182 lines
20 KiB
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182 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="10FF4E7FD33E5D948060BD90AAA0BF86" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. besleticus besleticus Kurnakov 1972" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="26" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="25" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
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<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
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<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
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<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
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<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-12-12</mods:number>
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<mods:detail type="volume">
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<mods:number>463</mods:number>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
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<treatment ID-GBIF-Taxon="182234822" LSID="urn:lsid:plazi:treatment:10FF4E7FD33E5D948060BD90AAA0BF86" httpUri="http://treatment.plazi.org/id/10FF4E7FD33E5D948060BD90AAA0BF86" lastPageId="6" lastPageNumber="26" pageId="4" pageNumber="25">
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<subSubSection pageId="4" pageNumber="25" type="nomenclature">
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName LSID="10FF4E7F-D33E-5D94-8060-BD90AAA0BF86" authority="besleticus Kurnakov, 1972" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus Kurnakov, 1972</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="4" pageNumber="25">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="4" pageNumber="25">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="4" pageNumber="25">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="4" pageNumber="25">, 4</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="4" pageNumber="25">, 7</figureCitation>
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<figureCitation captionStart="Figures 16–21" captionStartId="F9" captionText="Figures 16 - 21. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus besleticus and Carabus (Archiplectes) besleticus mtsaranus. 16 - 18 Carabus (Archiplectes) besleticus besleticus 19 - 21 Carabus (Archiplectes) besleticus mtsaranus." pageId="4" pageNumber="25">, 16-18</figureCitation>
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<figureCitation captionStart="Figures 42–49" captionStartId="F13" captionText="Figures 42 - 49. Dorsal habitus, Carabus (Archiplectes) besleticus besleticus, Abkhazia, Mt. Birtzkha." pageId="4" pageNumber="25">, 42-49</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="25" type="reference_group">
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Neoplectes) satyrus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Neoplectes" subSpecies="besleticus">Carabus (Neoplectes) satyrus besleticus</taxonomicName>
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Kurnakov, 1972: 112 (
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<normalizedToken originalValue="“Vallée">"Vallee</normalizedToken>
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de la
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<normalizedToken originalValue="Besléta”">Besleta"</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName class="Insecta" family="Carabidae" genus="Neoplectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoplectes reitteri subsp. gaskoi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="reitteri" subSpecies="gaskoi">Neoplectes reitteri gaskoi</taxonomicName>
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Kenyeri, 1975: 113.
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) satyrus besleticus</taxonomicName>
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:
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<bibRefCitation author="Gottwald, J" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="23" pageNumber="44" pagination="278 - 314" refId="B5" refString="Gottwald, J, 1985. Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae). Acta Entomologica Bohemoslovaca 824: 278 - 314" title="Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae)." volume="824" year="1985">Gottwald 1985</bibRefCitation>
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: 310;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="Catalogue of Palaearctic Coleoptera" pageId="23" pageNumber="44" pagination="118 - 201" refId="B2" refString="Bousquet, Y, Brezina, B, Davies, A, Farkac, J, Smetana, A, 2003. Tribe Carabini Latreille, 1802. Catalogue of Palaearctic Coleoptera Vol. 1: 118 - 201" title="Tribe Carabini Latreille, 1802." volume="Vol. 1" year="2003">Bousquet et al. 2003</bibRefCitation>
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: 132;
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<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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2008
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</bibRefCitation>
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: 40.
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) satyrus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Tribax" subSpecies="besleticus">Carabus (Tribax) satyrus besleticus</taxonomicName>
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:
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<bibRefCitation author="Deuve, T" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="23" pageNumber="44" refId="B3" refString="Deuve, T, 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers, Sofia-Moscow" title="Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae)." year="2004">Deuve 2004</bibRefCitation>
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: 274.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="25" type="comparative material examined">
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<paragraph pageId="4" pageNumber="25">Comparative material examined.</paragraph>
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<paragraph pageId="4" pageNumber="25">
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176 specimens were examined (41 specimens measured, 11 male genitalia preparations studied): 59 males, 73 females, Abkhazia, Sukhum Distr. near Sukhum city, N slope of Mt. Birtzkha, 200 m, 26.IV-25.VI.2013, leg. D. Fominykh, A. Bondarenko (cFDD, cBAS, cSIA, cKVM); 1 female, Abkhazia, Sukhum Distr., vicinities of Village Abzhakva, left bank of River Basla, 80-90 m, box and beech forest site, 05-16.V.2013, leg. I. Solodovnikov, S. Solodovnikova, V. Kotsur, N. Pichugin (cSIA); 1 male, Abkhazia, Sukhum Distr., vicinities of Village Kveno-Linda, N slope of Mt. Birtskha, 120-140 m, box and beech forest site,
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<geoCoordinate degrees="43" direction="north" minutes="02" orientation="latitude" precision="1" seconds="49.81" value="43.04717">N43°02'49.81"</geoCoordinate>
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,
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<geoCoordinate degrees="041" direction="east" minutes="04" orientation="longitude" precision="1" seconds="01.23" value="41.06701">E041°04'01.23"</geoCoordinate>
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, 05-13.05.2012, leg. I. Solodovnikov (cSIA); 11 males, 10 females, Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova, V. Kotsur, S. Pavlyuchuk, N. Pichugin (cSIA, cKVM, cPSM, cPNYu); 4 males, 12 females, Abkhazia, Sukhum Distr., right bank of River Kelasur, E slope of Mt. Anykhapaara, beech, alder, chestnut forest site, 135 m,
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<geoCoordinate degrees="42" direction="north" minutes="59" orientation="latitude" precision="1" seconds="30.67" value="42.99185">42°59'30.67"N</geoCoordinate>
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/
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<geoCoordinate degrees="041" direction="east" minutes="05" orientation="longitude" precision="1" seconds="59.66" value="41.099903">041°05'59.66"E</geoCoordinate>
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, 11.V-08.VII.2014, leg. I. Solodovnikov, S. Solodovnikova, S. Pavlyuchuk (cSIA, cPSM); 2 males, 3 females, Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, right bank of River Basla, beech, linden, box, elder forest site, 160 m,
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<geoCoordinate degrees="43" direction="north" minutes="02" orientation="latitude" precision="1" seconds="57.59" value="43.04933">43°02'57.59"N</geoCoordinate>
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/
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<geoCoordinate degrees="041" direction="east" minutes="03" orientation="longitude" precision="1" seconds="48.67" value="41.06352">041°03'48.67"E</geoCoordinate>
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, 11.V-08.VII.2014, leg. I. Solodovnikov, S. Solodovnikova, S. Pavlyuchuk (cSIA, cPSM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25" type="description">
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<paragraph pageId="4" pageNumber="25">Description.</paragraph>
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<paragraph pageId="4" pageNumber="25">
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Large form, males 32.0-38.0 (34.7) mm, females 38.0-40.75 (39.0) mm long, robust. Underside black, dorsum normally with bright metallic luster, various colours of green, dark blue, bronze, crimson, violet, seldom black, with transitional color forms in males, females matte. Body massive, less slender than in
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
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.
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</paragraph>
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<paragraph lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25">
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Head not inflated. Pronotum of variable shape, from subcordate to transverse, lateral sides not to faintly sinuated. PW/PL = 1.26-1.53 (1.37) in males and 1.25-1.62 (1.46)
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<pageBreakToken pageId="5" pageNumber="26" start="start">in</pageBreakToken>
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females, base rather broad, PW/PB = 1.03-1.33 (1.17) in males and 1.11-1.31 (1.19) in females, hind angles distinctly protruding backwards and faintly sidewards, rounded apically. Median groove distinct, microsculpture forming transversal rugosity, gradually strengthening towards median groove. Elytra strongly elongate to ovate, more elongate in males, with inconspicuous depression in the middle in both males and females. EL/EW = 1.62-1.86 (1.75) in males and 1.64-1.86 (1.78) in females. Elytral sculpture nearly identical in males and females, forming faint and smoothed series of elongated links. The main morphometric measurements of the studied populations are given in Table
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="5" pageNumber="26" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="26">Lamella of aedeagus elongate, with a prominent hollow on the right side before the apex. Saccellus large, conical, smoothly rounded apically, paracellar lobe hardly developed, endophallus with symmetrical position of inner structures (when viewed frontally). Ostium lobe large, of blunted conical shape.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="26" type="differential diagnosis">
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<paragraph pageId="5" pageNumber="26">Differential diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="26">
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Habitually strongly resembles
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
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from which it differs in the less elongate and more rounded body shape, the different shape of pronotum, smoothed elytral sculpture with longer links, the different shapes of pronotum and male genitalia. The morphometric characters of this subspecies are illustrated in Figures
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="5" pageNumber="26">1</figureCitation>
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-
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="5" pageNumber="26">4</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="26" type="distribution">
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<paragraph pageId="5" pageNumber="26">Distribution.</paragraph>
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<paragraph pageId="5" pageNumber="26">Populates a low mountain forest belt in an interfluve of the River Gumista - East Gumista and Kelasur.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26" type="habitat">
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<paragraph pageId="5" pageNumber="26">Habitat.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26">
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Prefers beech and beech-chestnut forest sites admixed with box, rich in ground litter on karstic landforms at altitudes from 50 to 500 m. Activity of imago proceeds from April until July.
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) apollo subsp. phoebus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="apollo" subGenus="Archiplectes" subSpecies="phoebus">
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<pageBreakToken pageId="6" pageNumber="27" start="start">Carabus</pageBreakToken>
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(Archiplectes) apollo phoebus
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</taxonomicName>
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Kurnakov, 1962,
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<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. rugatus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="rugatus">Carabus (Sphodristocarabus) armeniacus rugatus</taxonomicName>
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Breuning, 1934 (natio
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<taxonomicName class="Insecta" family="Carabidae" form="novotnyorum" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="form">
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<emphasis italics="true" pageId="6" pageNumber="27">novotnyorum</emphasis>
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</taxonomicName>
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Mandl, 1975), and
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
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Motschulsky, 1844, occur together with this subspecies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |