162 lines
15 KiB
XML
162 lines
15 KiB
XML
<document id="69F6CF1B0ACFAB88ABC2CB88D694352A" ENCODING="UTF-8" ID-CLB-Dataset="52661" ID-GBIF-Dataset="2cc6a882-3bfb-4232-a6c8-c46d4f1cf25f" ModsDocID="z01213p001" checkinTime="1247155641890" checkinUser="thomas" docAuthor="Wilson J. E. M. Costa" docDate="2006" docId="117C9DAF8504DFAF7376A3038C51EFA8" docLanguage="en" docName="2006_Costa_gg1.xml" docOrigin="Zootaxa 1213" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:3415A121-707B-4676-9259-4FD5CE1C3323" docTitle="Austrolebias varzeae" docType="treatment" docVersion="13" lastPageNumber="95" masterDocId="7116976AA5DF5C9AF7BCD64CDBAACDB0" masterDocTitle="The South American annual killifish genus Austrolebias (Teleostei: Cyprinodontiformes: Rivulidae): phylogenetic relationships, descriptive morphology and taxonomic revision." masterLastPageNumber="162" masterPageNumber="1" pageNumber="93" updateTime="1694017982350" updateUser="ExternalLinkService">
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<mods:titleInfo id="D43BD63AAB916BEF14979489082EFD93">
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<mods:title id="AB0E235BFE95529670F6752BAB07E2BC">The South American annual killifish genus Austrolebias (Teleostei: Cyprinodontiformes: Rivulidae): phylogenetic relationships, descriptive morphology and taxonomic revision.</mods:title>
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<mods:name id="29A176A22717883B17135F81B9394DD5" type="personal">
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<mods:roleTerm id="7D54FF453A0D6D17B0349E7312D113A4">Author</mods:roleTerm>
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<mods:namePart id="123B4E0A24ECCD01050DB3286DAF88FB">Wilson J. E. M. Costa</mods:namePart>
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<mods:typeOfResource id="20DDBDFC7BF6FFEB92986C5449E63921">text</mods:typeOfResource>
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<mods:title id="7D49239504EC903DC2FCE5D8081FF279">Zootaxa</mods:title>
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<mods:part id="749D9B4314B0BD30285A8DCAB2E1EAB2">
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<mods:date id="971F6B7B9C8846285EBEBE26A15F7909">2006</mods:date>
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<mods:number id="81EC99CDA9951BCD02B7BAC4E67D8791">1213</mods:number>
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<mods:classification id="78BC0A8EB52E015DD4F27C40049BA9A8">journal article</mods:classification>
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<mods:identifier id="63F11859D9A0A9D333A245A32E500989" type="ZooBank-PUB">z01213p001</mods:identifier>
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<treatment id="117C9DAF8504DFAF7376A3038C51EFA8" ID-DOI="http://doi.org/10.5281/zenodo.6263742" ID-GBIF-Taxon="100119122" ID-Zenodo-Dep="6263742" LSID="urn:lsid:plazi:treatment:117C9DAF8504DFAF7376A3038C51EFA8" httpUri="http://treatment.plazi.org/id/117C9DAF8504DFAF7376A3038C51EFA8" lastPageNumber="95" pageNumber="93">
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<subSubSection id="5D890ECA632E09906890D5C08E64852B" type="nomenclature">
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<paragraph id="E46B5C5BD18EA12AD339A590805ADBAB" pageNumber="93">
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<taxonomicName id="4E2810A8B0AC9A85B788CB36DC9938A5" ID-CoL="JWWW" ID-ENA="2491938" LSID-ZBK="urn:lsid:zoobank.org:act:CC046AF7-14F6-448A-82E8-F3978CFBCB36" family="Rivulidae" genus="Austrolebias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Austrolebias varzeae Costa, Reis & Behr 2004:14" order="Cyprinodontiformes" pageNumber="93" phylum="Chordata" rank="species" species="varzeae">
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<pageStartToken id="065DDE99AF7633FD27A48AAB2D5C65C9" pageNumber="93">Austrolebias</pageStartToken>
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varzeae
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</taxonomicName>
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Costa, Reis & Behr
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</paragraph>
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</subSubSection>
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<subSubSection id="E9E5717A20E5E52627F2F234BEFE724A" type="description">
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<paragraph id="EFC7F3A3BD1F0ED0330148DBAA38B5DF" pageNumber="93">(Fig. 36)</paragraph>
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</subSubSection>
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<subSubSection id="2661CD013723FD38D63D4D493F75C65B" type="reference_group">
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<paragraph id="9DE65A88CFF50D552066260B1C1B50F1" pageNumber="93">
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<taxonomicName id="190F9F835051064EDAC9243672E5081C" LSID-ZBK="urn:lsid:zoobank.org:act:CC046AF7-14F6-448A-82E8-F3978CFBCB36" family="Rivulidae" genus="Austrolebias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Austrolebias varzeae Costa, Reis & Behr 2004:14" order="Cyprinodontiformes" pageNumber="93" phylum="Chordata" rank="species" species="varzeae">Austrolebias varzeae</taxonomicName>
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Costa, Reis & Behr, 2004: 14 (
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<materialsCitation id="98EED429CE277C0BA36836C36DAF2E4B" ID-GBIF-Occurrence="923891658" collectionCode="MCP" country="Brazil" latitude="-28.3" location="type locality: temporary lagoon close to rio da Varzea, Fazenda dos Branda, Carazinho, rio Uruguay basin" longitude="-52.8" specimenCode="MCP 29641" stateProvince="Rio Grande do Sul" typeStatus="holotype">
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<location id="721B511A15BE68D1878BF1A73B362AF1" LSID="urn:lsid:plazi:treatment:117C9DAF8504DFAF7376A3038C51EFA8:721B511A15BE68D1878BF1A73B362AF1" country="Brazil" latitude="-28.3" longitude="-52.8" name="type locality: temporary lagoon close to rio da Varzea, Fazenda dos Branda, Carazinho, rio Uruguay basin" stateProvince="Rio Grande do Sul">
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type locality: temporary lagoon close to rio da
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<normalizedToken id="9D1931CE11DFF80EC642B96F50331727" originalValue="Várzea">Varzea</normalizedToken>
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, Fazenda dos Branda, Carazinho, rio Uruguay basin
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</location>
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,
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<geoCoordinate id="E3C09A2E7D52BB94A7F5FF65A39E803C" direction="south" orientation="latitude" value="-28.3">28º18’ S</geoCoordinate>
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<geoCoordinate id="BAE33EEEA7DFB20D574E4C847C945DC9" direction="west" orientation="longitude" value="-52.8">52º 48’W</geoCoordinate>
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,
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<collectingRegion id="4E8E9180E4F9FA1571423C199FCC457E">Rio Grande do Sul</collectingRegion>
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,
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<collectingCountry id="006246025A3E0C04415BCC3C5CC9EA25">Brazil</collectingCountry>
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;
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<typeStatus id="4FF3EC54A7CD21BB111AFD59423C1B1C">holotype</typeStatus>
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:
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<specimenCode id="B9E6A53DD0B73B4BB65DFD7427D54E36">
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<collectionCode id="E1A06E68CF1D560835FDA8FC62AE88F1">MCP</collectionCode>
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29641
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</specimenCode>
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</materialsCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3A09E1EF00EE300654F94259E36CCC5A" type="materials_examined">
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<paragraph id="A42CE016BFAF150A99F08665165B6AD0" pageNumber="93">Material examined</paragraph>
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<paragraph id="77E35EFF53FCFB5367470E09FB9354EA" pageNumber="93">
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<materialsCitation id="DBD4B3FE7081E245BAFED83FA1A798BA" ID-GBIF-Occurrence="923891640" collectingDate="1999-08-26" collectionCode="MCP, UFRJ" collectorName="M. N. Xavier" country="Brazil" latitude="-28.3" location="temporary lagoon close to rio da Varzea, Fazenda dos Branda, Carazinho, rio Uruguay basin" longitude="-52.8" specimenCode="MCP 29641, MCP 23667, UFRJ 5431, UFRJ 5432" stateProvince="Rio Grande do Sul" typeStatus="holotype, paratypes">
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<collectingCountry id="95F1CD3C869E9CA9F6C0AD2546042885">Brazil</collectingCountry>
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:
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<collectingRegion id="2374445860F84082B478AC2A6EE98E56">Rio Grande do Sul</collectingRegion>
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:
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<specimenCode id="62CA782B6E729918BB825ABC5FE26A36">
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<collectionCode id="C18797CBD29BEB1226B119B57FDA1E48">MCP</collectionCode>
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29641
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</specimenCode>
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, male
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<typeStatus id="C685201F02790ED4D670E15EF97A4B0A">holotype</typeStatus>
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, 33.7 mm SL;
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<specimenCode id="A8135AB1528F42C3B007633D632CD248">
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<collectionCode id="4C0F9C30895F23C2EC4C838DADE17B6B">MCP</collectionCode>
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23667
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</specimenCode>
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, 9
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<typeStatus id="F53A9E5736E1CEB51F7AA0A654D96CCE">paratypes</typeStatus>
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;
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<specimenCode id="7157EDC976E09B0F3CEA6563E8C131DD">
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<collectionCode id="E4FA0CCD6D08BCB2A8B76DE5F87F2F5F">UFRJ</collectionCode>
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5431
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</specimenCode>
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, 2
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<typeStatus id="035CF0AEAAD67B33D72C83F71F02B93A">paratypes</typeStatus>
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;
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<specimenCode id="C99ED1ABFB8F019723D05EC67B98F973">
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<collectionCode id="588101371E8E5CD65CD7514F9FEC7CF9">UFRJ</collectionCode>
