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<document id="BE992A3030F86647A813149E39FD0EAF" ID-DOI="10.5281/zenodo.157906" ID-GBIF-Dataset="1747e372-76a3-4a7d-b64c-fbbfa0b7e9c8" ID-ISSN="1175­5326" ID-Zenodo-Dep="157906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461023883182" checkinUser="plazi" docAuthor="Saether, Ole A." docDate="2004" docId="125B084CFFC5FFFE9261233CD812F490" docLanguage="en" docName="zt00595.pdf" docOrigin="Zootaxa 595" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Procladius (Holotanypus) crassinervis (Zetterstedt) sensu Pinder 1978" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="EE627034FFDCFFE79369210DDC13F11B" masterDocTitle="The chironomids (Diptera, Chironomidae) described by Lundström (1915) from arctic Siberia, with a redescription of Derotanypus sibiricus (Kruglova &amp; Chernovskii)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="26" updateTime="1698571168210" updateUser="plazi">
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<mods:title id="A0C6A8FCB416556D08CD925A7528D6F2">The chironomids (Diptera, Chironomidae) described by Lundström (1915) from arctic Siberia, with a redescription of Derotanypus sibiricus (Kruglova &amp; Chernovskii)</mods:title>
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<subSubSection id="D2E8EAD1FFC5FFFE9261233CD86BF350" box="[264,1144,561,587]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="9A4DB95AFFC5FFFE9261233CD86BF350" blockId="25.[264,1144,561,587]" box="[264,1144,561,587]" pageId="25" pageNumber="26">
<heading id="C1050E36FFC5FFFE9261233CD86BF350" bold="true" box="[264,1144,561,587]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="5DF2C2D9FFC5FFFE9261233CD86BF350" ID-CoL="898CM" authority="(Zetterstedt) sensu Pinder (1978)" authorityName="(Zetterstedt) sensu Pinder" authorityYear="1978" box="[264,1144,561,587]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crassinervis" subGenus="Holotanypus">
<emphasis id="A8866548FFC5FFFE9261233CD86BF350" bold="true" box="[264,1144,561,587]" pageId="25" pageNumber="26">
<emphasis id="A8866548FFC5FFFE9261233CDD99F350" bold="true" box="[264,394,561,587]" italics="true" pageId="25" pageNumber="26">Procladius</emphasis>
(
<emphasis id="A8866548FFC5FFFE92F5233CDE25F350" bold="true" box="[412,566,561,587]" italics="true" pageId="25" pageNumber="26">Holotanypus</emphasis>
)
<emphasis id="A8866548FFC5FFFE912E233CDEC4F350" bold="true" box="[583,727,561,587]" italics="true" pageId="25" pageNumber="26">crassinervis</emphasis>
(Zetterstedt) sensu
<bibRefCitation id="FE63C4ABFFC5FFFE90A7233CD86BF350" author="Pinder" box="[974,1144,561,587]" pageId="25" pageNumber="34" refString="Pinder, L. C. V. (1978) A key to adult males of British Chironomidae. Part 1. The key; part 2. Illustrations of the hypopygia. Freshwater Biological Association, Scientific Publication 37, 169 pp., 189 figs." type="book" year="1978">Pinder (1978)</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D2E8EAD1FFC5FFFE9261237FD812F490" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="9A4DB95AFFC5FFFE9261237FDEF3F393" blockId="25.[264,1322,625,749]" box="[264,736,625,649]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="BAE29E74FFC5FFFE9261237FDEF3F393" box="[264,736,625,649]" pageId="25" pageNumber="26">
<taxonomicName id="5DF2C2D9FFC5FFFE9261237FDECFF393" ID-CoL="898CM" authority="Zetterstedt, 1838: 817" authorityName="Zetterstedt" authorityPageNumber="817" authorityYear="1838" box="[264,732,625,649]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crassinervis">
<emphasis id="A8866548FFC5FFFE9261237FDDFBF392" box="[264,488,626,649]" italics="true" pageId="25" pageNumber="26">Tanypus crassinervis</emphasis>
<treatmentCitation id="1B539F4BFFC5FFFE9287237CDECFF393" author="Zetterstedt" box="[494,732,625,648]" page="817" pageId="25" pageNumber="26" year="1838">
<bibRefCitation id="FE63C4ABFFC5FFFE9287237CDEB5F393" author="Zetterstedt" box="[494,678,625,648]" pageId="25" pageNumber="35" refString="Zetterstedt, J. W. (1838) Diptera .. In: Insecta Lapponica, Sect. 3, 477 - 868." type="book chapter" year="1838">Zetterstedt, 1838</bibRefCitation>
: 817
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9A4DB95AFFC5FFFE9261239FDF8CF3F6" blockId="25.[264,1322,625,749]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="BAE29E74FFC5FFFE9261239FDF8CF3F6" infered="true" pageId="25" pageNumber="26">
<taxonomicName id="5DF2C2D9FFC5FFFE9261239FDF3DF3B3" ID-CoL="8FCV3" authority="Zett. sensu Lundstrom (1915: 24)" authorityName="Zett. sensu Lundstrom" authorityPageNumber="24" authorityYear="1915" box="[264,814,657,681]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="A8866548FFC5FFFE9261239FDDACF3B2" box="[264,447,658,681]" italics="true" pageId="25" pageNumber="26">Tanypus signatus</emphasis>
Zett. sensu
<treatmentCitation id="1B539F4BFFC5FFFE9129239CDF3DF3B3" author="Lundstrom" box="[576,814,657,680]" page="24" pageId="25" pageNumber="26" year="1915">
<bibRefCitation id="FE63C4ABFFC5FFFE9129239CDF3DF3B3" author="Lundstrom" box="[576,814,657,680]" pageId="25" pageNumber="34" refString="Lundstrom, C. (1915) Resultats scientifiques de lExpedition Polaire Russe en 1900 - 1903, sous la direction du Baron E. Toll. Section E: Zoologie. Volume II, livr. 8. Diptera Nematocera aus den arctischen Gegenden Sibiriens. Memoires de lAcademie Imperiale des Sciences, Classe Physico-Mathematique, VIII Serie, 29 (8), 1 - 33." type="journal article" year="1915">Lundström (1915: 24)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,?not
<bibRefCitation id="FE63C4ABFFC5FFFE9007239CD836F3B3" author="Zetterstedt" box="[878,1061,657,680]" pageId="25" pageNumber="35" refString="Zetterstedt, J. W. (1850) Diptera Scandinaviae disposita et descripta. Vol. 9, 3367 - 3710." type="journal article" year="1850">Zetterstedt, 1850</bibRefCitation>
.
<emphasis id="A8866548FFC5FFFE926123BEDDC1F3D1" box="[264,466,691,714]" italics="true" pageId="25" pageNumber="26">Material examined</emphasis>
.
<collectingCountry id="E2E5F9CAFFC5FFFE92B623BEDE51F3D1" box="[479,578,691,714]" name="Russia" pageId="25" pageNumber="26">RUSSIA</collectingCountry>
: New Siberian Islands, “Holzgebirge”, 2 ɗ,
<date id="EE4C9F9AFFC5FFFE977023BED89EF3D1" box="[1049,1165,691,714]" pageId="25" pageNumber="26" value="1903-06-23">23 vi 1903</date>
, upper reaches of Wosnessenje River, 2 ɗ,
<date id="EE4C9F9AFFC5FFFE910823DBDEF8F3F6" box="[609,747,726,749]" pageId="25" pageNumber="26" value="1903-07-03" valueMax="1903-07-04" valueMin="1903-07-03">34 vii 1903</date>
, M. I. Brussnew.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9A4DB95AFFC5FFFE92612214D812F490" blockId="25.[264,1323,793,1419]" pageId="25" pageNumber="26">
The genus
<taxonomicName id="5DF2C2D9FFC5FFFE92E22214DE4AF228" ID-CoL="9G2QS" authority="Skuse" authorityName="Skuse" box="[395,601,793,819]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A8866548FFC5FFFE92E22214DE18F229" box="[395,523,793,818]" italics="true" pageId="25" pageNumber="26">Procladius</emphasis>
Skuse
</taxonomicName>
is in need of revision. According to
<bibRefCitation id="FE63C4ABFFC5FFFE97652214D8A2F228" author="Pinder" box="[1036,1201,793,819]" pageId="25" pageNumber="34" refString="Pinder, L. C. V. (1978) A key to adult males of British Chironomidae. Part 1. The key; part 2. Illustrations of the hypopygia. Freshwater Biological Association, Scientific Publication 37, 169 pp., 189 figs." type="book" year="1978">Pinder (1978)</bibRefCitation>
, however, the present specimens cannot belong to
<taxonomicName id="5DF2C2D9FFC5FFFE9183224CD843F240" ID-CoL="8FCV3" authority="Zetterstedt, 1850" authorityName="Zetterstedt" authorityYear="1850" box="[746,1104,833,859]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="A8866548FFC5FFFE9183224CDF78F241" box="[746,875,833,858]" italics="true" pageId="25" pageNumber="26">P. signatus</emphasis>
(
<bibRefCitation id="FE63C4ABFFC5FFFE9015224CD85BF240" author="Zetterstedt" box="[892,1096,833,859]" pageId="25" pageNumber="35" refString="Zetterstedt, J. W. (1850) Diptera Scandinaviae disposita et descripta. Vol. 9, 3367 - 3710." type="journal article" year="1850">Zetterstedt, 1850</bibRefCitation>
)
</taxonomicName>
since that species has a cluster of small teeth apically on the phallapodeme lacking in these specimens.
