420 lines
46 KiB
XML
420 lines
46 KiB
XML
<document id="6F09E64B0CFA3EFF1341E3BF0692EE8E" ID-CLB-Dataset="32684" ID-DOI="10.11646/zootaxa.4291.2.5" ID-GBIF-Dataset="5b969e11-336f-49e4-a134-ba402d653c6f" ID-ISSN="1175-5326" ID-Zenodo-Dep="829627" ID-ZooBank="6B6DE509-9F1B-4127-93F4-441C8C2F7B02" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500111838680" checkinUser="plazi" docAuthor="Xue, Jing, Li, Xiaochen, Wang, Lizhi, Xian, Panpan & Chen, Huiqing" docDate="2017" docId="110F87DC366F4C2DFAB3FD246AA6FD95" docLanguage="en" docName="zootaxa.4291.2.5.pdf" docOrigin="Zootaxa 4291 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bryochoerus liupanensis Xue, Li, Wang, Xian & Chen, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="327" masterDocId="ED36FFA4366E4C2EFA24FF8E6E0EFF82" masterDocTitle="Bryochoerus liupanensis sp. nov. and Pseudechiniscus chengi. sp. nov. (Tardigrada: Heterotardigrada: Echiniscidae) from China" masterLastPageNumber="334" masterPageNumber="324" pageNumber="325" updateTime="1698465578722" updateUser="ExternalLinkService">
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<mods:mods id="1B4F82A7755237669CAF14E2990B39DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B96AF590BEE3DA961B6140A42386BFD3">
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<mods:title id="90AC069DC80B268650C959D99047D25E">Bryochoerus liupanensis sp. nov. and Pseudechiniscus chengi. sp. nov. (Tardigrada: Heterotardigrada: Echiniscidae) from China</mods:title>
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</mods:titleInfo>
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<mods:name id="8FE2F97F2C5F415402EF32E984149ADA" type="personal">
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<mods:role id="B5E5E81DAF2AD64E660279979B6ABF3C">
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<mods:roleTerm id="9A138AFCB40F715D1930E0333295178D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92797A4E1BDDB15C35D874CA588F3F5F">Xue, Jing</mods:namePart>
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</mods:name>
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<mods:name id="5B122B31435968E8D9AED8808492BA95" type="personal">
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<mods:roleTerm id="3FCD30D3F4F1DE97AB88CD175C4BA0D0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="225D9C6C40BCB92471117F27A9AB8C24">Li, Xiaochen</mods:namePart>
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</mods:name>
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<mods:name id="1EFA091A92739A680E42C6AEAFE67D0B" type="personal">
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<mods:roleTerm id="C1FFE76CF09E3E88DBDC8194717FAEC9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E76F9A9B56A65B39D883605AD2987DD">Wang, Lizhi</mods:namePart>
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</mods:name>
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<mods:name id="BFE4F8C117381CF1792854BDD915DF23" type="personal">
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<mods:roleTerm id="33426FB26596F6E00D6748D35169D0B7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0685E9E59ADD8AA97952312481C9D792">Xian, Panpan</mods:namePart>
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</mods:name>
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<mods:name id="1440FE4624C2C31D1A2B443288B34FB1" type="personal">
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<mods:role id="513379B0B8D5892B183BC40FC30383B1">
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<mods:roleTerm id="3F8B926C384BD8150C7970D366446EE2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2E5DB6BB1905DA08CD1AA816EA5C785C">Chen, Huiqing</mods:namePart>
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<mods:typeOfResource id="73586A2DDEB816D3A037413AE6077EF1">text</mods:typeOfResource>
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<mods:title id="9159CF074C8A6BD284B2146E15A7E6A0">Zootaxa</mods:title>
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<mods:part id="84FF34275D577A2C422CE014D92C7303">
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<mods:date id="45EA117C082AABDE37E3853D1ED981AD">2017</mods:date>
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<mods:detail id="14171877AC65A7E304E27F97C9834F2A" type="volume">
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<mods:number id="66CC87723D921A9F78A230E6FF476A46">4291</mods:number>
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</mods:detail>
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<mods:detail id="D24682DCFA6FBD4568AF9778E636615F" type="issue">
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<mods:number id="61ECD740ED27681388BA632ECC0459E9">2</mods:number>
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<mods:classification id="4D4183AFC1D3EB4CDF98EF0C39F30F4D">journal article</mods:classification>
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<mods:identifier id="D53E452F9572876952B041E310358EDE" type="CLB-Dataset">32684</mods:identifier>
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<mods:identifier id="B3E7068E19B0053CA7BC30DF706A9759" type="DOI">10.11646/zootaxa.4291.2.5</mods:identifier>
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<mods:identifier id="55549DAD2C3AFF6590B560A19BDD6B88" type="GBIF-Dataset">5b969e11-336f-49e4-a134-ba402d653c6f</mods:identifier>
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<mods:identifier id="662CD09FA0AA0892BB43D41BAB44CDDE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B3DF679A24B5396E3AEAEE130AA4C560" type="Zenodo-Dep">829627</mods:identifier>
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<mods:identifier id="0C106E6CD94EAAA0F7B4DE18B5467E3D" type="ZooBank">6B6DE509-9F1B-4127-93F4-441C8C2F7B02</mods:identifier>
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</mods:mods>
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<treatment id="110F87DC366F4C2DFAB3FD246AA6FD95" ID-DOI="http://doi.org/10.5281/zenodo.6022530" ID-GBIF-Taxon="132549712" ID-Zenodo-Dep="6022530" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95" httpUri="http://treatment.plazi.org/id/110F87DC366F4C2DFAB3FD246AA6FD95" lastPageId="3" lastPageNumber="327" pageId="1" pageNumber="325">
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<subSubSection id="D1BC6541366F4C2FFAB3FD246F7CFD64" pageId="1" pageNumber="325" type="nomenclature">
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<paragraph id="991936CA366F4C2FFAB3FD246C26FD46" blockId="1.[151,552,682,742]" box="[151,552,682,708]" pageId="1" pageNumber="325">
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<heading id="C25181A6366F4C2FFAB3FD246C26FD46" bold="true" box="[151,552,682,708]" fontSize="11" level="1" pageId="1" pageNumber="325" reason="1">
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<emphasis id="ABD2EAD8366F4C2FFAB3FD246C26FD46" bold="true" box="[151,552,682,708]" pageId="1" pageNumber="325">
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<taxonomicName id="5EA64D49366F4C2FFAB3FD246FCDFD46" ID-CoL="3c9a1d8c-0b87-4b31-a142-6ca9ca5a18fd" box="[151,451,682,708]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="325" phylum="Tardigrada" rank="species" species="liupanensis" status="sp. nov.">
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||
<emphasis id="ABD2EAD8366F4C2FFAB3FD246FCDFD46" bold="true" box="[151,451,682,708]" italics="true" pageId="1" pageNumber="325">Bryochoerus liupanensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="B0E157A3366F4C2FFBEFFD246C26FD46" box="[459,552,682,708]" pageId="1" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="991936CA366F4C2FFAB3FD436F7CFD64" blockId="1.[151,552,682,742]" box="[151,370,717,742]" pageId="1" pageNumber="325">
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(
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<tableCitation id="D4240371366F4C2FFABAFD436EFDFD64" box="[158,243,717,742]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Measurements (in µm) of Bryochoerus liupanensis sp. nov." pageId="1" pageNumber="325">Table 1</tableCitation>
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,
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<figureCitation id="019D2A4F366F4C2FFB25FD436F67FD64" box="[257,361,717,742]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1951,1973]" captionStartId-1="3.[151,250,1812,1834]" captionStartId-2="4.[151,250,1610,1632]" captionTargetBox-0="[291,1293,905,1929]" captionTargetBox-1="[151,1436,589,1791]" captionTargetBox-2="[151,1436,193,1588]" captionTargetId-0="figure@2.[291,1293,905,1929]" captionTargetId-1="figure@3.[151,1436,589,1791]" captionTargetId-2="figure@4.[151,1436,193,1588]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Bryochoerus liupanensis sp. nov. (holotype). a, habitus in dorsal view under 400 × magnification focused to show the cuticular sculpture on the dorsal plate; b, habitus in ventral view at 400 × magnification focused to show the ventral side of body and the claws. Scale bars = 30 µm." captionText-1="FIGURE 2. Bryochoerus liupanensis sp. nov. a, anterior part of body in dorsal view under 1000 × focused to show the cuticular sculpture on the dorsal plates (holotype); b, habitus in dorsal view under oil immersion lens at 1000 × magnification focused to show the two segmental plates after the third median plate (paratype, slide number xnc 03003); c, posterior part of body in dorsal view under 1000 × focused to show the cuticular sculpture on the dorsal plates (holotype). I – IV = segmental plates, hp = head plate, lp 1 a, lp 1 b, lp 2 a, lp 2 b, lp 3 a and lp 3 b = lateral plates, m 1, m 2 and m 3 = median plates, m 1 a, m 1 b, m 2 a, m 2 b, m 3 a and m 3 b = intersegmental plates. Scale bars = 30 µm." captionText-2="FIGURE 3. Bryochoerus liupanensis sp. nov. (holotype). a, Claws under 1000 × focused to show the spur on the internal claws on the second pair of legs, the arrows point at the spur; b, hind legs under 1000 × focused to show the sculptured let plate, the arrow points at cuticular sculpture. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/829629/files/figure.png" httpUri-1="https://zenodo.org/record/829631/files/figure.png" httpUri-2="https://zenodo.org/record/829633/files/figure.png" pageId="1" pageNumber="325">Figs. 1–3</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D1BC6541366F4C2FFAB3FC9B6CA7FC60" pageId="1" pageNumber="325" type="materials_examined">
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<paragraph id="991936CA366F4C2FFAB3FC9B6D34FCAC" blockId="1.[151,1437,789,2002]" box="[151,826,789,814]" pageId="1" pageNumber="325">
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<emphasis id="ABD2EAD8366F4C2FFAB3FC9B6F4BFCAC" bold="true" box="[151,325,789,814]" pageId="1" pageNumber="325">Type material.</emphasis>
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Holotype: Female, slide number: xnc02006.
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</paragraph>
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<paragraph id="991936CA366F4C2FFAE3FCB76A25FCF4" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
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||
<materialsCitation id="29CE3C97366F4C2FFAE3FCB76A25FCF4" ID-GBIF-Occurrence="1572326341" country="China" county="Jingyuan County" elevation="1986" latitude="35.360428" location="Liupanshan National Forest Park" longLatPrecision="1" longitude="106.31239" municipality="Liupanshan National Forest" pageId="1" pageNumber="325" specimenCount="1" stateProvince="Ningxia Hui" typeStatus="holotype">
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||
<emphasis id="ABD2EAD8366F4C2FFAE3FCB76F63FCD0" bold="true" box="[199,365,825,850]" pageId="1" pageNumber="325">
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||
<typeStatus id="461D8868366F4C2FFAE3FCB76F0CFCD0" box="[199,258,825,850]" pageId="1" pageNumber="325">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<geoCoordinate id="FC92500D366F4C2FFB58FCB76FF2FCD0" box="[380,508,825,850]" direction="north" orientation="latitude" pageId="1" pageNumber="325" precision="1" value="35.36043">35.36043N</geoCoordinate>
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||
,
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||
<geoCoordinate id="FC92500D366F4C2FF82BFCB76C99FCD0" box="[527,663,825,850]" direction="east" orientation="longitude" pageId="1" pageNumber="325" precision="1" value="106.31239">106.31239E</geoCoordinate>
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||
(
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||
<geoCoordinate id="FC92500D366F4C2FF88AFCB76D64FCD3" box="[686,874,825,850]" direction="north" orientation="latitude" pageId="1" pageNumber="325" precision="1" value="35.360428">35°21′37.55″ N</geoCoordinate>
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||
,
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||
<geoCoordinate id="FC92500D366F4C2FF959FCB76A3AFCD0" box="[893,1076,825,850]" direction="east" orientation="longitude" pageId="1" pageNumber="325" precision="1" value="106.31239">106°18′44.60″E</geoCoordinate>
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||
),
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||
<elevation id="128BD1F9366F4C2FFE6BFCB76AADFCD0" box="[1103,1187,825,850]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="1" pageNumber="325" unit="m" value="1986.0">
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||
<quantity id="5E5E9B2F366F4C2FFE6BFCB76AADFCD0" box="[1103,1187,825,850]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="1" pageNumber="325" unit="m" value="1986.0">1,986m</quantity>
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</elevation>
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asl, the
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<location id="9C796011366F4C2FFF3DFCB76F8FFCF4" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95:9C796011366F4C2FFF3DFCB76F8FFCF4" country="China" county="Jingyuan County" latitude="35.360428" longLatPrecision="1" longitude="106.31239" municipality="Liupanshan National Forest" name="Liupanshan National Forest Park" pageId="1" pageNumber="325" stateProvince="Ningxia Hui">
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<collectingMunicipality id="797DACB0366F4C2FFF3DFCB76F4AFCF7" pageId="1" pageNumber="325">Liupanshan National Forest</collectingMunicipality>
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||
Park
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</location>
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||
,
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<collectingCounty id="70784E46366F4C2FFBA9FCD36C43FCF4" box="[397,589,861,886]" pageId="1" pageNumber="325">Jingyuan County</collectingCounty>
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,
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<collectingRegion id="5B62F828366F4C2FF873FCD36CEDFCF4" box="[599,739,861,886]" country="China" name="Ningxia" pageId="1" pageNumber="325">Ningxia Hui</collectingRegion>
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Autonomous Region,
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<collectingCountry id="E1B1765A366F4C2FF9C4FCD36A29FCF4" box="[992,1063,861,886]" name="China" pageId="1" pageNumber="325">China</collectingCountry>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="991936CA366F4C2FFAE3FC0C6DDDFC18" blockId="1.[151,1437,789,2002]" box="[199,979,897,922]" pageId="1" pageNumber="325">
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<typeStatus id="461D8868366F4C2FFAE3FC0C6F37FC1B" box="[199,313,898,921]" pageId="1" pageNumber="325" type="paratype">
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||
<emphasis id="ABD2EAD8366F4C2FFAE3FC0C6F37FC1B" box="[199,313,898,921]" italics="true" pageId="1" pageNumber="325">Paratypes</emphasis>
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||
</typeStatus>
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: slide number: xnc03003, xnc03005, xnc03006, xnc03009.
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||
</paragraph>
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||
<paragraph id="991936CA366F4C2FFAE3FC2B6CA7FC60" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
|
||
<materialsCitation id="29CE3C97366F4C2FFAE3FC2B6CA7FC60" ID-GBIF-Occurrence="1572326347" country="China" county="Life Sciences" location="Li's" municipality="College" pageId="1" pageNumber="325" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
|
||
<emphasis id="ABD2EAD8366F4C2FFAE3FC2B6F80FC3C" bold="true" box="[199,398,933,958]" pageId="1" pageNumber="325">
|
||
<typeStatus id="461D8868366F4C2FFAE3FC2B6F0CFC3C" box="[199,258,933,958]" pageId="1" pageNumber="325">Type</typeStatus>
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||
repository.
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||
</emphasis>
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||
<location id="9C796011366F4C2FFBBFFC2B6FC8FC3C" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95:9C796011366F4C2FFBBFFC2B6FC8FC3C" box="[411,454,933,958]" country="China" county="Life Sciences" municipality="College" name="Li's" pageId="1" pageNumber="325" stateProvince="Shaanxi">Li’s</location>
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||
tardigrade collection,
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||
<location id="9C796011366F4C2FF8F0FC2B6DBCFC3C" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95:9C796011366F4C2FF8F0FC2B6DBCFC3C" box="[724,946,933,958]" country="China" county="Life Sciences" municipality="College" name="Molecular Ecology" pageId="1" pageNumber="325" stateProvince="Shaanxi">Molecular Ecology</location>
|
||
,
|
||
<location id="9C796011366F4C2FF9E0FC286A46FC3F" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95:9C796011366F4C2FF9E0FC286A46FC3F" box="[964,1096,934,957]" country="China" county="Life Sciences" municipality="College" name="Department" pageId="1" pageNumber="325" stateProvince="Shaanxi">Department</location>
|
||
of
|
||
<location id="9C796011366F4C2FFE5DFC2B6ADAFC3C" LSID="urn:lsid:plazi:treatment:110F87DC366F4C2DFAB3FD246AA6FD95:9C796011366F4C2FFE5DFC2B6ADAFC3C" box="[1145,1236,933,958]" country="China" county="Life Sciences" municipality="College" name="Biology" pageId="1" pageNumber="325" stateProvince="Shaanxi">Biology</location>
|
||
,
|
||
<collectingMunicipality id="797DACB0366F4C2FFEC1FC2B6B32FC3C" box="[1253,1340,933,958]" pageId="1" pageNumber="325">College</collectingMunicipality>
|
||
of
|
||
<collectingCounty id="70784E46366F4C2FFF49FC2B6EF7FC60" pageId="1" pageNumber="325">Life Sciences</collectingCounty>
|
||
of
|
||
<collectingRegion id="5B62F828366F4C2FFB3AFC476F76FC60" box="[286,376,969,994]" country="China" name="Shaanxi" pageId="1" pageNumber="325">Shaanxi</collectingRegion>
|
||
Normal University,
|
||
<collectingCountry id="E1B1765A366F4C2FF87AFC476CABFC60" box="[606,677,969,994]" name="China" pageId="1" pageNumber="325">China</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1BC6541366F4C2FFAE3FC636AABFB5C" pageId="1" pageNumber="325" type="diagnosis">
|
||
<paragraph id="991936CA366F4C2FFAE3FC636AABFB5C" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
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||
<emphasis id="ABD2EAD8366F4C2FFAE3FC636F4FFB84" bold="true" box="[199,321,1005,1030]" pageId="1" pageNumber="325">Diagnosis.</emphasis>
|
||
Apart from the head appendages including cirrus
|
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<emphasis id="ABD2EAD8366F4C2FF9A2FC606DECFB87" box="[902,994,1006,1029]" italics="true" pageId="1" pageNumber="325">internus</emphasis>
|
||
, cephalic papillae and cirrus
|
||
<emphasis id="ABD2EAD8366F4C2FFF12FC606B9BFB87" box="[1334,1429,1006,1029]" italics="true" pageId="1" pageNumber="325">externus</emphasis>
|
||
, only lateral cirrus
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||
<emphasis id="ABD2EAD8366F4C2FFB40FB9C6F7BFBAB" box="[356,373,1042,1065]" italics="true" pageId="1" pageNumber="325">A</emphasis>
|
||
with clava present; dorsal plates and lateral plates with double sculpture including large bright roundish granule-like dots and small dark roundish dots; median plates 1 and 2 divided transversely into four parts, median plate 3 divided transversely into three parts; two lateral plates on each side of all median plates,
|
||
<quantity id="5E5E9B2F366F4C2FFF07FBD46B52FBF0" box="[1315,1372,1113,1138]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="1" pageNumber="325" unit="in" value="12.0">12 in</quantity>
|
||
total; paired plates with a transverse smooth band; terminal plate with two longitudinal ridges which divided the terminal plate into three parts, terminal plate without notches; ventral side of body without plates or platelets; papilla not visible and dentate collar absent on legs IV; small spurs present on internal claws on all legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1BC6541366F4C2FFAE3FB676C2FF8E4" pageId="1" pageNumber="325" type="description">
|
||
<paragraph id="991936CA366F4C2FFAE3FB676C12F930" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
|
||
<emphasis id="ABD2EAD8366F4C2FFAE3FB676F57FA80" bold="true" box="[199,345,1257,1282]" pageId="1" pageNumber="325">Description.</emphasis>
|
||
Colour orange. Eye spots absent or not visible after slide preparation. Apart from head appendages, including cirrus
|
||
<emphasis id="ABD2EAD8366F4C2FFBFBFA806C35FAA7" box="[479,571,1294,1317]" italics="true" pageId="1" pageNumber="325">internus</emphasis>
|
||
, cirrus
|
||
<emphasis id="ABD2EAD8366F4C2FF8B4FA806CE1FAA7" box="[656,751,1294,1317]" italics="true" pageId="1" pageNumber="325">externus</emphasis>
|
||
and cephalic papillae (secondary clava), only lateral cirrus
|
||
<emphasis id="ABD2EAD8366F4C2FFFAFFA806B92FAA7" box="[1419,1436,1294,1317]" italics="true" pageId="1" pageNumber="325">A</emphasis>
|
||
with clava (primary clava) are present; cephalic papillae drop-shaped; stylet supports not visible, pharyngeal bulb roundish; dorsal plates comprise of head plate, scapular plate, first median plate, first paired plates, second median plate, second paired plates, third median plate, and terminal plate; all median plates divided transversely, median plate 1 and median plate 2 divided into four parts, while median plate 3 divided into three parts, the normal parts of all median plates are the same as
|
||
<taxonomicName id="5EA64D49366F4C2FF865FA4F6CDEFA5B" box="[577,720,1473,1497]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="1" pageNumber="325" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="ABD2EAD8366F4C2FF865FA4F6CDEFA5B" box="[577,720,1473,1497]" italics="true" pageId="1" pageNumber="325">Bryodelphax</emphasis>
|
||
</taxonomicName>
|
||
species, however all median plates are followed with two intersegmental plates, two lateral plates present on each side of all median plates,
|
||
<quantity id="5E5E9B2F366F4C2FFE03FA686A6EFA7C" box="[1063,1120,1509,1534]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="1" pageNumber="325" unit="in" value="12.0">12 in</quantity>
|
||
total, two paired plates with a transverse smooth band, terminal plate with two longitudinal ridges which divide terminal plate into three parts, terminal plate without incisions; all plates, including dorsal plates and lateral plates, with “double sculpture”, including large bright roundish pores and small dark roundish dots that are pillars within the cuticle, pores are scarce and distributed loosely whereas dark dots are arranged regularly and densely (
|
||
<figureCitation id="019D2A4F366F4C2FFE59F9FB6AC2F90C" box="[1149,1228,1653,1678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1973]" captionTargetBox="[291,1293,905,1929]" captionTargetId="figure@2.[291,1293,905,1929]" captionTargetPageId="2" captionText="FIGURE 1. Bryochoerus liupanensis sp. nov. (holotype). a, habitus in dorsal view under 400 × magnification focused to show the cuticular sculpture on the dorsal plate; b, habitus in ventral view at 400 × magnification focused to show the ventral side of body and the claws. Scale bars = 30 µm." httpUri="https://zenodo.org/record/829629/files/figure.png" pageId="1" pageNumber="325">Figs. 1</figureCitation>
|
||
a, 2); ventral side without plate or platelets (
|
||
<figureCitation id="019D2A4F366F4C2FFB98F9176FF5F930" box="[444,507,1689,1714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1951,1973]" captionTargetBox="[291,1293,905,1929]" captionTargetId="figure@2.[291,1293,905,1929]" captionTargetPageId="2" captionText="FIGURE 1. Bryochoerus liupanensis sp. nov. (holotype). a, habitus in dorsal view under 400 × magnification focused to show the cuticular sculpture on the dorsal plate; b, habitus in ventral view at 400 × magnification focused to show the ventral side of body and the claws. Scale bars = 30 µm." httpUri="https://zenodo.org/record/829629/files/figure.png" pageId="1" pageNumber="325">Fig. 1</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<paragraph id="991936CA366F4C2FFAE3F9336CBAF89C" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
|
||
Spine on legs I and papilla on legs IV not visible under light microscope. Dentate collar absent on legs IV. Internal claws on all legs with small spur, external claws spurless (
|
||
<figureCitation id="019D2A4F366F4C2FF9A1F96F6DC9F978" box="[901,967,1761,1786]" captionStart="FIGURE 3" captionStartId="4.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1588]" captionTargetId="figure@4.[151,1436,193,1588]" captionTargetPageId="4" captionText="FIGURE 3. Bryochoerus liupanensis sp. nov. (holotype). a, Claws under 1000 × focused to show the spur on the internal claws on the second pair of legs, the arrows point at the spur; b, hind legs under 1000 × focused to show the sculptured let plate, the arrow points at cuticular sculpture. Scale bars = 10 µm." httpUri="https://zenodo.org/record/829633/files/figure.png" pageId="1" pageNumber="325">Fig. 3</figureCitation>
|
||
a), the upper part of all legs were covered with plate with sculpture of small dots (
|
||
<figureCitation id="019D2A4F366F4C2FF870F88B6C98F89C" box="[596,662,1797,1822]" captionStart="FIGURE 3" captionStartId="4.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1588]" captionTargetId="figure@4.[151,1436,193,1588]" captionTargetPageId="4" captionText="FIGURE 3. Bryochoerus liupanensis sp. nov. (holotype). a, Claws under 1000 × focused to show the spur on the internal claws on the second pair of legs, the arrows point at the spur; b, hind legs under 1000 × focused to show the sculptured let plate, the arrow points at cuticular sculpture. Scale bars = 10 µm." httpUri="https://zenodo.org/record/829633/files/figure.png" pageId="1" pageNumber="325">Fig. 3</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<paragraph id="991936CA366F4C2FFAE3F8A46F78F8C0" blockId="1.[151,1437,789,2002]" box="[199,374,1833,1858]" pageId="1" pageNumber="325">Eggs unknown.</paragraph>
|
||
<paragraph id="991936CA366F4C2FFAE3F8C06C2FF8E4" blockId="1.[151,1437,789,2002]" box="[199,545,1869,1894]" pageId="1" pageNumber="325">
|
||
For measurements see
|
||
<tableCitation id="D4240371366F4C2FFBE3F8C36C15F8E4" box="[455,539,1869,1894]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Measurements (in µm) of Bryochoerus liupanensis sp. nov." pageId="1" pageNumber="325">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1BC6541366F4C2DFAE3F8FF6D7AFF32" lastPageId="3" lastPageNumber="327" pageId="1" pageNumber="325" type="discussion">
|
||
<paragraph id="991936CA366F4C2FFAE3F8FF6C94F850" blockId="1.[151,1437,789,2002]" pageId="1" pageNumber="325">
|
||
<emphasis id="ABD2EAD8366F4C2FFAE3F8FF6F35F808" bold="true" box="[199,315,1905,1930]" pageId="1" pageNumber="325">Remarks.</emphasis>
|
||
Please see
|
||
<bibRefCitation id="FD374B3B366F4C2FFB9BF8FF6D25F808" author="Michalczyk" box="[447,811,1905,1930]" pageId="1" pageNumber="325" refString="Michalczyk, L. & Kaczmarek, L. (2006) Revision of the Echiniscus bigranulatus group with a description of a new species Echiniscus madonnae (Tardigrada: Heterotardigrada: Echiniscidae) from South America. Zootaxa, 1154, 1 - 26." type="journal article" year="2006">Michalczyk & Kaczmarek 2006</bibRefCitation>
|
||
or
|
||
<bibRefCitation id="FD374B3B366F4C2FF977F8FF6A5BF808" author="Kaczmarek" box="[851,1109,1905,1930]" pageId="1" pageNumber="325" refString="Kaczmarek, L., Zawierucha, K., Smykla, J. & Michalczyk, L. (2012) Tardigrada of the Revdalen (Spitsbergen) with the descriptions of two new species: Bryodelphax parvuspolaris (Heterotardigrada) and Isohypsibius coulsoni (Eutardigrada). Polar Biology, 35, 1013 - 1026." type="journal article" year="2012">
|
||
Kaczmarek
|
||
<emphasis id="ABD2EAD8366F4C2FF9FFF8FC6A02F80B" box="[987,1036,1905,1929]" italics="true" pageId="1" pageNumber="325">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
for more information on the anatomy of ‘double sculpturing’. Whether the spine on legs I and papilla on legs IV are indeed absent, has to be confirmed by SEM observations in the future.
|
||
</paragraph>
|
||
<caption id="CDD96642366C4C2CFAB3FF166D85FF2D" box="[151,907,151,175]" pageId="2" pageNumber="326">
|
||
<paragraph id="991936CA366C4C2CFAB3FF166D85FF2D" blockId="2.[151,907,151,175]" box="[151,907,151,175]" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFAB3FF166F04FF2D" bold="true" box="[151,266,152,175]" pageId="2" pageNumber="326">TABLE 1.</emphasis>
|
||
Measurements (in µm) of
|
||
<taxonomicName id="5EA64D49366C4C2CF802FF196D20FF2D" box="[550,814,151,175]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="326" phylum="Tardigrada" rank="species" species="liupanensis" status="sp. nov.">
|
||
<emphasis id="ABD2EAD8366C4C2CF802FF196D20FF2D" box="[550,814,151,175]" italics="true" pageId="2" pageNumber="326">Bryochoerus liupanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0E157A3366C4C2CF911FF166D85FF2D" box="[821,907,152,175]" pageId="2" pageNumber="326" rank="species">
|
||
<emphasis id="ABD2EAD8366C4C2CF911FF166D8AFF2D" bold="true" box="[821,900,152,175]" pageId="2" pageNumber="326">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="991936CA366C4C2CFABBFF446B4AFF62" blockId="2.[159,1348,202,255]" box="[159,1348,202,224]" pageId="2" pageNumber="326">Structure xnc02006 xnc03003 xnc03005 xnc03006 xnc03009</paragraph>
|
||
<paragraph id="991936CA366C4C2DFBD2FF676D7AFF32" blockId="2.[159,1348,202,255]" lastBlockId="3.[151,1436,151,536]" lastPageId="3" lastPageNumber="327" pageId="2" pageNumber="326">
|
||
female female female female female
|
||
<emphasis id="ABD2EAD8366D4C2DFAE3FF196F40FF32" bold="true" box="[199,334,151,176]" pageId="3" pageNumber="327">Etymology.</emphasis>
|
||
The new species is named after the
|
||
<typeStatus id="461D8868366D4C2DF8C7FF166D1DFF32" box="[739,787,152,176]" pageId="3" pageNumber="327">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="991936CA366C4C2CFBD2FE9D6B65FCC0" pageId="2" pageNumber="326">
|
||
<table id="EBA6C46A366CB3D1FABBFE9D6B76FCC0" box="[159,1400,275,834]" colsContinueIn="7.[159,1395,275,834]" gridcols="11" gridrows="4" pageId="2" pageNumber="326">
|
||
<tr id="27963488366CB3D1FABBFE9D6B76FEAA" box="[159,1400,275,296]" gridrow="0" pageId="2" pageNumber="326" rowspan-0="1">
|
||
<th id="64475DF4366CB3D1FBD2FE9D6C2BFEAA" box="[502,549,275,296]" gridcol="1" gridrow="0" pageId="2" pageNumber="326">µm</th>
|
||
<th id="64475DF4366CB3D1F874FE9D6C87FEAA" box="[592,649,275,296]" gridcol="2" gridrow="0" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF874FE9D6C7FFEAA" box="[592,625,275,296]" italics="true" pageId="2" pageNumber="326">psc</emphasis>
|
||
</th>
|
||
<th id="64475DF4366CB3D1F88DFE9D6CD6FEAA" box="[681,728,275,296]" gridcol="3" gridrow="0" pageId="2" pageNumber="326">µm</th>
|
||
<th id="64475DF4366CB3D1F927FE9D6D32FEAA" box="[771,828,275,296]" gridcol="4" gridrow="0" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF927FE9D6D2AFEAA" box="[771,804,275,296]" italics="true" pageId="2" pageNumber="326">psc</emphasis>
|
||
</th>
|
||
<th id="64475DF4366CB3D1F94DFE9D6D99FEAA" box="[873,919,275,296]" gridcol="5" gridrow="0" pageId="2" pageNumber="326">µm</th>
|
||
<th id="64475DF4366CB3D1F9E6FE9D6DF5FEAA" box="[962,1019,275,296]" gridcol="6" gridrow="0" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF9E6FE9D6DEDFEAA" box="[962,995,275,296]" italics="true" pageId="2" pageNumber="326">psc</emphasis>
|
||
</th>
|
||
<th id="64475DF4366CB3D1FE00FE9D6A5DFEAA" box="[1060,1107,275,296]" gridcol="7" gridrow="0" pageId="2" pageNumber="326">µm</th>
|
||
<th id="64475DF4366CB3D1FE5BFE9D6AB6FEAA" box="[1151,1208,275,296]" gridcol="8" gridrow="0" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFE5BFE9D6AAFFEAA" box="[1151,1185,275,296]" italics="true" pageId="2" pageNumber="326">psc</emphasis>
|
||
</th>
|
||
<th id="64475DF4366CB3D1FEC4FE9D6B01FEAA" box="[1248,1295,275,296]" gridcol="9" gridrow="0" pageId="2" pageNumber="326">µm</th>
|
||
<th id="64475DF4366CB3D1FF1BFE9D6B76FEAA" box="[1343,1400,275,296]" gridcol="10" gridrow="0" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFF1BFE9D6B6EFEAA" box="[1343,1376,275,296]" italics="true" pageId="2" pageNumber="326">psc</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="27963488366CB3D1FABBFEB26B76FDA2" box="[159,1400,316,544]" gridrow="1" pageId="2" pageNumber="326">
|
||
<th id="64475DF4366CB3D1FABBFEB26FA2FDA2" box="[159,428,316,544]" gridcol="0" gridrow="1" pageId="2" pageNumber="326">
|
||
Body length Scapular plate length Cirrus
|
||
<emphasis id="ABD2EAD8366C4C2CFAC0FE1E6F39FE27" box="[228,311,400,421]" italics="true" pageId="2" pageNumber="326">internus</emphasis>
|
||
Cephalic papilla Cirrus
|
||
<emphasis id="ABD2EAD8366C4C2CFAC0FE6C6F34FE75" box="[228,314,482,503]" italics="true" pageId="2" pageNumber="326">externus</emphasis>
|
||
Clava
|
||
</th>
|
||
<td id="64475DF4366CB3D1FBD2FEB26C2BFDA2" box="[502,549,316,544]" gridcol="1" gridrow="1" pageId="2" pageNumber="326">163 17.3 5.1 3.1 9.5 3.0</td>
|
||
<td id="64475DF4366CB3D1F874FEB26C87FDA2" box="[592,649,316,544]" gridcol="2" gridrow="1" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF874FE006C72FDA2" box="[592,636,398,544]" italics="true" pageId="2" pageNumber="326">29.5 17.9 54.9 17.3</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F88DFEB26CD6FDA2" box="[681,728,316,544]" gridcol="3" gridrow="1" pageId="2" pageNumber="326">134 16.7 3.5 3.0 8.6 3.4</td>