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5432
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</specimenCode>
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, 2
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<typeStatus id="EC5702DC48F6BC77C26D401467E27F47">paratypes</typeStatus>
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(c&s);
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<location id="67FE49AA80BD4C326C3601F940AD8F2F" LSID="urn:lsid:plazi:treatment:117C9DAF8504DFAF7376A3038C51EFA8:67FE49AA80BD4C326C3601F940AD8F2F" country="Brazil" latitude="-28.3" longitude="-52.8" name="temporary lagoon close to rio da Varzea, Fazenda dos Branda, Carazinho, rio Uruguay basin" stateProvince="Rio Grande do Sul">
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temporary lagoon close to rio da
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<normalizedToken id="40AEF5BB089641ABD44CB057DCD50069" originalValue="Várzea">Varzea</normalizedToken>
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, Fazenda dos Branda, Carazinho, rio Uruguay basin
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</location>
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;
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<collectorName id="4850B3092E0B0446E8C8C72A39C20212">M. N. Xavier</collectorName>
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,
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<date id="0D2F51188B2A07F32148C5AE7FF6EE18" day="26" month="08" value="1999-08-26" year="1999">
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<collectingDate id="3235E3299E3D13826ABE647742C16C9F" day="26" month="08" value="1999-08-26" year="1999">26 Aug. 1999</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9C7C41596AF648B8A451F8FBF27E652E" type="diagnosis">
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<paragraph id="144287768E9515694740479D0CF5D79A" pageNumber="93">Diagnosis</paragraph>
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<paragraph id="82D05AB2FEC0AF8F6CF3C9A4B9207F94" pageNumber="93">Distinguished from all other congeners by the combination of the following features: dark gray bars on flank in males, the three anteriormost bars distinctively darker and narrower, no contact organs on male anal fin, pelvic-fin bases separated by small interspace, anal fin short in females, urogenital papilla not attached to anal fin, dorsal-fin origin anterior to anal-fin origin, dorsal-fin rays 22-24 in males, 16-18 in females, anal-fin rays 23-26 in males, 18-19 in females, and longitudinal series scales 28-30.</paragraph>
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</subSubSection>
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<subSubSection id="22F1820207EF8D906458AF60A8D8DD37" type="description">
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||
<paragraph id="9DC9A3F1419A9E47BF513D9868690C1A" pageNumber="93">Description</paragraph>
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<paragraph id="B27B76B9819BC203F4589C79AA10B648" pageNumber="93">Morphometric data appear in Table 6. Males larger than females, largest male examined 33.7 mm SL, largest female 25.8 mm SL. Dorsal profile slightly concave on head, convex from nape to end of dorsal-fin base, approximately straight on caudal peduncle; no distinctive adipose ridge on frontal region. Ventral profile convex from lower jaw to end of anal-fin base, nearly straight on caudal peduncle. Greatest body depth at level of pelvic-fin base. Body deep and moderately compressed. Snout blunt and jaws short.</paragraph>
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<paragraph id="5013764CA20FC3AAF6EE84EFB07EE7D0" pageNumber="93">Tip of both dorsal and anal fins rounded. Anteromedian rays of anal fin of females not lengthened; distal portion of anal fin thickened in females. Caudal fin rounded. Pectoral fins elliptical, short, posterior margin on vertical between pelvic-fin base and urogenital papilla. Tip of each pelvic fin reaching base of 2nd anal-fin ray. Pelvic-fin bases in close proximity. Urogenital papilla not attached to anal fin. Anal-fin origin on vertical through base of 2nd or 3rd dorsal-fin ray; dorsal-fin origin between neural spines of 8th and 9th vertebrae in males, between neural spines of 10th and 11th vertebrae in females. Anal-fin origin between pleural ribs of 9th and 10th vertebrae in males, between pleural ribs of 10th and 11th vertebrae in females. Dorsal-fin rays 22-24 in males, 16-18 in females; anal-fin rays 23-26 in males, 18-19 in females; caudal-fin rays 28-29; pectoral-fin rays 12; pelvicfin rays 5.</paragraph>
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<paragraph id="3AAD6139C2B82302FFC72D59A15F3405" pageNumber="94">
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<pageStartToken id="94EDFD72EEEB411BFDBD16A8C8E6DD58" pageNumber="94">Scales</pageStartToken>
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large and cycloid. Trunk and head entirely scaled, except ventral surface of head. Sometimes few scales extending over anal-fin base; no scales on dorsal-fin base; three rows of scales on caudal-fin base. Frontal squamation F-patterned; E-scales not overlapping medially; scales arranged in transverse pattern. Longitudinal series of scales 28-30, scales regularly arranged; transverse series of scales 16-17; scale rows around caudal peduncle 20. One contact organ on each scale of ventral portion of flanks in males. Row of contact organs on three uppermost pectoral-fin rays in males. No contact organ on unpaired fins.