<taxonomicName id="5DF2C2D9FFC5FFFE967D2264DD86F2B1" ID-CoL="898CM" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crassinervis">
<emphasis id="A8866548FFC5FFFE967D2264D930F299" box="[1300,1315,873,898]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="A8866548FFC5FFFE9261229CDD86F2B1" box="[264,405,913,938]" italics="true" pageId="25" pageNumber="26">crassinervis</emphasis>
</taxonomicName>
was regarded as a synonym of
<taxonomicName id="5DF2C2D9FFC5FFFE9074229CD88DF2B0" ID-CoL="8DGS7" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[797,1182,913,939]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="culiciformis">
<emphasis id="A8866548FFC5FFFE9074229CDFDAF2B1" box="[797,969,913,938]" italics="true" pageId="25" pageNumber="26">P. culiciformis</emphasis>
(
<bibRefCitation id="FE63C4ABFFC5FFFE90B2229CD885F2B0" author="Linnaeus" box="[987,1174,913,939]" pageId="25" pageNumber="34" refString="Linnaeus, C. (1767) Systema naturae, per regna tria naturae, secundum classes, ordines, genera, species, cum carateribus, differentiis, synonymis, locis, Ed. 12. (rev.), Holmiae [= Stockholm] 1 (2), 533 - 1327." type="journal article" year="1767">Linnaeus, 1767</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="FE63C4ABFFC5FFFE97B8229CDD8FF2C8" author="Roback" pageId="25" pageNumber="34" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback (1971: 175)</bibRefCitation>
, whereas he regarded
<emphasis id="A8866548FFC5FFFE91D622B4DF2FF2C9" box="[703,828,953,978]" italics="true" pageId="25" pageNumber="26">P. c h o re u s</emphasis>
(
<bibRefCitation id="FE63C4ABFFC5FFFE903E22B4D817F2C8" author="Meigen" box="[855,1028,953,979]" pageId="25" pageNumber="34" refString="Meigen, J. W. (1804) Klassifikation und Beschreibung der europaischen zwei-flugeligen Insecten (Diptera Linn.). Braunschweig [= Brunswick], 1, 1 - 314." type="journal article" year="1804">Meigen, 1804</bibRefCitation>
) as a separate species. According to
<bibRefCitation id="FE63C4ABFFC5FFFE92DA22ECDE9DF2E0" author="Kobayashi" box="[435,654,993,1019]" pageId="25" pageNumber="33" refString="Kobayashi, T. (1998) Seasonal changes in body size and male genital structures of Procladius choreus (Diptera: Chironomidae: Tanypodinae). Aquatic Insects, 20, 165 - 172." type="journal article" year="1998">Kobayashi (1998)</bibRefCitation>
, however, male genital structures can vary seasonally, with the gonostylus having a longer heel in spring and fall than in summer. Kobayashi regards
<taxonomicName id="5DF2C2D9FFC5FFFE920E253CDE00F551" ID-CoL="898CM" box="[359,531,1073,1098]" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crassinervis">
<emphasis id="A8866548FFC5FFFE920E253CDE00F551" box="[359,531,1073,1098]" italics="true" pageId="25" pageNumber="26">P. crassinervis</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="5DF2C2D9FFC5FFFE918F253CDF73F551" ID-CoL="8DGS4" box="[742,864,1073,1098]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="choreus">
<emphasis id="A8866548FFC5FFFE918F253CDF73F551" box="[742,864,1073,1098]" italics="true" pageId="25" pageNumber="26">P. choreus</emphasis>
</taxonomicName>
, but he explicitly identified
<emphasis id="A8866548FFC5FFFE97D8253CD934F551" box="[1201,1319,1073,1098]" italics="true" pageId="25" pageNumber="26">P. c h o re u s</emphasis>
after
<bibRefCitation id="FE63C4ABFFC5FFFE922F2554DE15F568" author="Edwards" box="[326,518,1113,1139]" pageId="25" pageNumber="33" refString="Edwards, F. W. (1929) British non-biting midges (Diptera, Chironomidae). Transactions of the Royal Entomological Society of London, 77, 279 - 430." type="journal article" year="1929">Edwards (1929)</bibRefCitation>
, and the latter, according to
<bibRefCitation id="FE63C4ABFFC5FFFE90312554D818F568" author="Roback" box="[856,1035,1113,1139]" pageId="25" pageNumber="34" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback (1971)</bibRefCitation>
, was a misidentification of
<taxonomicName id="5DF2C2D9FFC5FFFE924E258CDDDCF581" ID-CoL="8DGS7" box="[295,463,1153,1178]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="culiciformis">
<emphasis id="A8866548FFC5FFFE924E258CDDDCF581" box="[295,463,1153,1178]" italics="true" pageId="25" pageNumber="26">P. culiciformis</emphasis>
</taxonomicName>
(L.). Thus, Kobayashi's (1998) &quot;
<taxonomicName id="5DF2C2D9FFC5FFFE9027258CDFD5F581" ID-CoL="8DGS4" box="[846,966,1153,1178]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="choreus">
<emphasis id="A8866548FFC5FFFE9027258CDFD5F581" box="[846,966,1153,1178]" italics="true" pageId="25" pageNumber="26">P. choreus</emphasis>
</taxonomicName>
&quot; is not necessarily in any conflict with what
<bibRefCitation id="FE63C4ABFFC5FFFE92AD25A4DE65F5D8" author="Roback" box="[452,630,1193,1219]" pageId="25" pageNumber="34" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback (1971)</bibRefCitation>
had treated under the name
<taxonomicName id="5DF2C2D9FFC5FFFE90BC25A4D892F5D9" ID-CoL="8DGS7" box="[981,1153,1193,1218]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="culiciformis">
<emphasis id="A8866548FFC5FFFE90BC25A4D892F5D9" box="[981,1153,1193,1218]" italics="true" pageId="25" pageNumber="26">P. culiciformis</emphasis>
</taxonomicName>
(L.). It is not known why
<bibRefCitation id="FE63C4ABFFC5FFFE92F225DCDE2CF5F0" author="Pinder" box="[411,575,1233,1259]" pageId="25" pageNumber="34" refString="Pinder, L. C. V. (1978) A key to adult males of British Chironomidae. Part 1. The key; part 2. Illustrations of the hypopygia. Freshwater Biological Association, Scientific Publication 37, 169 pp., 189 figs." type="book" year="1978">Pinder (1978)</bibRefCitation>
did not follow
<bibRefCitation id="FE63C4ABFFC5FFFE919125DCDFBAF5F0" author="Roback" box="[760,937,1233,1259]" pageId="25" pageNumber="34" refString="Roback, S. S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monographs of the Academy of Natural Sciences of Philadelphia, 17, 1 - 410." type="journal article" year="1971">Roback (1971)</bibRefCitation>
in the use of the name
<emphasis id="A8866548FFC5FFFE97AB25DCDD40F409" italics="true" pageId="25" pageNumber="26">P. c u l i c i ­ formis</emphasis>
(L.), but comparing the two authors' keys suggests that Pinder's &quot;
<taxonomicName id="5DF2C2D9FFC5FFFE971525F4DD8FF420" ID-CoL="898CM" authority="Zetterstedt" authorityName="Zetterstedt" class="Insecta" family="Chironomidae" genus="Tanypus" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crassinervis">
<emphasis id="A8866548FFC5FFFE971525F4D938F409" box="[1148,1323,1273,1298]" italics="true" pageId="25" pageNumber="26">P. crassinervis</emphasis>
(Zetterstedt)
</taxonomicName>
&quot; is Roback's
<taxonomicName id="5DF2C2D9FFC5FFFE912D242CDEE2F421" ID-CoL="8DGS7" box="[580,753,1313,1338]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="culiciformis">
<emphasis id="A8866548FFC5FFFE912D242CDEE2F421" box="[580,753,1313,1338]" italics="true" pageId="25" pageNumber="26">P. culiciformis</emphasis>
</taxonomicName>
(L.). Apparently consistent with this, it seems possible (judging from Kobayashi's work) that in fact there is only a single species in all of this. If so, then the name
<taxonomicName id="5DF2C2D9FFC5FFFE9159247CDECAF491" ID-CoL="8DGS7" box="[560,729,1393,1418]" class="Insecta" family="Chironomidae" genus="Procladius" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="culiciformis">
<emphasis id="A8866548FFC5FFFE9159247CDECAF491" box="[560,729,1393,1418]" italics="true" pageId="25" pageNumber="26">P. culiciformis</emphasis>
</taxonomicName>
(L.) would have priority.
</paragraph>
</subSubSection>
</treatment>
</document>