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123 lines
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<document id="6322EA68610CBC435A9B3E55A012318A" ID-DOI="10.11646/zootaxa.5403.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561570" ID-ZooBank="A9E9D993-74C3-49CC-9202-DB1132EEF113" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1706103461433" checkinUser="plazi" docAuthor="Mah, Christopher L., Kogure, Yoichi, Fujita, Toshihiko & Higashiji, Takuo" docDate="2024" docId="123887CCFFAAFFAEFF0EFDCF7F1A1173" docLanguage="en" docName="zootaxa.5403.1.1.pdf" docOrigin="Zootaxa 5403 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calliaster Gray 1840" docType="treatment" docVersion="1" lastPageNumber="3" masterDocId="EE01FFB4FFA8FFACFF99FFA57C731474" masterDocTitle="New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions" masterLastPageNumber="41" masterPageNumber="1" pageNumber="3" updateTime="1706202430652" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="C39F5934C9E0BCF5B5AAAA1774EC39EA">New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions</mods:title>
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<mods:name id="85F112E7E4CFD9AEAB294F25CBF10A0C" type="personal">
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<mods:namePart id="B0E373FABA099184B12D684A3A492BDC">Mah, Christopher L.</mods:namePart>
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<mods:affiliation id="0E409027C79F28414AD42934B1E0AB4F">Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D. C. USA</mods:affiliation>
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<mods:namePart id="C4B58C2483312BD80AAFCA606E2BF056">Kogure, Yoichi</mods:namePart>
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<mods:nameIdentifier id="4E4E6AC725CEFF9CA036CB6BF62D4ABB" type="ORCID">0009-0009-9935-8208</mods:nameIdentifier>
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<mods:affiliation id="D6F743A38C0AC2D9C373DEE831DD2F19">Fisheries Resources Institute, Japan Fisheries Research and Education Agency, 1 - 5939 - 22 Suido-cho, Chuo, Niigata, Niigata 951 - 8121, Japan</mods:affiliation>
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<mods:nameIdentifier id="4FE1541167E91AAC19921DDA62FDCBB7" type="email">kogure_yoichi52@fra.go.jp</mods:nameIdentifier>
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<mods:namePart id="F44B5E1E0F432FD7988C45CC6A05EB1A">Fujita, Toshihiko</mods:namePart>
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<mods:affiliation id="D3DBF9CF7B73BE99F611348A6BDC160B">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan</mods:affiliation>
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<mods:nameIdentifier id="C4FEE40E30EAEFF2F54F0E4A5B97B1F7" type="email">fujita@kahaku.go.jp</mods:nameIdentifier>
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<mods:namePart id="4FECD10D6A26C689F82658CC1E6C6A84">Higashiji, Takuo</mods:namePart>
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<mods:affiliation id="F92C6674A70116E5FC910250F6D2F7BB">Okinawa Churaumi Aquarium, 424 Ishikawa, Motobu, Okinawa 905 - 0206, Japan</mods:affiliation>
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<mods:nameIdentifier id="1FAE75EA06B658F106DF43362B44FC02" type="email">t-higashiji@okichura.jp</mods:nameIdentifier>
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<mods:title id="14C4CAC95BDE06281F96E001864BA75F">Zootaxa</mods:title>
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<mods:date id="7435E9B753BDAE28BC275E5E015CA5EA">2024</mods:date>
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<mods:number id="76F40DE442ED5D81E1CA5224E6C430A9">2024-01-18</mods:number>
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<mods:number id="EF72112AC1B96EE68989FD260ACD5094">5403</mods:number>
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<treatment id="123887CCFFAAFFAEFF0EFDCF7F1A1173" LSID="urn:lsid:plazi:treatment:123887CCFFAAFFAEFF0EFDCF7F1A1173" httpUri="http://treatment.plazi.org/id/123887CCFFAAFFAEFF0EFDCF7F1A1173" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="D28B6551FFAAFFAEFF0EFDCF7EE516F0" box="[151,662,618,645]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="9A2E36DAFFAAFFAEFF0EFDCF7EE516F0" blockId="2.[151,662,618,645]" box="[151,662,618,645]" pageId="2" pageNumber="3">
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<heading id="C16681B6FFAAFFAEFF0EFDCF7EE516F0" bold="true" box="[151,662,618,645]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="A8E5EAC8FFAAFFAEFF0EFDCF7EE516F0" bold="true" box="[151,662,618,645]" pageId="2" pageNumber="3">
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<taxonomicName id="5D914D59FFAAFFAEFF0EFDCF7DE416F0" ID-CoL="87DSF" authority="Gray 1840" authorityName="Gray" authorityYear="1840" box="[151,407,618,645]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="genus">
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<emphasis id="A8E5EAC8FFAAFFAEFF0EFDCF7D7F16F1" bold="true" box="[151,268,618,645]" italics="true" pageId="2" pageNumber="3">Calliaster</emphasis>
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<bibRefCitation id="FE004B2BFFAAFFAEFE8DFDCF7DE416F0" author="Gray, J. E." box="[276,407,618,644]" pageId="2" pageNumber="3" pagination="275 - 290" refId="ref18074" refString="Gray, J. E. (1840) XXXII. A synopsis of the genera and species of the class Hypostoma (Asterias, Linnaeus). Annals of the Magazine of Natural History, 6, 275 - 290. https: // doi. org / 10.1080 / 03745484009443296" type="journal article" year="1840">Gray 1840</bibRefCitation>
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</taxonomicName>
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(