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</paragraph>
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<paragraph id="E6ADE60550FBBFE468869AE3175FF0DC" pageNumber="94">Cephalic neuromasts: supraorbital 17-18, parietal 1-2, anterior rostral 1, posterior rostral 1, infraorbital 3 + 20, preorbital 2, otic 3, post-otic 4-5, supratemporal 1, median opercular 1, ventral opercular 2, preopercular 21-23, mandibular 15-16, lateral mandibular 4.</paragraph>
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<paragraph id="79FB62A75C2C25DA6E315E0761BE866B" pageNumber="94">Basihyal subtriangular, width about 80 % of length; basihyal cartilage long, about 65 % of total basihyal length, with pronounced lateral projection. Six branchiostegal rays. Three to five teeth on second pharyngobranchial. Gill-rakers on first branchial arch 3 + 9. Dermosphenotic ossification absent. Ventral process of posttemporal long. Total vertebrae 27-28.</paragraph>
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<caption id="27508A2B962516E3B46102542E3B9670">
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<paragraph id="EA1730E5E8F1B3482652C0F915EA1657" pageNumber="94">
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FIGURE 36.
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<taxonomicName id="D66C63DFA673EDD6889FC824045E4C9D" LSID-ZBK="urn:lsid:zoobank.org:act:CC046AF7-14F6-448A-82E8-F3978CFBCB36" family="Rivulidae" genus="Austrolebias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Austrolebias varzeae Costa, Reis & Behr 2004:14" order="Cyprinodontiformes" pageNumber="94" phylum="Chordata" rank="species" species="varzeae">Austrolebias varzeae</taxonomicName>
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, MCP 29641, male holotype, 33.7 mm SL; Brazil: Rio Grande do Sul: Carazinho.
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</paragraph>
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</caption>
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<paragraph id="49267204A670EA24CBCD7F78DE4A8D86" pageNumber="94">Coloration</paragraph>
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<paragraph id="6B26C423280FB3F7569813E55828D179" pageNumber="94">Males: sides of body bluish gray, with 9-11 dark gray bars, anterior three bars darker and narrower than interspace, remaining bars faint and wider than interspace. Urogenital papilla dark gray. Opercular and infraorbital bluish gray; nearly rectangular dark gray infraorbital bar, subtriangular dark gray supraorbital bar. Iris orangish brown, with dark brown bar through center of eye. Unpaired and pelvic fins dark bluish gray; white dots on entire dorsal fin, basal portion of anal fin, and dorsal portion of caudal fin; black distal line on anal fin. Pectoral fins hyaline.</paragraph>
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<paragraph id="60E604D95FE5E0D2C106EBE905E7D5FC" pageNumber="94">Females: sides of body light yellowish brown, with round to slightly vertically elongated dark gray spots, darker on anterocentral portion of flank; no distinctive spot on caudal peduncle. Opercular region pale greenish golden. Iris orangish brown, with gray bar through center of eye. Infraorbital and supraorbital bars dark gray. Unpaired fins hyaline, with small gray spots on basal portion of anal fin; paired fins hyaline.</paragraph>
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</subSubSection>
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<subSubSection id="BF4B5964DB84ECA3B941E52AEC9BBA58" type="distribution">
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<paragraph id="1EC3AC54238862990B2B7C811F2B48DB" pageNumber="95">
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<pageStartToken id="E59C4B214530231D03C441622F105DC9" pageNumber="95">Distribution</pageStartToken>
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</paragraph>
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<paragraph id="A2A857A92C7857EAA94635A87A0C1CE4" pageNumber="95">
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Known only from rio da
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<normalizedToken id="9122CE8AF7B5E7518110FAC39DEBE6A0" originalValue="Várzea">Varzea</normalizedToken>
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drainage, upper rio Uruguay basin, southern Brazil (Fig. 8).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |