AE57C11182E98" colspan="1" pageId="0" pageNumber="221" rowspan="1">serrated</td>
|
||
<td id="D1D96DF50A0B9A58033B9E79DC25C842" colspan="1" pageId="0" pageNumber="221" rowspan="1">no scale bases, almost not shagreened</td>
|
||
<td id="EBCE762F16B42B1463C7C04193942EA9" colspan="1" pageId="0" pageNumber="221" rowspan="1">10-20</td>
|
||
<td id="4B65EED9964FA4404EF461DE753D7141" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 6 rarely 10-12</td>
|
||
<td id="89E2ADB0929D540467AD5B2CC018AB92" colspan="1" pageId="0" pageNumber="221" rowspan="1">smooth</td>
|
||
<td id="CE4F633B2911F60091A411933708E408" colspan="1" pageId="0" pageNumber="221" rowspan="1">long triangular, pointed</td>
|
||
<td id="834744086AB4493493862A722ADB156C" colspan="1" pageId="0" pageNumber="221" rowspan="1">spines only on border</td>
|
||
</tr>
|
||
<tr id="67E6C6B41375C8895F62C16C52CA3981" pageId="0" pageNumber="221">
|
||
<td id="1D3585FD4AD3029C9E21950E99F7B99B" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="2DC7FFFDC7135EEF2483BD1C57CE8401" baseAuthorityName="Thomas & Kazanci" baseAuthorityYear="1989" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites kars" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="kars">
|
||
<emphasis id="62B02E8233053FB20073B87E8FBBC015" italics="true" pageId="0" pageNumber="221">Alainites kars</emphasis>
|
||
</taxonomicName>
|
||
(Thomas & Kazanci, 1989)
|
||
</td>
|
||
<td id="22AB7D8E779F87A46C71F0E4DF604F0D" colspan="1" pageId="0" pageNumber="221" rowspan="1">Turkey</td>
|
||
<td id="ADCF7225CB08FC079CAC12E1AF9260E2" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="E2E88869B5CFB90837EF726D9E795352" colspan="1" pageId="0" pageNumber="221" rowspan="1">teeth absent</td>
|
||
<td id="6DA0D7157A6DEA20ADF571E272AC3F19" colspan="1" pageId="0" pageNumber="221" rowspan="1">no scale bases, almost not shagreened</td>
|
||
<td id="5090A72DD5DD21B70E3D18A88DB942E7" colspan="1" pageId="0" pageNumber="221" rowspan="1">> 40 in two rows</td>
|
||
<td id="E5019881B9F1575E07B2F4BBA67BABD4" colspan="1" pageId="0" pageNumber="221" rowspan="1">5-9</td>
|
||
<td id="988433528C9AB4CB64AF05E25EE174A1" colspan="1" pageId="0" pageNumber="221" rowspan="1">slightly shagreened</td>
|
||
<td id="28C6A96454FB19252F63081782B89B77" colspan="1" pageId="0" pageNumber="221" rowspan="1">triangular pointed</td>
|
||
<td id="0F32C10AB270CC9B6FCCEED624334431" colspan="1" pageId="0" pageNumber="221" rowspan="1">spines on entire surface or just on apex</td>
|
||
</tr>
|
||
<tr id="A73EE50915027853A1C02735DA4E9B69" pageId="0" pageNumber="221">
|
||
<td id="5B861C5B71E6191596C043061C41F4D9" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="88B806806B2611F4E6E055934DEDF24D" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites muticus" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="muticus">
|
||
<emphasis id="CEF43D2699EF4D6C7494036A14A1ADBD" italics="true" pageId="0" pageNumber="221">Alainites muticus</emphasis>
|
||
</taxonomicName>
|
||
(Linnaeus, 1758)
|
||
</td>
|
||
<td id="B82768DC55CC5A5A2E7757A99518E6B3" colspan="1" pageId="0" pageNumber="221" rowspan="1">Palaearctic</td>
|
||
<td id="8A7E106BC7545F73D484F514A4C52CB8" colspan="1" pageId="0" pageNumber="221" rowspan="1">7</td>
|
||
<td id="5D2907D12E98225B095A501DF7CAE4D1" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 10 small teeth</td>
|
||
<td id="AE955CD3FF6B992120550541843E8CF7" colspan="1" pageId="0" pageNumber="221" rowspan="1">rare scale bases</td>
|
||
<td id="E23A988EC9F428AAEB25818958D88B17" colspan="1" pageId="0" pageNumber="221" rowspan="1">14</td>
|
||
<td id="15EE5DF7E540590129843F8DCB612549" colspan="1" pageId="0" pageNumber="221" rowspan="1">8</td>
|
||
<td id="45B1628421AA1F1BC82BFE257EE1855E" colspan="1" pageId="0" pageNumber="221" rowspan="1">slightly shagreened</td>
|
||
<td id="C8EBD71414C97D7B51CF15FB03EAFD5D" colspan="1" pageId="0" pageNumber="221" rowspan="1">short triangular broad basally</td>
|
||
<td id="BD7E475C5BD0474DD21EC75E1477309A" colspan="1" pageId="0" pageNumber="221" rowspan="1">spines only on border</td>
|
||
</tr>
|
||
<tr id="6655AE79248AE3DEAFE756205438BB74" pageId="0" pageNumber="221">
|
||
<td id="DC31FF7168540A97B0FB0D650D4E7204" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="4E5BAA1160FD954A0581BACA012AE3E1" baseAuthorityName="Muller-Liebenau" baseAuthorityYear="1974" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites navasi" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="navasi">
|
||
<emphasis id="B0E9A8BA87C691D56F7CD18F98BD542D" italics="true" pageId="0" pageNumber="221">Alainites navasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="20C6088DD49015313002E833206BC17F" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
|
||
, 1974)
|
||
</td>
|
||
<td id="62A025E9A20231FE70F0EC01558EAB12" colspan="1" pageId="0" pageNumber="221" rowspan="1">Iberian Peninsula</td>
|
||
<td id="23DF0AB4487284A528467B702E92778E" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="BAE4684F35FBDAFED2FD5B8E8ACFC505" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 10 small teeth</td>
|
||
<td id="E34EDCF7629109F074478C33D1E52D3F" colspan="1" pageId="0" pageNumber="221" rowspan="1">?</td>
|
||
<td id="EA891DD8F35F68544490A502077BB4D9" colspan="1" pageId="0" pageNumber="221" rowspan="1">26</td>
|
||
<td id="A364551CA0AE763B89783AA681867C39" colspan="1" pageId="0" pageNumber="221" rowspan="1">21</td>
|
||
<td id="F1D4276521E1DAA29D3551E961F18EB7" colspan="1" pageId="0" pageNumber="221" rowspan="1">smooth</td>
|
||
<td id="2CBF92A081EB5AC326E71D9B32AF4349" colspan="1" pageId="0" pageNumber="221" rowspan="1">short triangular</td>
|
||
<td id="B4E0802161EDF0617A8530671A7B0832" colspan="1" pageId="0" pageNumber="221" rowspan="1">covered by spines</td>
|
||
</tr>
|
||
<tr id="291E547E7ADEAEAA74708026DFDD6179" pageId="0" pageNumber="221">
|
||
<td id="9201FF992279B79BE9CE1447572A581B" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="91FA131676D4327EF4BB3F938AE2F376" baseAuthorityName="Thomas & Sartori" baseAuthorityYear="1991" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites oukaimeden" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="oukaimeden">
|
||
<emphasis id="B7050FC732BF1E02F446FD4DE7A39658" italics="true" pageId="0" pageNumber="221">Alainites oukaimeden</emphasis>
|
||
</taxonomicName>
|
||
(Thomas & Sartori, 1992)
|
||
</td>
|
||
<td id="FEB79855D6A252BD688EFE9AD3A1A711" colspan="1" pageId="0" pageNumber="221" rowspan="1">Morocco (High Atlas)</td>
|
||
<td id="EFA4BB0E683FB8781BD52BF163ED5962" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="2111445B65456589E6F350139CC2CC0A" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 10 small teeth</td>
|
||
<td id="D26ED4D0C905326BDA9E62C80F8A15B2" colspan="1" pageId="0" pageNumber="221" rowspan="1">shagreened</td>
|
||
<td id="34AF17E0C8A3055A9EA72B5254E2E2F9" colspan="1" pageId="0" pageNumber="221" rowspan="1">20</td>
|
||
<td id="C778D2BD60439B2C1930425E4B4D16DF" colspan="1" pageId="0" pageNumber="221" rowspan="1">8</td>
|
||
<td id="E9810047B7C3D9DBCABD922FCC15F288" colspan="1" pageId="0" pageNumber="221" rowspan="1">strongly shagreened</td>
|
||
<td id="F4BB0F8FBA1E39A8E749F1C67A35FD66" colspan="1" pageId="0" pageNumber="221" rowspan="1">long, relatively narrow</td>
|
||
<td id="F820BABEB5F3CBF341F19BFB3E4244FB" colspan="1" pageId="0" pageNumber="221" rowspan="1">covered by spines</td>
|
||
</tr>
|
||
<tr id="C23D148080CC692F95D5AFA499AE71C1" pageId="0" pageNumber="221">
|
||
<td id="78CD5A35621A998E943D9A573AE59CF7" colspan="1" pageId="0" pageNumber="221" rowspan="1">
|
||
<taxonomicName id="25B5781793911BB9F4FE50FF3D865EE5" authorityName="Thomas" authorityYear="1994" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites sadati" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="sadati">
|
||
<emphasis id="6B14D182FE08DE4B65304B62230FC955" italics="true" pageId="0" pageNumber="221">Alainites sadati</emphasis>
|
||
</taxonomicName>
|
||
Thomas, 1994
|
||
</td>
|
||
<td id="6A351A340636039A24E67AC448B50B11" colspan="1" pageId="0" pageNumber="221" rowspan="1">Algeria, Tunisia</td>
|
||
<td id="3BA40CF6B0F4F7579D4E8CE12EAD4BED" colspan="1" pageId="0" pageNumber="221" rowspan="1">6</td>
|
||
<td id="76954D0ADCBE04D04C65930CBB56FB31" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 10 small teeth</td>
|
||
<td id="F9EC987EC9541769AC0E48DC1A5FD23D" colspan="1" pageId="0" pageNumber="221" rowspan="1">no scale bases, almost not shagreened</td>
|
||
<td id="BE4592880899FC840201827B2AF8415D" colspan="1" pageId="0" pageNumber="221" rowspan="1">~ 25</td>
|
||
<td id="D2D0E85B67B4765214042F6CDC370C32" colspan="1" pageId="0" pageNumber="221" rowspan="1">6-9</td>
|
||
<td id="11ADB8CBAC65F5D55398E04951C224D5" colspan="1" pageId="0" pageNumber="221" rowspan="1">slightly shagreened</td>
|
||
<td id="E708001F22907AB5F81813DF61574C73" colspan="1" pageId="0" pageNumber="221" rowspan="1">medium triangular</td>
|
||
<td id="D7AFE18DC90B89C5A38EEC6FC0E9566F" colspan="1" pageId="0" pageNumber="221" rowspan="1">covered by spines</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ACDE884EE832A56CBBA891EAAC6AC941" pageId="0" pageNumber="221" type="description">
|
||
<paragraph id="4C70023CFFDF353D95957277E18EFE8F" pageId="0" pageNumber="221">Morphological description.</paragraph>
|
||
<paragraph id="4DC6E67045AC546DF0BEC69C8F00D25D" pageId="0" pageNumber="221">
|
||
<emphasis id="4CFDD24F5FB89108018E3ACE3BE36732" bold="true" pageId="0" pageNumber="221">
|
||
<emphasis id="D74DC7F0A34840087AD8644FBBA5838F" italics="true" pageId="0" pageNumber="221">Nymph</emphasis>
|
||
.