|
||
<td id="64475DF4366CB3D1F927FEB26D32FDA2" box="[771,828,316,544]" gridcol="4" gridrow="1" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF927FE006D21FDA2" box="[771,815,398,544]" italics="true" pageId="2" pageNumber="326">21.0 18.0 51.5 20.4</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F94DFEB26D99FDA2" box="[873,919,316,544]" gridcol="5" gridrow="1" pageId="2" pageNumber="326">125 10.9 3.4 2.5 7.1 3.8</td>
|
||
<td id="64475DF4366CB3D1F9E6FEB26DF5FDA2" box="[962,1019,316,544]" gridcol="6" gridrow="1" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF9E6FE006DE0FDA2" box="[962,1006,398,544]" italics="true" pageId="2" pageNumber="326">31.2 22.9 65.1 34.9</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FE00FEB26A5DFDA2" box="[1060,1107,316,544]" gridcol="7" gridrow="1" pageId="2" pageNumber="326">151 17.9 4.7 3.1 9.3 2.8</td>
|
||
<td id="64475DF4366CB3D1FE5BFEB26AB6FDA2" box="[1151,1208,316,544]" gridcol="8" gridrow="1" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFE5BFE006AA5FDA2" box="[1151,1195,398,544]" italics="true" pageId="2" pageNumber="326">26.3 17.3 52.0 15.6</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FEC4FEB26B01FDA2" box="[1248,1295,316,544]" gridcol="9" gridrow="1" pageId="2" pageNumber="326">155 18.5 4.5 2.7 10.5 4.2</td>
|
||
<td id="64475DF4366CB3D1FF1BFEB26B76FDA2" box="[1343,1400,316,544]" gridcol="10" gridrow="1" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFF1BFE006B65FDA2" box="[1343,1387,398,544]" italics="true" pageId="2" pageNumber="326">24.3 14.6 56.8 22.7</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="27963488366CB3D1FABBFDBA6B76FC9A" box="[159,1400,564,792]" gridrow="2" pageId="2" pageNumber="326">
|
||
<th id="64475DF4366CB3D1FABBFDBA6FA2FC9A" box="[159,428,564,792]" gridcol="0" gridrow="2" pageId="2" pageNumber="326">
|
||
Cirrus
|
||
<emphasis id="ABD2EAD8366C4C2CFAC0FDBA6EFDFDCB" box="[228,243,564,585]" italics="true" pageId="2" pageNumber="326">A</emphasis>
|
||
Cirrus
|
||
<emphasis id="ABD2EAD8366C4C2CFAC0FDD06EFDFDF1" box="[228,243,606,627]" italics="true" pageId="2" pageNumber="326">A</emphasis>
|
||
/Body length ratio Cirrus
|
||
<emphasis id="ABD2EAD8366C4C2CFAC0FD066EF1FD1F" box="[228,255,648,669]" italics="true" pageId="2" pageNumber="326">int</emphasis>
|
||
/
|
||
<emphasis id="ABD2EAD8366C4C2CFB22FD066F2DFD1F" box="[262,291,648,669]" italics="true" pageId="2" pageNumber="326">ext</emphasis>
|
||
length ratio Claws I Claws II Claws III
|
||
</th>
|
||
<td id="64475DF4366CB3D1FBD2FDBA6C2BFC9A" box="[502,549,564,792]" gridcol="1" gridrow="2" pageId="2" pageNumber="326">20.8 13% 54% 8.1 9.2 9.5</td>
|
||
<td id="64475DF4366CB3D1F874FDBA6C87FC9A" box="[592,649,564,792]" gridcol="2" gridrow="2" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF874FDBA6C72FC9A" box="[592,649,564,792]" italics="true" pageId="2" pageNumber="326">120.2 46.8 53.2 54.9</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F88DFDBA6CD6FC9A" box="[681,728,564,792]" gridcol="3" gridrow="2" pageId="2" pageNumber="326">23.5 18% 41% 8.7 8.8 8.8</td>
|
||
<td id="64475DF4366CB3D1F927FDBA6D32FC9A" box="[771,828,564,792]" gridcol="4" gridrow="2" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF927FDBA6D21FC9A" box="[771,828,564,792]" italics="true" pageId="2" pageNumber="326">140.7 52.1 52.7 52.7</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F94DFDBA6D99FC9A" box="[873,919,564,792]" gridcol="5" gridrow="2" pageId="2" pageNumber="326">15.9 48% 48% 7.7 7.8 8.0</td>
|
||
<td id="64475DF4366CB3D1F9E6FDBA6DF5FC9A" box="[962,1019,564,792]" gridcol="6" gridrow="2" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF9E6FDBA6DE0FC9A" box="[962,1019,564,792]" italics="true" pageId="2" pageNumber="326">145.9 70.6 71.6 73.4</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FE00FDBA6A5DFC9A" box="[1060,1107,564,792]" gridcol="7" gridrow="2" pageId="2" pageNumber="326">23.6 16% 51% 8.5 8.2 8.0</td>
|
||
<td id="64475DF4366CB3D1FE5BFDBA6AB6FC9A" box="[1151,1208,564,792]" gridcol="8" gridrow="2" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFE5BFDBA6AA5FC9A" box="[1151,1208,564,792]" italics="true" pageId="2" pageNumber="326">131.8 47.5 45.8 44.7</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FEC4FDBA6B01FC9A" box="[1248,1295,564,792]" gridcol="9" gridrow="2" pageId="2" pageNumber="326">29.5 19% 43% 8.6 8.5 8.0</td>
|
||
<td id="64475DF4366CB3D1FF1BFDBA6B76FC9A" box="[1343,1400,564,792]" gridcol="10" gridrow="2" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFF1BFDBA6B65FC9A" box="[1343,1400,564,792]" italics="true" pageId="2" pageNumber="326">159.5 46.5 45.9 43.2</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="27963488366CB3D1FABBFCA26B76FCC0" box="[159,1400,812,834]" gridrow="3" pageId="2" pageNumber="326">
|
||
<th id="64475DF4366CB3D1FABBFCA26FA2FCC0" box="[159,428,812,834]" gridcol="0" gridrow="3" pageId="2" pageNumber="326">Claws IV</th>
|
||
<td id="64475DF4366CB3D1FBD2FCA26C2BFCC0" box="[502,549,812,834]" gridcol="1" gridrow="3" pageId="2" pageNumber="326">9.4</td>
|
||
<td id="64475DF4366CB3D1F874FCA26C87FCC0" box="[592,649,812,834]" gridcol="2" gridrow="3" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF874FCA26C72FCC0" box="[592,636,812,834]" italics="true" pageId="2" pageNumber="326">54.3</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F88DFCA26CD6FCC0" box="[681,728,812,834]" gridcol="3" gridrow="3" pageId="2" pageNumber="326">8.8</td>
|
||
<td id="64475DF4366CB3D1F927FCA26D32FCC0" box="[771,828,812,834]" gridcol="4" gridrow="3" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF927FCA26D21FCC0" box="[771,815,812,834]" italics="true" pageId="2" pageNumber="326">52.7</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1F94DFCA26D99FCC0" box="[873,919,812,834]" gridcol="5" gridrow="3" pageId="2" pageNumber="326">8.8</td>
|
||
<td id="64475DF4366CB3D1F9E6FCA26DF5FCC0" box="[962,1019,812,834]" gridcol="6" gridrow="3" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CF9E6FCA26DE0FCC0" box="[962,1006,812,834]" italics="true" pageId="2" pageNumber="326">80.7</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FE00FCA26A5DFCC0" box="[1060,1107,812,834]" gridcol="7" gridrow="3" pageId="2" pageNumber="326">8.9</td>
|
||
<td id="64475DF4366CB3D1FE5BFCA26AB6FCC0" box="[1151,1208,812,834]" gridcol="8" gridrow="3" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFE5BFCA26AA5FCC0" box="[1151,1195,812,834]" italics="true" pageId="2" pageNumber="326">49.7</emphasis>
|
||
</td>
|
||
<td id="64475DF4366CB3D1FEC4FCA26B01FCC0" box="[1248,1295,812,834]" gridcol="9" gridrow="3" pageId="2" pageNumber="326">8.3</td>
|
||
<td id="64475DF4366CB3D1FF1BFCA26B76FCC0" box="[1343,1400,812,834]" gridcol="10" gridrow="3" pageId="2" pageNumber="326">44.9</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="CDD96642366C4C2CFAB3F8116C3FF870" httpUri="https://zenodo.org/record/829629/files/figure.png" pageId="2" pageNumber="326" targetBox="[291,1293,905,1929]" targetPageId="2">
|
||
<paragraph id="991936CA366C4C2CFAB3F8116C3FF870" blockId="2.[151,1436,1951,2034]" pageId="2" pageNumber="326">
|
||
<emphasis id="ABD2EAD8366C4C2CFAB3F8116F1DF836" bold="true" box="[151,275,1951,1973]" pageId="2" pageNumber="326">FIGURE 1.</emphasis>
|
||
<taxonomicName id="5EA64D49366C4C2CFB3EF8116C1BF837" box="[282,533,1951,1973]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="2" pageNumber="326" phylum="Tardigrada" rank="species" species="liupanensis" status="sp. nov.">
|
||
<emphasis id="ABD2EAD8366C4C2CFB3EF8116C1BF837" box="[282,533,1951,1973]" italics="true" pageId="2" pageNumber="326">Bryochoerus liupanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0E157A3366C4C2CF838F8116C62F837" box="[540,620,1951,1973]" pageId="2" pageNumber="326" rank="species">
|
||
<emphasis id="ABD2EAD8366C4C2CF838F8116C68F836" bold="true" box="[540,614,1951,1972]" pageId="2" pageNumber="326">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(holotype). a, habitus in dorsal view under 400× magnification focused to show the cuticular sculpture on the dorsal plate; b, habitus in ventral view at 400× magnification focused to show the ventral side of body and the claws. Scale bars = 30 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D1BC6541366D4C2DFAE3FF326AA6FD95" pageId="3" pageNumber="327" type="diagnosis">
|
||
<paragraph id="991936CA366D4C2DFAE3FF326AA6FD95" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="327">
|
||
<emphasis id="ABD2EAD8366D4C2DFAE3FF326FC1FF57" bold="true" box="[199,463,188,213]" pageId="3" pageNumber="327">Differential diagnosis.</emphasis>
|
||
The monotypic genus
|
||
<taxonomicName id="5EA64D49366D4C2DF8F7FF326D6CFF56" box="[723,866,188,212]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="ABD2EAD8366D4C2DF8F7FF326D6CFF56" box="[723,866,188,212]" italics="true" pageId="3" pageNumber="327">Bryochoerus</emphasis>
|
||
</taxonomicName>
|
||
contained only the nominate species
|
||
<taxonomicName id="5EA64D49366D4C2DFF29FF326F15FF7A" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="species" species="intermedius">
|
||
<emphasis id="ABD2EAD8366D4C2DFF29FF326F15FF7A" italics="true" pageId="3" pageNumber="327">Bryochoerus intermedius</emphasis>
|
||
</taxonomicName>
|
||
. There are three subspecies in
|
||
<taxonomicName id="5EA64D49366D4C2DF8BFFF6F6D4AFF7A" box="[667,836,224,248]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="species" species="intermedius">
|
||
<emphasis id="ABD2EAD8366D4C2DF8BFFF6F6D4AFF7A" box="[667,836,224,248]" italics="true" pageId="3" pageNumber="327">B. intermedius</emphasis>
|
||
</taxonomicName>
|
||
. They are
|
||
<taxonomicName id="5EA64D49366D4C2DF9EBFF6F6B58FF7A" authority="Murray, 1910" authorityName="Murray" authorityYear="1910" box="[975,1366,224,248]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="subSpecies" species="intermedius" subSpecies="intermedius">
|
||
<emphasis id="ABD2EAD8366D4C2DF9EBFF6F6A98FF7A" box="[975,1174,224,248]" italics="true" pageId="3" pageNumber="327">B. i. intermedius</emphasis>
|
||
(
|
||
<bibRefCitation id="FD374B3B366D4C2DFE8EFF6E6B40FF7A" author="Murray" box="[1194,1358,224,248]" pageId="3" pageNumber="327" refString="Murray, J. (1910) Part V Tardigrada. In: British Antarctic Expedition 1907 -, under the command of Sir E. H. Shackleton, C. F. O. Reports on the scientific investigations. Fol. 1. Biology. William Heinemann Publisher, London, pp. 83 - 187." type="journal article" year="1910">Murray, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="ABD2EAD8366D4C2DFF4CFF6F6F07FE9E" italics="true" pageId="3" pageNumber="327">B. i. hawaiicus</emphasis>
|
||
(
|
||
<bibRefCitation id="FD374B3B366D4C2DFB3BFE8A6FB4FE9E" author="Thulin" box="[287,442,260,285]" pageId="3" pageNumber="327" refString="Thulin, G. (1928) Uber die phylogenie und das system der tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article" year="1928">Thulin, 1928</bibRefCitation>
|
||
) and
|
||
<emphasis id="ABD2EAD8366D4C2DF82CFE8B6C83FE9E" box="[520,653,260,284]" italics="true" pageId="3" pageNumber="327">B. i. laevis</emphasis>
|
||
(
|
||
<bibRefCitation id="FD374B3B366D4C2DF880FE8A6D47FE9F" author="Marcus" box="[676,841,260,285]" pageId="3" pageNumber="327" refString="Marcus, E. (1936) Tardigrada. In: Schulze, F. E., Kukenthal, W. & Heider, K. (Eds.), Das Tierreich. Fol. 66. Walter de Gruyter, Berlin und Leipzig, pp. 1 - 340." type="journal article" year="1936">Marcus, 1936</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="FD374B3B366D4C2DF943FE8A6A24FE9E" author="Kristensen" box="[871,1066,260,285]" pageId="3" pageNumber="327" refString="Kristensen, R. M. (1987) Generic revision of the Echiniscidae (Heterotardigrada), with a discussion of the origin of the family. In: Bertolani, R. (Ed.), Biology of tardigrades. Selected symposia and monographs U. Z. I. Fol. 1. Mucchi, Modena, pp. 261 - 335." type="journal article" year="1987">Kristensen 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="FD374B3B366D4C2DFE1AFE8A6B9EFE9E" author="Ramazzotti" box="[1086,1424,260,285]" pageId="3" pageNumber="327" refString="Ramazzotti, G. & Maucci, W. (1983) Il Phylum Tardigrada. 3 rd ed. Memorie dell'Isituto Italiano di Idrobiologia, 41, 1 - 1012. [CW Beasley, English translation]" type="journal article" year="1983">Ramazzotti & Maucci 1983</bibRefCitation>
|
||
).
|
||
<taxonomicName id="5EA64D49366D4C2DFAB3FEA66FBCFEC2" box="[151,434,296,320]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="species" species="liupanensis" status="sp. nov.">
|
||
<emphasis id="ABD2EAD8366D4C2DFAB3FEA66FBCFEC2" box="[151,434,296,320]" italics="true" pageId="3" pageNumber="327">Bryochoerus liupanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0E157A3366D4C2DFB9BFEA96C10FEC2" box="[447,542,295,320]" pageId="3" pageNumber="327" rank="species">
|
||
<emphasis id="ABD2EAD8366D4C2DFB9BFEA96C10FEC2" bold="true" box="[447,542,295,320]" pageId="3" pageNumber="327">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="5EA64D49366D4C2DF8E7FEA66A7AFEC2" box="[707,1140,296,320]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="subSpecies" species="intermedius" subSpecies="intermedius">
|
||
<emphasis id="ABD2EAD8366D4C2DF8E7FEA66A7AFEC2" box="[707,1140,296,320]" italics="true" pageId="3" pageNumber="327">Bryochoerus intermedius intermedius</emphasis>
|
||
</taxonomicName>
|
||
in having orange colour instead of colourless, or yellowish, or else greenish-yellow as in
|
||
<taxonomicName id="5EA64D49366D4C2DF95EFEC36A38FEE6" box="[890,1078,332,356]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="subSpecies" species="intermedius" subSpecies="intermedius">
|
||
<emphasis id="ABD2EAD8366D4C2DF95EFEC36A38FEE6" box="[890,1078,332,356]" italics="true" pageId="3" pageNumber="327">B. i. intermedius</emphasis>
|
||
</taxonomicName>
|
||
; a different cuticular sculpture; presence of lateral plates; lacking papilla and dentate collar on legs IV; and presence of spurs on the internal claws of all legs. The new species differs from
|
||
<emphasis id="ABD2EAD8366D4C2DF841FE1B6D00FE2E" box="[613,782,404,428]" italics="true" pageId="3" pageNumber="327">B. i. hawaiicus</emphasis>
|
||
in having different cuticular sculpture; presence of lateral plates; and presence of spurs on the internal claws of all legs.
|
||
<taxonomicName id="5EA64D49366D4C2DF97FFE366A7CFE52" box="[859,1138,440,464]" class="Heterotardigrada" family="Echiniscidae" genus="Bryochoerus" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="327" phylum="Tardigrada" rank="species" species="liupanensis" status="sp. nov.">
|
||
<emphasis id="ABD2EAD8366D4C2DF97FFE366A7CFE52" box="[859,1138,440,464]" italics="true" pageId="3" pageNumber="327">Bryochoerus liupanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0E157A3366D4C2DFE5EFE396ADBFE52" box="[1146,1237,439,464]" pageId="3" pageNumber="327" rank="species">
|
||
<emphasis id="ABD2EAD8366D4C2DFE5EFE396ADBFE52" bold="true" box="[1146,1237,439,464]" pageId="3" pageNumber="327">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<emphasis id="ABD2EAD8366D4C2DFF49FE376ED6FE76" italics="true" pageId="3" pageNumber="327">B. i. laevis</emphasis>
|
||
in having a different cuticular sculpture; presence of lateral plates in a different shape; a facetted terminal plate; lacking spine on the first pair of legs; and lacking papilla and dentate collar on legs IV.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |