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<mods:title id="09CBA2974BEED5A4DF8589CD848A22A5">Revision of the genus Tanycypris (Ostracoda, Cypricercinae) with the description of Tanycypris alfonsi n. sp., and an identification key to the genus</mods:title>
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<mods:affiliation id="528D370BCCB3D5FCBF8C81F93B23043E">Aquatic Systems Biology Unit, Department of Ecology and Ecosystem Management, Technische Universitaet Muenchen, Muehlenweg 22, 85350 Freising, Germany. Department of Earth and Environmental Sciences, Palaeontology and Geobiology, GeoBio-Center, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany.</mods:affiliation>
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<mods:affiliation id="A478415B69620DA1A27C5D2918DB5456">Aquatic Systems Biology Unit, Department of Ecology and Ecosystem Management, Technische Universitaet Muenchen, Muehlenweg 22, 85350 Freising, Germany. Department of Earth and Environmental Sciences, Palaeontology and Geobiology, GeoBio-Center, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany.</mods:affiliation>
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<treatment id="1231A341540E2E631DBBCBBEFC5DCB41" ID-DOI="http://doi.org/10.5281/zenodo.5117777" ID-GBIF-Taxon="183717263" ID-Zenodo-Dep="5117777" LSID="urn:lsid:plazi:treatment:1231A341540E2E631DBBCBBEFC5DCB41" httpUri="http://treatment.plazi.org/id/1231A341540E2E631DBBCBBEFC5DCB41" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
<subSubSection id="D28241DC540E2E621DBBCBBEFD07CCC8" box="[151,696,1244,1270]" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="9A271257540E2E621DBBCBBEFD07CCC8" blockId="20.[151,696,1244,1270]" box="[151,696,1244,1270]" pageId="20" pageNumber="421">
<heading id="C16FA53B540E2E621DBBCBBEFD07CCC8" bold="true" box="[151,696,1244,1270]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<taxonomicName id="5D9869D4540E2E621DBBCBBEFD07CCC8" authority="(Krampner, 1928)" baseAuthorityName="Krampner" baseAuthorityYear="1928" box="[151,696,1244,1270]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="telavivensis">
<emphasis id="A8ECCE45540E2E621DBBCBBEFD07CCC8" bold="true" box="[151,696,1244,1270]" pageId="20" pageNumber="421">
<emphasis id="A8ECCE45540E2E621DBBCBBEFE6ECCC8" bold="true" box="[151,465,1244,1270]" italics="true" pageId="20" pageNumber="421">Herpetocypris telavivensis</emphasis>
(Krampner, 1928)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540E2E621DBBCA41FCEACD48" pageId="20" pageNumber="421" type="reference_group">
<paragraph id="9A271257540E2E621DBBCA41FC7ECD06" blockId="20.[151,1002,1314,1398]" box="[151,961,1314,1336]" pageId="20" pageNumber="421">
Nomen nudum 1928a
<emphasis id="A8ECCE45540E2E621CBBCA41FD05CD06" box="[407,698,1314,1336]" italics="true" pageId="20" pageNumber="421">
<taxonomicName id="5D9869D4540E2E621CBBCA41FD1FCD06" baseAuthorityName="Krampner" baseAuthorityYear="1928" box="[407,672,1314,1336]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="telavivensis">Herpetocypris telavivensis</taxonomicName>
</emphasis>
Krampner: 284, 286, 288.
</paragraph>
<paragraph id="9A271257540E2E621DBBCA23FC55CD69" blockId="20.[151,1002,1314,1398]" box="[151,1002,1345,1367]" pageId="20" pageNumber="421">
* 1928b
<taxonomicName id="5D9869D4540E2E621CB5CA20FCA9CD69" authority="Krampner" authorityName="Krampner" authorityYear="1928" box="[409,790,1345,1367]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="telavivensis">
<emphasis id="A8ECCE45540E2E621CB5CA20FD19CD69" box="[409,678,1345,1367]" italics="true" pageId="20" pageNumber="421">Herpetocypris telavivensis</emphasis>
Krampner
</taxonomicName>
: 267270, figs 111.
</paragraph>
<paragraph id="9A271257540E2E621C7FCA02FCEACD48" blockId="20.[151,1002,1314,1398]" box="[339,853,1376,1398]" pageId="20" pageNumber="421">
1992
<taxonomicName id="5D9869D4540E2E621CA0CA03FDC9CD48" baseAuthorityName="Krampner" baseAuthorityYear="1928" box="[396,630,1376,1398]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="telavivensis">
<emphasis id="A8ECCE45540E2E621CA0CA03FDC9CD48" box="[396,630,1376,1398]" italics="true" pageId="20" pageNumber="421">Tanycypris telavivensis</emphasis>
</taxonomicName>
—Martens
<emphasis id="A8ECCE45540E2E621FC5CA03FCA3CD48" box="[745,796,1376,1398]" italics="true" pageId="20" pageNumber="421">et al.</emphasis>
: 114.
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540E2E621DBBCAC0FEC1CEE5" pageId="20" pageNumber="421" type="diagnosis">
<paragraph id="9A271257540E2E621DBBCAC0FC84CE3D" blockId="20.[151,1437,1442,2007]" pageId="20" pageNumber="421">
<emphasis id="A8ECCE45540E2E621DBBCAC0FEAECD85" bold="true" box="[151,273,1442,1467]" inLineHeading="true" pageId="20" pageNumber="421" reason="1">Diagnosis.</emphasis>
L =
<quantity id="5D60BFB2540E2E621C62CAC0FE64CD84" box="[334,475,1442,1467]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="20" pageNumber="421" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
; H =
<quantity id="5D60BFB2540E2E621F0CCAC0FDD9CD84" box="[544,614,1442,1467]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="421" unit="mm" value="1.0">1 mm</quantity>
; W =
<quantity id="5D60BFB2540E2E621F9DCAC0FCA4CD84" box="[689,795,1442,1467]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="421" unit="mm" value="0.75">0.75 mm</quantity>
; Carapace in dorsal view egg-shaped, yellowish-brown, translucent; LV and RV differing in shape, with LV&gt;RV. CR asymmetric, with left trunk thin and curved, and right trunk stout and straight. Males unknown (
<bibRefCitation id="FE096FA6540E2E621F46CA88FC90CE3D" author="Krampner, A." box="[618,815,1514,1539]" pageId="20" pageNumber="421" pagination="281 - 288" refId="ref13486" refString="Krampner, A. (1928 b) Wandlungen der Kruster - und Rotatorien-Fauna eines perennierenden und eines temporalen Tumpels bei Tel Aviv (Palastina) im Verlaufe eines Jahres. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 21, 281 - 288. http: // dx. doi. org / 10.1002 / iroh. 19280190306" type="journal article" year="1928">Krampner 1928b</bibRefCitation>
).
</paragraph>
<paragraph id="9A271257540E2E621DEBC96CFEC1CEE5" blockId="20.[151,1437,1442,2007]" pageId="20" pageNumber="421">
<emphasis id="A8ECCE45540E2E621DEBC96CFE98CE19" bold="true" box="[199,295,1550,1575]" inLineHeading="true" pageId="20" pageNumber="421" reason="1">History.</emphasis>
<bibRefCitation id="FE096FA6540E2E621C18C96DFDB4CE19" author="Krampner, A." box="[308,523,1550,1575]" pageId="20" pageNumber="421" pagination="268 - 270" refId="ref13440" refString="Krampner, A. (1928 a) Eine neue Art der Herpetocypris - syriaca - Gruppe. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 21, 268 - 270. http: // dx. doi. org / 10.1002 / iroh. 19280210207" type="journal article" year="1928">Krampner (1928a)</bibRefCitation>
mentioned this species for the first time in an ecological publication on a temporary freshwater pool near Tel Aviv without any taxonomic description, thus producing a nomen nudum. She acknowledged the help of Vincent Brehm (Lunz,
<collectingCountry id="E28F52C7540E2E621FE9C934FCA3CE51" box="[709,796,1622,1647]" name="Austria" pageId="20" pageNumber="421">Austria</collectingCountry>
), to whom she had sent her material for identification. A short time later, she published the taxonomic description of
<taxonomicName id="5D9869D4540E2E621E10C919FBD8CEAD" authorityName="Krampner" authorityYear="1928" box="[828,1127,1658,1683]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="telavivensis">
<emphasis id="A8ECCE45540E2E621E10C919FBD8CEAD" box="[828,1127,1658,1683]" italics="true" pageId="20" pageNumber="421">Herpetocypris telavivensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="FE096FA6540E2E62198FC919FAD9CEAD" author="Krampner, A." box="[1187,1382,1658,1683]" pageId="20" pageNumber="421" pagination="281 - 288" refId="ref13486" refString="Krampner, A. (1928 b) Wandlungen der Kruster - und Rotatorien-Fauna eines perennierenden und eines temporalen Tumpels bei Tel Aviv (Palastina) im Verlaufe eines Jahres. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 21, 281 - 288. http: // dx. doi. org / 10.1002 / iroh. 19280190306" type="journal article" year="1928">Krampner 1928b</bibRefCitation>
). No further record of this species has been published ever since and
<bibRefCitation id="FE096FA6540E2E621E5DC9FDFBDDCE89" author="Martens, K. &amp; Ortal, R. &amp; Meisch, C." box="[881,1122,1694,1719]" pageId="20" pageNumber="421" pagination="95 - 114" refId="ref13739" refString="Martens, K., Ortal, R. &amp; Meisch, C. (1992) The ostracod fauna of Mamilla Pool (Jerusalem, Israel) (Crustacea, Ostracoda). Zoology in the Middle East, 7, 95 - 114. http: // dx. doi. org / 10.1080 / 09397140.1992.10637628" type="journal article" year="1992">
Martens
<emphasis id="A8ECCE45540E2E621EF9C9FDFBB0CE89" box="[981,1039,1694,1719]" italics="true" pageId="20" pageNumber="421">et al.</emphasis>
(1992)
</bibRefCitation>
transferred it to
<taxonomicName id="5D9869D4540E2E62180EC9FDFA23CE88" authorityName="Triebel" authorityYear="1959" box="[1314,1436,1695,1718]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="20" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540E2E62180EC9FDFA23CE88" box="[1314,1436,1695,1718]" italics="true" pageId="20" pageNumber="421">Tanycypris</emphasis>
</taxonomicName>
without explanation.
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540E2E621DEBC985FB12CF8D" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="9A271257540E2E621DEBC985FB12CF8D" blockId="20.[151,1437,1442,2007]" pageId="20" pageNumber="421">
<materialsCitation id="2AF0180A540E2E621DEBC985FB16CF8D" ID-GBIF-Occurrence="3325982310" location="British Museum of National History" pageId="20" pageNumber="421" specimenCount="1" stateProvince="Tel Aviv">
In the frame of this study, several attempts were made to track down Krampners material. It is not present in the Vulkani-Institute (
<collectingRegion id="585CDCB5540E2E621CBAC868FDBDCF1D" box="[406,514,1802,1827]" country="Israel" name="Tel Aviv" pageId="20" pageNumber="421">Tel Aviv</collectingRegion>
), nor in the University collections of
<collectingRegion id="585CDCB5540E2E621EEBC868FB5ECF1D" box="[967,1249,1802,1827]" isEnumeration="true" pageId="20" pageNumber="421">Tel Aviv and Jerusalem</collectingRegion>
. Our focus was therefore on finding Vincent Brehms collection, to whom Krampner had given her material for identification. According to a short online biography of Brehm (
<bibRefCitation id="FE096FA6540E2E621FFFC830FC34CF55" author="Adamicka, P." box="[723,907,1874,1899]" pageId="20" pageNumber="421" refId="ref12596" refString="Adamicka, P. (2010) Biography of Vincenz Brehm (in German). Available from: http: // de. wikipedia. org / wiki / Vincenz _ Brehm (Accessed 17 Jan. 2014)" type="url" year="2010">Adamicka 2010</bibRefCitation>
) the limnologist Vladimir Kořinek transferred some of Brehms collection to the
<location id="9F47448C540E2E621F09C814FC7BCFB1" LSID="urn:lsid:plazi:treatment:1231A341540E2E631DBBCBBEFC5DCB41:9F47448C540E2E621F09C814FC7BCFB1" box="[549,964,1910,1935]" name="British Museum of National History" pageId="20" pageNumber="421" stateProvince="Tel Aviv">British Museum of National History</location>
, London, in 1988. However, this transfer was not confirmed by the NHM, nor were any of Brehms ostracods found in their collection
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540E2E631DEBC8DCFDFCCABB" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="description">
<paragraph id="9A271257540E2E631DEBC8DCFC8AC8C6" blockId="20.[151,1437,1442,2007]" lastBlockId="21.[151,1437,151,896]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
Until the 1950s, Brehm closely collaborated with Hans Wolfgang Schäfer, limnologist and ostracodologist in Berlin. Schäfer immigrated to
<collectingCountry id="E28F52C7540F2E631CDFCFF5FD3BC88E" box="[499,644,151,176]" name="South Africa" pageId="21" pageNumber="422">South Africa</collectingCountry>
in 1954 and soon continued ostracod research as an employee of the then Department of Agricultural Technical Services. However, Schäfer ceased publishing on ostracods and information on his further career has long been unavailable.
</paragraph>
<paragraph id="9A271257540F2E631DEBCE66FC34C9CB" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="422">
An indication of the whereabouts of the Brehm collection emerged when investigating the archive of Erich Triebels correspondence, housed in the Senckenberg Institute (Frankfurt,
<collectingCountry id="E28F52C7540F2E631EF7CE45FBF7C97E" box="[987,1096,295,320]" name="Germany" pageId="21" pageNumber="422">Germany</collectingCountry>
). Triebel, founder and head of the micropaleontological section from 1939 to 1969, corresponded with researchers worldwide. The correspondence with Schäfer includes a letter sent from Pretoria in 1961, where Schäfer claims to have the largest private ostracod collection, particularly because Vincent Brehm had given him all ostracod material he owned. In the same letter, Schäfer explained that his collection and documents would go to the Transvaal Museum in Pretoria (now Ditsong National Museum of Natural History) after his death.
</paragraph>
<paragraph id="9A271257540F2E631DEBCE9DFDFCCABB" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="422">Attempts to find the Schäfer collection, and thus the Brehm material, in the Ditsong Museum have not (yet) been successful. For the time being, Brehms material cannot be tracked down and thus the material Krampner gave to Brehm must be considered lost. An evaluation of the generic assignment must therefore be carried out based on the original description only.</paragraph>
</subSubSection>
<subSubSection id="D28241DC540F2E631DEBCDEDFC5DCB41" pageId="21" pageNumber="422" type="discussion">
<paragraph id="9A271257540F2E631DEBCDEDFC15CB06" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="422">
<emphasis id="A8ECCE45540F2E631DEBCDEDFEE0CA96" bold="true" box="[199,351,655,680]" inLineHeading="true" pageId="21" pageNumber="422" reason="1">Conclusions.</emphasis>
Following Krampners description and illustrations (1928b), three characters of
<taxonomicName id="5D9869D4540F2E6319D9CDF3FA23CA96" authorityName="Krampner" authorityYear="1928" box="[1269,1436,655,680]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="telavivensis">
<emphasis id="A8ECCE45540F2E6319D9CDF3FA23CA96" box="[1269,1436,655,680]" italics="true" pageId="21" pageNumber="422">H. telavivensis</emphasis>
</taxonomicName>
are clearly different from
<taxonomicName id="5D9869D4540F2E631C90CDD7FD89CAF2" authorityName="Triebel" authorityYear="1959" box="[444,566,693,716]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540F2E631C90CDD7FD89CAF2" box="[444,566,693,716]" italics="true" pageId="21" pageNumber="422">Tanycypris</emphasis>
</taxonomicName>
: in dorsal view, the carapace is egg-shaped (not elliptical, or torpedo-shaped as in
<taxonomicName id="5D9869D4540F2E631DF8CDBAFEF1CAD1" authorityName="Triebel" authorityYear="1959" box="[212,334,728,751]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540F2E631DF8CDBAFEF1CAD1" box="[212,334,728,751]" italics="true" pageId="21" pageNumber="422">Tanycypris</emphasis>
</taxonomicName>
), the valve outline in lateral view is too rectangular for the genus
<emphasis id="A8ECCE45540F2E631910CDBAFB02CAD1" box="[1084,1213,728,751]" italics="true" pageId="21" pageNumber="422">
<taxonomicName id="5D9869D4540F2E631910CDBAFB06CAD1" authorityName="Triebel" authorityYear="1959" box="[1084,1209,728,751]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">Tanycypris</taxonomicName>
,
</emphasis>
and the asymmetry of the caudal rami is not typical of
<taxonomicName id="5D9869D4540F2E631F04CD9FFD1DCB2A" authorityName="Triebel" authorityYear="1959" box="[552,674,765,788]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540F2E631F04CD9FFD1DCB2A" box="[552,674,765,788]" italics="true" pageId="21" pageNumber="422">Tanycypris</emphasis>
</taxonomicName>
, either. In contrast, this asymmetry has been regarded as typical of the Near Eastern species of
<taxonomicName id="5D9869D4540F2E631CE2CC42FDD1CB09" authorityName="Brady &amp; Norman" authorityYear="1889" box="[462,622,800,823]" class="Ostracoda" family="Cyprididae" genus="Herpetocypris" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540F2E631CE2CC42FDD1CB09" box="[462,622,800,823]" italics="true" pageId="21" pageNumber="422">Herpetocypris</emphasis>
</taxonomicName>
by
<bibRefCitation id="FE096FA6540F2E631FB5CC42FC19CB06" author="Krampner, A." box="[665,934,799,824]" pageId="21" pageNumber="422" pagination="281 - 288" refId="ref13486" refString="Krampner, A. (1928 b) Wandlungen der Kruster - und Rotatorien-Fauna eines perennierenden und eines temporalen Tumpels bei Tel Aviv (Palastina) im Verlaufe eines Jahres. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 21, 281 - 288. http: // dx. doi. org / 10.1002 / iroh. 19280190306" type="journal article" year="1928">Krampner (1928b: 270)</bibRefCitation>
.
</paragraph>
<paragraph id="9A271257540F2E631DEBCC26FC5DCB41" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="422">
A final assessment, however, can only be provided if Krampners material reappears in the future. For the time being, we here reject an assignment of this species to the genus
<taxonomicName id="5D9869D4540F2E631E4DCC0AFC64CB41" authorityName="Triebel" authorityYear="1959" box="[865,987,872,895]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="21" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540F2E631E4DCC0AFC64CB41" box="[865,987,872,895]" italics="true" pageId="21" pageNumber="422">Tanycypris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>