|
||
</emphasis>
|
||
Length. Female body 6.0-7.9 mm; cerci 4.5-5.5 mm; median caudal filament 1.3-1.4 mm (ca 2/3 of cerci); Male body 6.0-6.7 mm; cerci 4.0-5.0 mm; median caudal filament ca 2/3 of cerci.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8CBA40E6E1F9ACA69977B396F96E932E" pageId="0" pageNumber="221" type="coloration">
|
||
<paragraph id="BC5258B7BB6B13973342EAA2C5AB64CD" pageId="0" pageNumber="221">Coloration.</paragraph>
|
||
<paragraph id="51B1CAA87C8B0D092F73BA03D7955388" pageId="0" pageNumber="221">
|
||
General coloration pale to medium brown (Figs
|
||
<figureCitation id="63999E90D0ECC2889C090B71E3A4A907" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alainites albai sp. nov., nymph habitus a female, dorsal view b female lateral view c male, dorsal view d lateral view of the first three gills." figureDoi="10.3897/zookeys.1176.107829.figure2" httpUri="https://binary.pensoft.net/fig/898274" pageId="0" pageNumber="221">2</figureCitation>
|
||
,
|
||
<figureCitation id="14AB48E9EDD21032C8E2AD38FF4EABF8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3</figureCitation>
|
||
). Head uniformly pale brown to brown with yellow vermiform marks on vertex and frons (Fig.
|
||
<figureCitation id="C0B7B24E69E17ACD6E7985DA8991FCA1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3a</figureCitation>
|
||
). Turbinate eyes in male
|
||
<normalizedToken id="3B3B525038868E207E2E3E7C43229B1A" originalValue="nymph’s">nymph's</normalizedToken>
|
||
purple-brown. Legs ecru except upper side of femora brown (Fig.
|
||
<figureCitation id="A2211313A792B6B5EB9233A7770D978A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alainites albai sp. nov., nymph habitus a female, dorsal view b female lateral view c male, dorsal view d lateral view of the first three gills." figureDoi="10.3897/zookeys.1176.107829.figure2" httpUri="https://binary.pensoft.net/fig/898274" pageId="0" pageNumber="221">2b</figureCitation>
|
||
). Thorax with some paler clear pattern (Fig.
|
||
<figureCitation id="3449CFB893DE55B3123C00382FCA1480" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3c</figureCitation>
|
||
). Abdominal tergites pale brown with a central, elongated, yellowish dot; distal part of tergite IX and whole tergite X pale brown to yellowish. Abdominal sternites yellowish to pale brown. Cerci ecru to pale brown without bands or pattern.
|
||
</paragraph>
|
||
<caption id="30837534F6A5281A4A02AA36DB946ECC" doi="10.3897/zookeys.1176.107829.figure2" httpUri="https://binary.pensoft.net/fig/898274" pageId="0" pageNumber="221" start="Figure 2" startId="F2">
|
||
<paragraph id="99038B507DC61423D1FE5BF823DF7E6B" pageId="0" pageNumber="221">
|
||
<emphasis id="644F6F7FFE6FE75073A502718EBFEDCF" bold="true" pageId="0" pageNumber="221">Figure 2.</emphasis>
|
||
<taxonomicName id="0C7C876397551B858E73A3E7E31D6305" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="5F130C403A521313A2BEAFCF3E6EF648" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph habitus
|
||
<emphasis id="A3F204661B98225B73DB5CD60F23259F" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
female, dorsal view
|
||
<emphasis id="20234CF51411DFC8EF63660191DC14FE" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
female lateral view
|
||
<emphasis id="0E916A9C56D9F4BB4ADB3789F8A4795B" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
male, dorsal view
|
||
<emphasis id="DE62A9406D7D3FCF3951FE9E66EC4069" bold="true" pageId="0" pageNumber="221">d</emphasis>
|
||
lateral view of the first three gills.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F8B38FBC878391686E61369F7850FB10" doi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221" start="Figure 3" startId="F3">
|
||
<paragraph id="793CEAD992C92FC6809C8195BE045B40" pageId="0" pageNumber="221">
|
||
<emphasis id="B7CBAD80BC64BE60C26C9BDD9BF64A12" bold="true" pageId="0" pageNumber="221">Figure 3.</emphasis>
|
||
<taxonomicName id="2CC6E8CC966EBB7C71813331ABF8D4DC" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="405345122B7D4D11AD90B88E57B29CA1" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph morphology
|
||
<emphasis id="DCCFDA8646C5FF44E703AFA2105F8E66" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
head frontal view
|
||
<emphasis id="99F3030D6D2AC1F05A3B05D59406E1A3" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
scape and pedicel
|
||
<emphasis id="80F7FA08D110E4A69FEE8340484E4B70" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
pronotum
|
||
<emphasis id="60EB90C3577DA6A6136733E9196F99F0" bold="true" pageId="0" pageNumber="221">d</emphasis>
|
||
labrum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="95F99E17F8FFDC4E8271825C1A050593" pageId="0" pageNumber="221">
|
||
<emphasis id="F4D7F011FF683D44DC63F1EF0D96D7BF" bold="true" pageId="0" pageNumber="221">
|
||
Head.
|
||
<emphasis id="02AF5838DB5AFE1D2EADE0C327FFE983" italics="true" pageId="0" pageNumber="221">Antennae</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="66FABDD16F896E2617F36004943A5B0F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3a</figureCitation>
|
||
) close to each other, with a narrow inter-antennal carina, scape with deep scale insertions, pedicel with deep scale insertions and few setae (Fig.
|
||
<figureCitation id="15C379A0C2E304383FE1CD0A5C5B7B07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A4A5F084601FFAD2F38BA5AFBC0F2DC" pageId="0" pageNumber="221">
|
||
Dorsal surface of labrum (Fig.
|
||
<figureCitation id="CB0CBE983B8D84E518CCE3AE3BCBF3BB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Alainites albai sp. nov., nymph morphology a head frontal view b scape and pedicel c pronotum d labrum." figureDoi="10.3897/zookeys.1176.107829.figure3" httpUri="https://binary.pensoft.net/fig/898275" pageId="0" pageNumber="221">3d</figureCitation>
|
||
) with one central long seta and distolateral arc of two long, simple, stout setae, and small fine setae scattered on surface; ventral surface with 5-8 submarginal small, pointed setae; distal margin fringed with ca 20 short, followed by eight or ten long, feathered setae.
|
||
</paragraph>
|
||
<paragraph id="B9975E4E5E88E6F0C1725C7DFAA14DD2" pageId="0" pageNumber="221">
|
||
<emphasis id="61BDDC2B949DA24EBA12E4B522C511ED" bold="true" italics="true" pageId="0" pageNumber="221">Right mandible</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2070090C76C971284FC354D461DFA8AF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Alainites albai sp. nov., nymph morphology a right mandible b spines between mola and prostheca bifid c left mandible d spines between mola and prostheca." figureDoi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221">4a</figureCitation>
|
||
) shagreened, with scattered fine setae and deep scale insertions; incisors composed of eight blunt, distinct denticles, outer- and inner denticles notably smaller than others; prostheca reduced and bifid with numerous thin setae; 10-16 teeth in the space between prostheca and mola (Fig.
|
||
<figureCitation id="53E495F52BE85E7A5A0548A60F5B7007" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Alainites albai sp. nov., nymph morphology a right mandible b spines between mola and prostheca bifid c left mandible d spines between mola and prostheca." figureDoi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221">4b</figureCitation>
|
||
), tuft of setae absent.
|
||
</paragraph>
|
||
<caption id="C7C21CAE88CB68308178CBED5360984A" doi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221" start="Figure 4" startId="F4">
|
||
<paragraph id="63F6DAC9733D973F62F2F3186E6181F9" pageId="0" pageNumber="221">
|
||
<emphasis id="560B1041B0A5E2E4DC592C129C0A6402" bold="true" pageId="0" pageNumber="221">Figure 4.</emphasis>
|
||
<taxonomicName id="E8D3225C1B1BDC41451EF38C150D4AAD" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="6DF0694C6B11A99FC6A44A1DEBE7D4A0" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph morphology
|
||
<emphasis id="96490032BF8B26826056267D5D57A1B5" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
right mandible
|
||
<emphasis id="91E83A701F451F350A5611012D74AF99" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
spines between mola and prostheca bifid
|
||
<emphasis id="37907FDBD53CBFFB3C620C92753FFB84" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
left mandible
|
||
<emphasis id="15AA7696E95FB4BD8AE1291854202519" bold="true" pageId="0" pageNumber="221">d</emphasis>
|
||
spines between mola and prostheca.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A09A9973BE2B5772F2699E146E163787" pageId="0" pageNumber="221">
|
||
<emphasis id="3DD6990705E917C3C6FEE4E84816FD9E" bold="true" italics="true" pageId="0" pageNumber="221">Left mandible</emphasis>
|
||
(Fig.
|
||
<figureCitation id="7A2528F110F0E5B6FFB5D7C36A8EE630" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Alainites albai sp. nov., nymph morphology a right mandible b spines between mola and prostheca bifid c left mandible d spines between mola and prostheca." figureDoi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221">4c</figureCitation>
|
||
) slightly shagreened, with sparse fine setae and deep scale insertions; incisors composed of seven blunt, distinct denticles, outer denticle conspicuously smaller than others; prostheca with one row of 11-15 medium denticles and a comb-like structure (Fig.
|
||
<figureCitation id="E943AF58AB3C96CA5DB307939CB60082" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Alainites albai sp. nov., nymph morphology a right mandible b spines between mola and prostheca bifid c left mandible d spines between mola and prostheca." figureDoi="10.3897/zookeys.1176.107829.figure4" httpUri="https://binary.pensoft.net/fig/898276" pageId="0" pageNumber="221">4d</figureCitation>
|
||
); edge between prostheca and mola with sparse spines particularly in apical half, without setae.
|
||
</paragraph>
|
||
<paragraph id="65210772E1BA15830A8F172EF5549931" pageId="0" pageNumber="221">
|
||
<emphasis id="99D944A284774D24DC2429C9511D2697" bold="true" italics="true" pageId="0" pageNumber="221">Hypopharynx</emphasis>
|
||
(Fig.
|
||
<figureCitation id="6DF6CD3D95A963D060C3ACBD3FB86A3C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Alainites albai sp. nov., nymph morphology a hypopharynx b maxilla c details of dentisetae and maxilla crown of setae d labium." figureDoi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221">5a</figureCitation>
|
||
) trilobed, apically covered with thin setae; lingua with small central protuberance; superlingua longer than lingua.
|
||
</paragraph>
|
||
<paragraph id="EC0DD81AE8614718B518DF261878559A" pageId="0" pageNumber="221">
|
||
<emphasis id="0DA142F3732DA6D4A51358221BF71DD6" bold="true" italics="true" pageId="0" pageNumber="221">Maxilla</emphasis>
|
||
(Fig.
|
||
<figureCitation id="3B4E96204F27C89E052168209657877D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Alainites albai sp. nov., nymph morphology a hypopharynx b maxilla c details of dentisetae and maxilla crown of setae d labium." figureDoi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221">5b</figureCitation>
|
||
) apex with three elongated acute and curved teeth and a toothlike dentisetae; crown with one row of small setae ending with stouter and longer ones (Fig.
|
||
<figureCitation id="E9756D5C85E23A5A11B8E57F87C09539" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Alainites albai sp. nov., nymph morphology a hypopharynx b maxilla c details of dentisetae and maxilla crown of setae d labium." figureDoi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221">5c</figureCitation>
|
||
); palp two-segmented, extending behind apex of galea-lacinia, length of segment I approximately 0.65
|
||
<normalizedToken id="EC4AC6671D1BB0E847F1725CBE126F6B" originalValue="×">x</normalizedToken>
|
||
segment II; segment II apically rounded, with few thin setae.
|
||
</paragraph>
|
||
<caption id="AC17A4D2661D40AA563303D2E8A6AA33" doi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221" start="Figure 5" startId="F5">
|
||
<paragraph id="DCF231916E565392D4F122D1C55E0446" pageId="0" pageNumber="221">
|
||
<emphasis id="AF6B7C31C16158FD318603BF064FDB3A" bold="true" pageId="0" pageNumber="221">Figure 5.</emphasis>
|
||
<taxonomicName id="E137516EA797FE675A760669AA8DF0E6" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="E2A891536D091570B6110735796AB2C8" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph morphology
|
||
<emphasis id="FC401EAE24D208F7A6CB79853DB93399" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
hypopharynx
|
||
<emphasis id="70A4E9ECEEA5D1403A42FEE2B851A04A" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
maxilla
|
||
<emphasis id="268B8FE1C038A7F44D7645B48F8905D2" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
details of dentisetae and maxilla crown of setae
|
||
<emphasis id="FE9C991A787024637F9F304B37964401" bold="true" pageId="0" pageNumber="221">d</emphasis>
|
||
labium.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9FFE7A8192D47B234108CEDAC60342DE" pageId="0" pageNumber="221">
|
||
<emphasis id="25566AEF5EF422526D9678C909555619" bold="true" italics="true" pageId="0" pageNumber="221">Labium</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2AEFE27D1CD68B6BEAC6E4DF270AF0FB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Alainites albai sp. nov., nymph morphology a hypopharynx b maxilla c details of dentisetae and maxilla crown of setae d labium." figureDoi="10.3897/zookeys.1176.107829.figure5" httpUri="https://binary.pensoft.net/fig/898277" pageId="0" pageNumber="221">5d</figureCitation>
|
||
) with glossae slightly shorter than paraglossae; inner margins of glossae with 7 or 9 stout medium setae, apical margin with ca 5-7 long stout setae, ventral surface with few thin scattered setae; dorsal surface with row of ca 6-9 medium setae; paraglossae of constant width, with three rows of 11 or 14 long, stout and feathery setae apically; labial palp three-segmented; segment I 0.8
|
||
<normalizedToken id="D4571F7F23D99686F09C349D75CB0EF3" originalValue="×">x</normalizedToken>
|
||
length of segments II and III combined; segment II with dorsal oblique row of 4-6 medium setae; segment III conical, nearly symmetrical, slightly pointed apically, covered with few medium stout setae and small pointed setae apically.
|
||
</paragraph>
|
||
<paragraph id="17A65F1602AA6BE068E15CC678BBF5E6" pageId="0" pageNumber="221">
|
||
<emphasis id="3967FFEF2A617A7AA613C372716AA792" bold="true" pageId="0" pageNumber="221">
|
||
<emphasis id="136ECA6D24A02607C326C11565A731F4" italics="true" pageId="0" pageNumber="221">Thorax</emphasis>
|
||
.
|
||
<emphasis id="F8B2B44D6E71A2CC984FC56142BBB958" italics="true" pageId="0" pageNumber="221">Forelegs</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="9CD277C12A5E91F57CDC82EED279D961" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">6</figureCitation>
|
||
): Trochanter with seven marginal short stout pointed setae; femur dorsal margin with one regular row of 10-15 long, robust setae, and two subapical stout and long setae; ventral margin with short stout pointed setae; lateral surface with sparse scale bases, mainly on apical half and along subdorsal area (Fig.
|
||
<figureCitation id="F80A140900D774A1766C842C75D1AF0F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">6a</figureCitation>
|
||
). Tibiae (Fig.
|
||
<figureCitation id="686D03A030A156D601582205FF69FE31" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">6b</figureCitation>
|
||
) dorsally with 6-10 short stout pointed setae and single apical seta; ventrally with marginal and submarginal short stout pointed setae, denser towards apical end; tibiopatellar suture absent; lateral surface with few short, stout, pointed setae and numerous scale bases. Tarsi bares dorsally; ventral margin with ca 15-24 pointed medium setae; lateral surface with numerous scale bases. Tarsal claws (Fig.
|
||
<figureCitation id="63A2814E83258909614F389E76CED11D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">6c</figureCitation>
|
||
) hooked with one row of 11-13 medium teeth, apical setae absent (Fig.
|
||
<figureCitation id="5DD54DB1ABD3B1268E6C52FE540647A3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Alainites albai sp. nov., nymph morphology a foreleg femur b foreleg tibia c foreleg tarsal claw, foreleg claw." figureDoi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221">6d</figureCitation>
|
||
). Mid and hindlegs similar to forelegs except femora dorsally with 13-17 pointed setae and tibiae with 21-24 setae dorsally.
|
||
</paragraph>
|
||
<caption id="0630CD893FE58B818CF342F811E02EE5" doi="10.3897/zookeys.1176.107829.figure6" httpUri="https://binary.pensoft.net/fig/898278" pageId="0" pageNumber="221" start="Figure 6" startId="F6">
|
||
<paragraph id="C3A6471194FEF61E6028AA7332862884" pageId="0" pageNumber="221">
|
||
<emphasis id="C1965DE728C86CA5726BA20AF9A7FB8D" bold="true" pageId="0" pageNumber="221">Figure 6.</emphasis>
|
||
<taxonomicName id="30ED53A4BDE240154655651DAC89F703" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="F78E05C2932EA9545F67199D3D6207E4" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph morphology
|
||
<emphasis id="E4466F505242BEA3A480485C95D5B683" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
foreleg femur
|
||
<emphasis id="B6FA663EB5A059DDC54898921128933D" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
foreleg tibia
|
||
<emphasis id="EA5F448BDD73C7D8B8F6C661A463072B" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
foreleg tarsal claw, foreleg claw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C72FC80F93D6E23E659E779F3A3A06B3" pageId="0" pageNumber="221">
|
||
<emphasis id="5F87E404B8F89A980CC340A900F27647" bold="true" italics="true" pageId="0" pageNumber="221">Hindwing.</emphasis>
|
||
Pads present.
|
||
</paragraph>
|
||
<paragraph id="6416BF305EE8EF17709324B18A1830DD" pageId="0" pageNumber="221">
|
||
<emphasis id="48929EB0BF183B26BACAB2287814C12E" bold="true" pageId="0" pageNumber="221">
|
||
<emphasis id="3EF978B06ABA863E8E0C7FC2E0A56AA4" italics="true" pageId="0" pageNumber="221">Abdomen</emphasis>
|
||
.
|
||
<emphasis id="8972AA1C2C61E67AEE81D37B0FBAF9CE" italics="true" pageId="0" pageNumber="221">Terga</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="DEDFE74854CDED9EDCF8B29C97ECD7CC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Alainites albai sp. nov., nymph morphology a distal margin of tergum IV b gill IV c gill V d left paraproct e right paraproct." figureDoi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221">7a</figureCitation>
|
||
) shagreened, with numerous scale bases, distal margin of tergite IV with triangular spines ~ 2
|
||
<normalizedToken id="B22BED0E91B499CEA59644681047751A" originalValue="×">x</normalizedToken>
|
||
longer than broad. Sternites with scale bases; posterior margin smooth without spination.
|
||
</paragraph>
|
||
<caption id="403AFF1750DB5D447684ABA7443C7AF9" doi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221" start="Figure 7" startId="F7">
|
||
<paragraph id="C4AAA932D845780721AA9003D789E2E3" pageId="0" pageNumber="221">
|
||
<emphasis id="86FEAB3FD8E2E2630B4F654EFD6C4236" bold="true" pageId="0" pageNumber="221">Figure 7.</emphasis>
|
||
<taxonomicName id="BBF5D2AD2B5A3B978B0CAD491A126561" authorityName="El Alami, Vuataz & Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="9514BA10F463F06D739150AB5C911857" italics="true" pageId="0" pageNumber="221">Alainites albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., nymph morphology
|
||
<emphasis id="512F945636D576BFA5AF5976130A188B" bold="true" pageId="0" pageNumber="221">a</emphasis>
|
||
distal margin of tergum IV
|
||
<emphasis id="05B25E46703F42D6E5A5D3E4C323B47F" bold="true" pageId="0" pageNumber="221">b</emphasis>
|
||
gill IV
|
||
<emphasis id="2C8E9AD8569FDB433AFEA2406C75F338" bold="true" pageId="0" pageNumber="221">c</emphasis>
|
||
gill V
|
||
<emphasis id="E079E10A54BF987843558BED8F0C9E16" bold="true" pageId="0" pageNumber="221">d</emphasis>
|
||
left paraproct
|
||
<emphasis id="D01FEEC8B422C3AACE7C91120236B1EB" bold="true" pageId="0" pageNumber="221">e</emphasis>
|
||
right paraproct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="63701BBE58F2A8F0114CA7411EF30E5E" pageId="0" pageNumber="221">
|
||
<emphasis id="3D5882E6823BC111D016473C4FF5D8D3" bold="true" italics="true" pageId="0" pageNumber="221">Gills</emphasis>
|
||
(Fig.
|
||
<figureCitation id="58A6AB4A3B07C8C2A5FDB5836959242B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Alainites albai sp. nov., nymph habitus a female, dorsal view b female lateral view c male, dorsal view d lateral view of the first three gills." figureDoi="10.3897/zookeys.1176.107829.figure2" httpUri="https://binary.pensoft.net/fig/898274" pageId="0" pageNumber="221">2d</figureCitation>
|
||
) on segments I-VII, elliptic, almost symmetrical and serrated all along costal and distal margins except proximal part and with few setae, gill I smaller than others, length of gill I longer than half of tergite II; gill IV (Fig.
|
||
<figureCitation id="E9B44D087CB2C41FF7381100764B0C6C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Alainites albai sp. nov., nymph morphology a distal margin of tergum IV b gill IV c gill V d left paraproct e right paraproct." figureDoi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221">7b</figureCitation>
|
||
) and gill V larger than others, (Fig.
|
||
<figureCitation id="8A12BCB9C9FB052DC481CE746E2C5B4F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Alainites albai sp. nov., nymph morphology a distal margin of tergum IV b gill IV c gill V d left paraproct e right paraproct." figureDoi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221">7c</figureCitation>
|
||
). Surface with pores and without any setae; tracheation well visible but with reduced ramification. Paraproct (Fig.
|
||
<figureCitation id="7F398D6F3214FEA616D703530C4D1AB6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Alainites albai sp. nov., nymph morphology a distal margin of tergum IV b gill IV c gill V d left paraproct e right paraproct." figureDoi="10.3897/zookeys.1176.107829.figure7" httpUri="https://binary.pensoft.net/fig/898279" pageId="0" pageNumber="221">7d, e</figureCitation>
|
||
) shagreened with scale bases and setae; margin with seven broad, triangular spines inner to prolongation and numerous medium spines outer to prolongation; lateral margins of prolongation with numerous medium to broad spines, surface covered with spines; cercotractor with scattered scale bases, distal margin with ca 20 medium spines.
|
||
</paragraph>
|
||
<paragraph id="07B27344A5E809BAB905303279A550F5" pageId="0" pageNumber="221">
|
||
<emphasis id="BD89BCDB20271B8F4736EEA9EF4DD566" bold="true" pageId="0" pageNumber="221">Imago.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E6F3792BBFFEEA8815062EDB41E9A189" pageId="0" pageNumber="221" type="molecular analysis">
|
||
<paragraph id="A5094562BAD30074BD93AB9AD7078BF7" pageId="0" pageNumber="221">Molecular analysis.</paragraph>
|
||
<paragraph id="51B6A6FA6CBBF9076633685BDCC500AB" pageId="0" pageNumber="221">
|
||
The COI data set was> 95% complete and included 35% of parsimony informative sites. Pairwise COI distances across all sequences ranged from 0% to 23.4%. The overall mean p-distance within MOTUs was 0.8% (mean range: 0.1%-2.3%), while the overall mean p-distance between MOTUs was 18.7% (mean range: 14.9%-22.6%). The maximum p-distance within MOTUs varied from 0.2% (
|
||
<taxonomicName id="7C41D5F8ABCD2683A89A4AD35E9626D8" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. gasithi" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="gasithi">
|
||
<emphasis id="A72086415A8FF63C59E4805E9B3E7857" italics="true" pageId="0" pageNumber="221">A. gasithi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47309691E7BDE316C4CE241BFA477D4E" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. kars" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="kars">
|
||
<emphasis id="F9670D2917E402D6B7ED18893151F0A0" italics="true" pageId="0" pageNumber="221">A. kars</emphasis>
|
||
</taxonomicName>
|
||
) to 2.8% (
|
||
<taxonomicName id="0FBBADE83B6179F04753F0C444C158FB" class="Insecta" genus="A." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A." order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4EA4A7FE780B98A6C63F8C93F67052F3" italics="true" pageId="0" pageNumber="221">A.</emphasis>
|
||
</taxonomicName>
|
||
sp. 2). The minimum p-distance between MOTUs ranged from 14.3% (
|
||
<taxonomicName id="9A117A320AF81169174AE6925206A35A" class="Insecta" genus="A." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A." order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E742BF66783628225439C8CA1C3819E3" italics="true" pageId="0" pageNumber="221">A.</emphasis>
|
||
</taxonomicName>
|
||
sp. 1-
|
||
<taxonomicName id="475F1DA7F8CC129D2E39515811AC0E07" class="Insecta" genus="A." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A." order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="08FFEA5C5553D3B2F34DB891F0F39DA5" italics="true" pageId="0" pageNumber="221">A.</emphasis>
|
||
</taxonomicName>
|
||
sp. 2) to 16.7% (
|
||
<taxonomicName id="219AE78802AFF4D018EF694CABFB31D0" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. kars" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="kars">
|
||
<emphasis id="A58F8C2773AFFD89AA76E6C22ADE5CAF" italics="true" pageId="0" pageNumber="221">A. kars</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName id="27950FC7FD88B8FF43D561D721F97425" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. gasithi" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="gasithi">
|
||
<emphasis id="03566CF20D5C3E680A488C524BE0FA16" italics="true" pageId="0" pageNumber="221">A. gasithi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2A4533DEB58F0A665284397A0F85569C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. kars" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="kars">
|
||
<emphasis id="9E6FA2E7C898870FA4C76E9D6F7D1CF7" italics="true" pageId="0" pageNumber="221">A. kars</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName id="45070F915023877D00652826362510ED" class="Insecta" genus="A." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A." order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9B79B4DF725061BB9D54C678C68AC731" italics="true" pageId="0" pageNumber="221">A.</emphasis>
|
||
</taxonomicName>
|
||
sp. 4). The seven sequences from
|
||
<taxonomicName id="45FAB9282429182C3148F3B656CC5D28" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. albai" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="species" species="albai">
|
||
<emphasis id="76B9835D1D746AC15FC0B27CA5791E84" italics="true" pageId="0" pageNumber="221">A. albai</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. formed a strongly supported monophyletic clade, identified as a distinct MOTU in the three species delimitation analyses (Fig.
|
||
<figureCitation id="45A934745CD4F5709E913BF7176B6415" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bayesian (BEAST) maximum clade credibility COI tree of the genus Alainites in the West Palearctic: branch ends labelled with GBIF codes in bold indicate newly sequenced specimens; the DTNHM 1449 - 21, TDAAT 323 - 19, and VMCOL 019 - 20 - VMCOL 021 - 20 codes are from BOLD; the other codes correspond to sequences obtained from GenBank. Colored vertical boxes indicate species delimitation hypothesis according to the ASAP, mPTP, and GMYC methods. For each MOTU, the corresponding species names (where available) and the country / region of origin is provided, with the newly described species and associated GBIF codes specified in bold. Circles on branches indicate Bayesian posterior probabilities> 0.9. Outgroup branches, labels, and species names are presented in grey." figureDoi="10.3897/zookeys.1176.107829.figure8" httpUri="https://binary.pensoft.net/fig/898280" pageId="0" pageNumber="221">8</figureCitation>
|
||
). All methods were fully congruent in delimitating the other MOTUs as well.
|
||
</paragraph>
|
||
<caption id="7FEEF52371D75183A935FA0924199EBF" doi="10.3897/zookeys.1176.107829.figure8" httpUri="https://binary.pensoft.net/fig/898280" pageId="0" pageNumber="221" start="Figure 8" startId="F8">
|
||
<paragraph id="3932965189E77DC5546DB919ECFCB00E" pageId="0" pageNumber="221">
|
||
<emphasis id="C8C7A2C25037099E262762B59CEF40E8" bold="true" pageId="0" pageNumber="221">Figure 8.</emphasis>
|
||
Bayesian (BEAST) maximum clade credibility COI tree of the genus
|
||
<taxonomicName id="EC586C2308781C981C16558D7636BC1B" class="Insecta" family="Baetidae" genus="Alainites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alainites" order="Ephemeroptera" pageId="0" pageNumber="221" phylum="Arthropoda" rank="genus">Alainites</taxonomicName>
|
||
in the West Palearctic: branch ends labelled with GBIF codes in bold indicate newly sequenced specimens; the DTNHM1449-21, TDAAT323-19, and VMCOL019-20-VMCOL021-20 codes are from BOLD; the other codes correspond to sequences obtained from GenBank. Colored vertical boxes indicate species delimitation hypothesis according to the ASAP, mPTP, and GMYC methods. For each MOTU, the corresponding species names (where available) and the country/region of origin is provided, with the newly described species and associated GBIF codes specified in bold. Circles on branches indicate Bayesian posterior probabilities> 0.9. Outgroup branches, labels, and species names are presented in grey.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="39D7DB11EA1B0C69D6F5365906BDEEFF" pageId="0" pageNumber="221" type="distribution">
|
||
<paragraph id="083940429B9A4D868D07EA63F5186928" pageId="0" pageNumber="221">Distribution</paragraph>
|
||
<paragraph id="77376ABA3ECFA9839ADF23D441B9F599" pageId="0" pageNumber="221">
|
||
(Fig.
|
||
<figureCitation id="9D9542B80647A008358914B10829ADEE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The sampling site localization of Alainites albai sp. nov., in the Rif domain." figureDoi="10.3897/zookeys.1176.107829.figure1" httpUri="https://binary.pensoft.net/fig/898273" pageId="0" pageNumber="221">1</figureCitation>
|
||
). Morocco.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="529015DC0F8932B69C00CF4AF2A36C98" pageId="0" pageNumber="221" type="etymology">
|
||
<paragraph id="87B1EFF61E933F98C5C30C1B663185D0" pageId="0" pageNumber="221">Etymology.</paragraph>
|
||
<paragraph id="5B1209B404D375E367062C51406F89FE" pageId="0" pageNumber="221">
|
||
The first author dedicates the name of this species to her former mentor, Professor Javier Alba-Tercedor, a specialist in the
|
||
<taxonomicName id="A99AA5C795D0ECE4B1988897C1CE7593" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="221" rank="order">Ephemeroptera</taxonomicName>
|
||
of the Iberian Peninsula. He contributed significantly to her training and specialization in the taxonomic study of the
|
||
<taxonomicName id="5B6A243B1AAD6EB32624886826168690" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="221" rank="order">Ephemeroptera</taxonomicName>
|
||
of Morocco.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |