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126 lines
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<document id="50FFE8FC8C14D04242C8CAF6D097B1F0" ID-DOI="http://dx.doi.org/10.3897/zookeys.577.7254" ID-GBIF-Dataset="bad453df-1230-42fb-98f6-30fdcaa45f51" ID-PMC="PMC4829886" ID-Pensoft-Pub="1313-2970-577-125" ID-PubMed="27110191" ID-ZooBank="327F242512D544E28CD6E5BC91773077" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-577-125" ModsDocOrigin="ZooKeys 577" ModsDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1459885117108" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2016" docId="11FB29A15557463248AC15A179AD68EC" docLanguage="en" docName="ZooKeys 577: 125-148" docOrigin="ZooKeys 577" docSource="http://dx.doi.org/10.3897/zookeys.577.7254" docTitle="Exocelina hintelmannae Shaverdo, Sagata & Balke 2005" docType="treatment" docVersion="5" lastPageNumber="130" masterDocId="FFECB328490DFFA1BA62B2070F4EC00F" masterDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="148" masterPageNumber="125" pageNumber="128" updateTime="1701316882021" updateUser="plazi">
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<mods:titleInfo id="3DA6DF662B0A58E7461D74B4A999E912">
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<mods:title id="2D21BE52B17BA9E096F7288B5F86674D">Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:name id="E6F66C090B17B430423F6C70027006AD" type="personal">
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<mods:namePart id="0B425BD04EBE3F7AF176BCF65D98B2B9">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="80F47797B41A4F435ACFFC7A343957D7">Sagata, Katayo</mods:namePart>
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<mods:name id="F80B80F03601ECB5579AEDF5F633F1A5" type="personal">
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<mods:roleTerm id="13871A37CA9F90BDF4A5286742ED1759">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9DBA909B825193648C92981384FBC161">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="47FB86EB2D14A6BEA0947F8FA92D3AB8">text</mods:typeOfResource>
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<mods:title id="A3EFEF7C4BF1A57ADEA1D618AA0BC688">ZooKeys</mods:title>
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<mods:part id="8C37BB58540E2220844CB319DEFB4D96">
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<mods:date id="2562377064BE674EAC85535CE2E3066A">2016</mods:date>
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<mods:detail id="F04854EFACF24A11B35E6CAACA08004D" type="volume">
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<mods:number id="15FE30CBF2D59AEAB588372929D672EC">577</mods:number>
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<mods:start id="B40681EE280709CDB2B7EEE0D10E7176">125</mods:start>
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<mods:end id="B6708AC881539A17564BD2B6A265A149">148</mods:end>
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<mods:url id="AFB450267BB1F7690BDE01651CB3892C">http://dx.doi.org/10.3897/zookeys.577.7254</mods:url>
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<mods:classification id="4011AA6125803BD12FEF970374CABBC6">journal article</mods:classification>
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<mods:identifier id="AE35A9F029D8768494FDCB9ECC07F03A" type="DOI">http://dx.doi.org/10.3897/zookeys.577.7254</mods:identifier>
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<mods:identifier id="DDC7A984FC648D8BD02810CFB561C902" type="Pensoft-Pub">1313-2970-577-125</mods:identifier>
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<treatment id="11FB29A15557463248AC15A179AD68EC" ID-GBIF-Taxon="127880284" LSID="urn:lsid:plazi:treatment:11FB29A15557463248AC15A179AD68EC" httpUri="http://treatment.plazi.org/id/11FB29A15557463248AC15A179AD68EC" lastPageId="5" lastPageNumber="130" pageId="3" pageNumber="128">
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<subSubSection id="C9F36D1229DE488F05C8AD459FD98F6A" pageId="3" pageNumber="128" type="multiple">
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<paragraph id="1D5D2C913BFCC7A92AC37272F1B4DA41" pageId="3" pageNumber="128">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="CA21F7C53F3E4F8D6ACEEDFC0263B510" pageId="3" pageNumber="128" type="nomenclature">
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<paragraph id="9FEEDF84018AB32E416A56B7ED802CE8" pageId="3" pageNumber="128">
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2.
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<taxonomicName id="E146DFA31B786FBF15CE8194465EA4EA" ID-CoL="3DLQ3" authority="Shaverdo, Sagata & Balke, 2005" authorityName="Shaverdo, Sagata & Balke" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hintelmannae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="hintelmannae">Exocelina hintelmannae (Shaverdo, Sagata & Balke, 2005)</taxonomicName>
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Figs 4, 9F, G
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</paragraph>
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</subSubSection>
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<subSubSection id="2B2F076B9FCD96AB85E92E66DD362452" pageId="4" pageNumber="129" type="reference_group">
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<paragraph id="6054EB05355098B24766D9F605B285AC" pageId="4" pageNumber="129">
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<taxonomicName id="6062BB4261E5354BB43591CA21BD7FAA" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes hintelmannae" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="hintelmannae">
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<pageBreakToken id="EF956316E137AEAD48AA6180938D0809" pageId="4" pageNumber="129" start="start">Papuadytes</pageBreakToken>
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hintelmannae
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</taxonomicName>
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Shaverdo, Sagata & Balke, 2005: 272.
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</paragraph>
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<paragraph id="BFB11BB68D33CD7435CC6541D4C4F0A2" pageId="4" pageNumber="129">
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<taxonomicName id="041B2B6E3FF1C15C7A1616A5FA8C7481" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hintelmannae" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="hintelmannae">Exocelina hintelmannae</taxonomicName>
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(Shaverdo, Sagata & Balke, 2005):
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<bibRefCitation id="DD10A48D1D05A037810646D6B91A4B12" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="13" pageNumber="138" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
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: 33 (comb. n.);
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<bibRefCitation id="CADE04C6924CF4087522A04A4A2FEC10" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="13" pageNumber="138" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, tab. 2. (MB 1367).
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</paragraph>
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</subSubSection>
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<subSubSection id="0BF22F184BC3E093A9CCBBA1535AF17B" pageId="4" pageNumber="129" type="type locality">
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<paragraph id="C5F7C1A0B767382C027C989223BE4E5C" pageId="4" pageNumber="129">Type locality.</paragraph>
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<paragraph id="08B301E96DED7B268CCE5BBC86B6EBD4" pageId="4" pageNumber="129">
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Papua New Guinea: border Simbu and Eastern Highlands Provinces, Crater Mountain, between Wara Sera Station and Herowana Village, Hulene River, approximately
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<geoCoordinate id="4BFDCCE2A745C19B62231D3B096EE70E" direction="south" orientation="latitude" precision="92" value="-6.7233334">06°43.4'S</geoCoordinate>
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;
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<geoCoordinate id="75D15979BD1C4725A11F155941D52C3F" direction="east" orientation="longitude" precision="92" value="145.09334">145°05.6'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="2B230301F1D505EB66D252603A90DB39" pageId="4" pageNumber="129" type="type material">
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<paragraph id="0E0742CE1AAAB997AD93BD19C354DA5F" pageId="4" pageNumber="129">Type material studied.</paragraph>
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<paragraph id="7CE9E0C4C6E797226C0F522B48667DBA" pageId="4" pageNumber="129">
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<typeStatus id="5BE3B7A2D82AE843CDA76F1DFE2A0EE8" pageId="4" pageNumber="129">Holotype</typeStatus>
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: male "264 DNA M Balke" [green], "PNG Simbu / EHPr. Crater Mountain, Sera - Herowana, Wara Hulene, 1000 m, 16IX2002, Balke & Sagata (PNG 17)", "Holotypus
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<taxonomicName id="57DB88E047A580E017CB3F47EB159124" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes hintelmannae" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="hintelmannae">Papuadytes hintelmannae</taxonomicName>
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sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (BMNH).
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<typeStatus id="597A681C7E1EBF81B90D9C6FFFB0723A" pageId="4" pageNumber="129">Paratypes</typeStatus>
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: 1 male "256 DNA M Balke" [green], "Papua New Guinea Simbu / EHPr. Crater Mountain, Wara Sera Station, 800 m, 14IX2002, Balke & Sagata (PNG 10)", "Paratypus
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<taxonomicName id="4AA69FD06C6C91A4E3BE71E9DF8D0C57" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes hintelmannae" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="hintelmannae">Papuadytes hintelmannae</taxonomicName>
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sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (NHMW). 1 male "260 DNA M Balke" [green], "PNG Simbu / EHPr. Crater Mountain, Sera - Herowana, upper Oh River, 1200 m, 15IX2002, Balke & Sagata (PNG 12)", "Paratypus
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<taxonomicName id="B4DDA7887B746D4F2930E701AC1995F7" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes hintelmannae" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="hintelmannae">Papuadytes hintelmannae</taxonomicName>
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sp.n. des. H. Shaverdo, K. Sagata & M. Balke, 2005" (NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection id="47C10B4DC4CDCE08642E923294AEFA27" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129" type="materials_examined">
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<paragraph id="8CDD05AF05DC2572FBF2D8959878A2B6" pageId="4" pageNumber="129">Additional material.</paragraph>
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<paragraph id="5F025C8339C4B6BF7888EA9BFEF521BC" lastPageId="5" lastPageNumber="130" pageId="4" pageNumber="129">
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Simbu/EHL: 1 female "Papua New Guinea: Simbu / EHPr. Crater Mountain, Wara Sera Station, 820m, 14IX2002, Balke & Sagata, (PNG 8)" (ZSM). 2 males, 6 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 009)" (ZSM). 17 males, 15 females "Papua New Guinea: Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 010)" (NARI, NHMW, ZSM). 8 males, 9 females "Papua New Guinea Simbu/EHP. Crater Mountain, Wara Sera Station, 800m, 14IX2002, Balke & Sagata, (PNG 10)" (NHMW, ZSM). 3 males, 4 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Sera - Herowana, Wara Pima, 900m, 15IX2002, Balke & Sagata (PNG 011)" (ZSM). 7 males, 14 females "Papua New Guinea: Crater Mountain, Sera - Herowana, upper Oh river, 1200m, 15IX2002, Balke & Sagata (PNG 012)" (NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1100m, 15IX2002, Balke & Sagata (PNG 013)" (ZSM). 8 males, 7 females "Papua New Guinea: Crater Mountain, Sera - Herowana, Jau river, 1000m, 15IX2002, Balke & Sagata (PNG 015)" (NHMW, ZSM). 12 males, 10 females "Papua New Guinea: Simbu/EHP. Crater Mountain, Sera - Herowana, Hulene river, 1000m, 16IX2002, Balke & Sagata (PNG 017)" (NHMW, ZSM). 25 males, 33 females "Papua New Guinea: Simbu / EHP. Crater Mountain, Herowana, Yawasa River, 1200m, 17.IX.2003, Balke & Sagata (PNG 019)" (NARI, NHMW, ZSM). Simbu: 8 males, 13 females "Papua New Guinea: Supa Haia, 1032, 10.xi.2002, K. Sagata, (WB1)" (NHMW, ZSM). EHL: 1 male "Stn. No. 177a", "NEW GUINEA: E. Highland Dist., Wanatabe Valley. Nr. Okapa, c. 5000 ft. 5.ii.1965", "M.E. Bacchus. B.M. 195-120" (BMNH). Gulf: 30 males, 30 females "Papua New Guinea: Gulf, Marawaka, nr. Ande, 1000m, 10.xi.2006, 07.03.598S 145.4.375E, Balke & Kinibel (PNG 89)", one of them an additional green label "DNA M.Balke 1367" (NHMW, ZSM). 32 males, 35 females "Papua New Guinea: Gulf Province, Marawaka, Mala, 1400m, 11.xi.2006, 07.05.664S 145.44.467E, Balke & Kinibel, (PNG 90)" (NARI, NHMW, ZSM). 16 male, 12 females "Papua New Guinea: Gulf, Marawaka, Andakombe towards Morobe, 1100m,
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<pageBreakToken id="D0377581B047BE4EFB05E040A988E054" pageId="5" pageNumber="130" start="start">12</pageBreakToken>
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.xi.200, 07.09.766S 145.46.333E, Balke & Kinibel (PNG 92)" (NHMW, ZSM). 1 male, 3 females "Papua New Guinea: Gulf, Marawaka, Andakombe towards Morobe, 1500m, 12.xi.200, 07.10.413S 145.49.555E, Balke & Kinibel (PNG 93)", the male with an additional green label "DNA M.Balke 1362" (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection id="E79D7986CCD7AEA4047797132FDAF1FB" pageId="5" pageNumber="130" type="additions to the description">
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<paragraph id="68CE8A5E0A2082262F2FF7AD5AE86F38" pageId="5" pageNumber="130">Additions to the description</paragraph>
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<paragraph id="938D2826912E0DEA4A777251739CD097" pageId="5" pageNumber="130">
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(original description in
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<bibRefCitation id="36A9D25D3114F84350D51A90915262BF" author="Shaverdo, HV" journalOrPublisher="Aquatic Insects" pageId="13" pageNumber="138" pagination="269 - 280" title="Five new species of the genus Papuadytes Balke, 1998 from New Guinea (Coleoptera: Dytiscidae)." url="10.1080/01650420500290169" volume="27" year="2005">Shaverdo et al. 2005</bibRefCitation>
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, p. 272). Size and shape: Beetles medium-sized (TL-H 3.4-4.15 mm, TL 3.75-4.55 mm, MW 1.8-2.2 mm;
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<typeStatus id="76AD078CF268922D9FA78F59DCFC8971" pageId="5" pageNumber="130">holotype</typeStatus>
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: TL-H 3.9 mm, TL 4.3 mm, MW 2.05 mm), see Fig. 4.
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</paragraph>
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<paragraph id="A6F0F54D7E93405B73838858465A418D" pageId="5" pageNumber="130">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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<paragraph id="C87B7440362D7B7270E2B9CC36C96CC6" pageId="5" pageNumber="130">Variability: Beetles vary in shape of the median lobe and its ventral sclerite: mainly shape and length of its left lobe (Fig. 9F, G).</paragraph>
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</subSubSection>
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<subSubSection id="CBC334FDE3799C7CF62493CACE9FBC13" pageId="5" pageNumber="130" type="distribution">
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<paragraph id="F23442F535EAEA9233E4A1FD7A214DCE" pageId="5" pageNumber="130">Distribution.</paragraph>
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<paragraph id="EC94424273C59EAF48AA90D0EB0A5487" pageId="5" pageNumber="130">Papua New Guinea: Simbu, Eastern Highlands, and Gulf Provinces (Fig. 14).</paragraph>
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</subSubSection>
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</treatment>
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</document> |