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<taxonomicName id="5D914D59FFAAFFAEFE31FDCF7E6E16F1" authorityName="Gray" authorityYear="1840" box="[424,541,618,645]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="genus">
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<emphasis id="A8E5EAC8FFAAFFAEFE31FDCF7E6E16F1" bold="true" box="[424,541,618,645]" italics="true" pageId="2" pageNumber="3">Calliaster</emphasis>
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</taxonomicName>
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“group”)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D28B6551FFAAFFAEFF0EFD0A7DA316B3" box="[151,464,687,711]" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="9A2E36DAFFAAFFAEFF0EFD0A7DA316B3" blockId="2.[151,464,687,711]" box="[151,464,687,711]" pageId="2" pageNumber="3">
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<bibRefCitation id="FE004B2BFFAAFFAEFF0EFD0A7D5616B3" author="Mah, C. L." box="[151,293,687,711]" pageId="2" pageNumber="3" pagination="1 - 116" refId="ref19491" refString="Mah, C. L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1 - 116. https: // doi. org / 10.11646 / zootaxa. 4539.1.1" type="journal article" year="2018">Mah 2018: 17</bibRefCitation>
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(full synonymy)
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</paragraph>
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</subSubSection>
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<subSubSection id="D28B6551FFAAFFAEFF0EFCBC7F1C107F" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="9A2E36DAFFAAFFAEFF0EFCBC7D631747" blockId="2.[151,272,793,819]" box="[151,272,793,819]" pageId="2" pageNumber="3">
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<heading id="C16681B6FFAAFFAEFF0EFCBC7D631747" bold="true" box="[151,272,793,819]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="A8E5EAC8FFAAFFAEFF0EFCBC7D631747" bold="true" box="[151,272,793,819]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9A2E36DAFFAAFFAEFF0EFCC77F1C107F" blockId="2.[151,1436,865,1036]" pageId="2" pageNumber="3">A highly variable goniasterid group whose primary characters include the presence of one to several enlarged subambulacral spines present in transverse series, enlarged conical pointed spines present on abactinal, marginal and actinal plates. Abactinal, marginal and actinal plate surfaces bare and smooth. Marginal plates form a prominent abactinal border in most genera but this varies among species (and see taxonomic notes below). Body stellate to strongly stellate with known occurrence only in the Indo-Pacific.</paragraph>
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</subSubSection>
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<subSubSection id="D28B6551FFAAFFAEFF0EFBF87F1A1173" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="9A2E36DAFFAAFFAEFF0EFBF87D6D1003" blockId="2.[151,286,1117,1143]" box="[151,286,1117,1143]" pageId="2" pageNumber="3">
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<heading id="C16681B6FFAAFFAEFF0EFBF87D6D1003" bold="true" box="[151,286,1117,1143]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="A8E5EAC8FFAAFFAEFF0EFBF87D6D1003" bold="true" box="[151,286,1117,1143]" pageId="2" pageNumber="3">Comments</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9A2E36DAFFAAFFAEFF0EFB037F1A1173" blockId="2.[151,1437,1189,1288]" pageId="2" pageNumber="3">
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A full summary of the taxonomic status of
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<taxonomicName id="5D914D59FFAAFFAEFDEBFB007E9110CB" authorityName="Gray" authorityYear="1840" box="[626,738,1189,1215]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="2" pageNumber="3" phylum="Echinodermata" rank="genus">
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<emphasis id="A8E5EAC8FFAAFFAEFDEBFB007E9110CB" box="[626,738,1189,1215]" italics="true" pageId="2" pageNumber="3">Calliaster</emphasis>
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</taxonomicName>
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and related genera was outlined by
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<bibRefCitation id="FE004B2BFFAAFFAEFBE9FB00789D10CB" author="Mah, C. L." box="[1136,1262,1189,1215]" pageId="2" pageNumber="3" pagination="1 - 116" refId="ref19491" refString="Mah, C. L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1 - 116. https: // doi. org / 10.11646 / zootaxa. 4539.1.1" type="journal article" year="2018">Mah (2018</bibRefCitation>
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,
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<bibRefCitation id="FE004B2BFFAAFFAEFB61FB00794610CB" author="Mah, C. L." box="[1272,1333,1189,1215]" pageId="2" pageNumber="3" pagination="1 - 75" refId="ref19657" refString="Mah, C. L. (2022) New Genera, Species and Occurrence of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the Tropical Pacific Ocean by the CAPSTONE Expedition. Zootaxa, 5164 (1), 1 - 75. https: // doi. org / 10.11646 / zootaxa. 5164.1.1" type="journal article" year="2022">2022</bibRefCitation>
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). Species in this genus are variable and boundaries between species concepts is difficult. Unpublished specimen records indicate that there are likely many more species than are known.
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</paragraph>
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</subSubSection>
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</treatment>
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