treatments-xml/data/10/2A/9C/102A9C71FF80FFCEFF46FB28FA8FF2C6.xml

<document id="5109F4E917AD34E0BB22614AC5127533" ID-DOI="10.5281/zenodo.195474" ID-GBIF-Dataset="58b19e40-396d-4195-b753-7e810f666a8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="195474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460227646048" checkinUser="plazi" docAuthor="Barrio-Amorós, César L., Brewer-Carías, Javier Mesa Charles &amp; Mcdiarmid, Roy W." docDate="2010" docId="102A9C71FF80FFCEFF46FB28FA8FF2C6" docLanguage="en" docName="zt02483p044.pdf" docOrigin="Zootaxa 2483" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pristimantis muchimuk Barrio-Amorós, Brewer-Carías &amp; Mcdiarmid, 2010, sp. nov." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="EC13E409FF81FFCBFFD1FFAAFFD0F56C" masterDocTitle="A new Pristimantis (Anura, Terrarana, Strabomantidae) from Churi-tepui in the Chimanta massif, Venezuelan Guayana" masterLastPageNumber="44" masterPageNumber="35" pageNumber="36" updateTime="1698238697682" updateUser="plazi">
<mods:mods id="3E8E9403C0A67AD7DD8828FAD7A31582" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="92EE175BE7C181E26ED478AB4F15FB48">
<mods:title id="9EFC90CED954E7495B1D9F72764AEDC2">A new Pristimantis (Anura, Terrarana, Strabomantidae) from Churi-tepui in the Chimanta massif, Venezuelan Guayana</mods:title>
</mods:titleInfo>
<mods:name id="9AC820FE1A95CB4FFA25FD4A79B7AD7D" type="personal">
<mods:role id="22C3D867CBD89CE78AD6141D94006ABB">
<mods:roleTerm id="692818796119A7112B9E7AAEC32D1A6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AE263C352D9007D1CE9C0D651E3240F">Barrio-Amorós, César L.</mods:namePart>
</mods:name>
<mods:name id="AA8BE4CE6C81EC58AFFA33C5C26B4399" type="personal">
<mods:role id="2D5C936A18B4F55A91A0D546D8911867">
<mods:roleTerm id="D735488DEF2F4E84D577852756A80100">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ABFD4106D32F4F2543BB504381379143">Brewer-Carías, Javier Mesa Charles</mods:namePart>
</mods:name>
<mods:name id="AE0C87A5247D320CF0E014DAA4252706" type="personal">
<mods:role id="EA737FA6E0929A1013673030A359DF75">
<mods:roleTerm id="453150681473C04161B2B597183075A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0C143D2AB3D9A73FECBC36A9C456134">Mcdiarmid, Roy W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CD8540E37879BAFB2254E694A952E1C3">text</mods:typeOfResource>
<mods:relatedItem id="27006C5B58B324C7352921AFE54482E0" type="host">
<mods:titleInfo id="65EAA16E65F4D12A115C18D1C8B8098D">
<mods:title id="5775E4F2890D2F08718F17924D79AB61">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B6D2D8523DB754360ED22E9B9CAE521">
<mods:date id="A3FDDB842BF03C7CAA220CE8AC74452E">2010</mods:date>
<mods:detail id="733803E8D04676A0D92E8C6944FE0438" type="volume">
<mods:number id="EBC94156AFF084D760F5AF46A27C54F9">2483</mods:number>
</mods:detail>
<mods:extent id="1B5716F7BAE5DEB3DACB99A9FAD8EA09" unit="page">
<mods:start id="5E7710E9116177CD237E697C3B8A4EF8">35</mods:start>
<mods:end id="9F5AB6599973D3284E9041287C1E5845">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="397211A08DAC7E010599F10E89F91BDD">journal article</mods:classification>
<mods:identifier id="3DC30AE5FB1E7DEBEF1873707C459A3D" type="DOI">10.5281/zenodo.195474</mods:identifier>
<mods:identifier id="ECAF122C0007185B251C72BF8DED7E66" type="GBIF-Dataset">58b19e40-396d-4195-b753-7e810f666a8e</mods:identifier>
<mods:identifier id="3B4E9DCD9FDD3C0A998D5E7B7311212A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="23C0D2B92CA8BEC083062718FE684A9D" type="Zenodo-Dep">195474</mods:identifier>
</mods:mods>
<treatment id="102A9C71FF80FFCEFF46FB28FA8FF2C6" ID-DOI="http://doi.org/10.5281/zenodo.6207677" ID-GBIF-Taxon="119386819" ID-Zenodo-Dep="6207677" LSID="urn:lsid:plazi:treatment:102A9C71FF80FFCEFF46FB28FA8FF2C6" httpUri="http://treatment.plazi.org/id/102A9C71FF80FFCEFF46FB28FA8FF2C6" lastPageId="5" lastPageNumber="40" pageId="1" pageNumber="36">
<subSubSection id="D0997EECFF80FFCAFF46FB28FEE6F1D3" pageId="1" pageNumber="36" type="nomenclature">
<paragraph id="983C2D67FF80FFCAFF46FB28FDC8F1F0" blockId="1.[151,536,1154,1215]" box="[151,536,1154,1180]" pageId="1" pageNumber="36">
<heading id="C3749A0BFF80FFCAFF46FB28FDC8F1F0" bold="true" box="[151,536,1154,1180]" fontSize="11" level="1" pageId="1" pageNumber="36" reason="1">
<emphasis id="AAF7F175FF80FFCAFF46FB28FDC8F1F0" bold="true" box="[151,536,1154,1180]" pageId="1" pageNumber="36">
<taxonomicName id="5F8356E4FF80FFCAFF46FB28FE63F1F0" box="[151,435,1154,1180]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF80FFCAFF46FB28FE63F1F0" bold="true" box="[151,435,1154,1180]" italics="true" pageId="1" pageNumber="36">Pristimantis muchimuk</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1C44C0EFF80FFCAFE6BFB28FDC8F1F0" box="[442,536,1154,1180]" pageId="1" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="983C2D67FF80FFCAFF46FB0FFEE6F1D3" blockId="1.[151,536,1154,1215]" box="[151,310,1189,1215]" pageId="1" pageNumber="36">
(
<figureCitation id="00B831E2FF80FFCAFF4EFB0FFEFCF1D3" box="[159,300,1189,1215]" captionStart-0="FIGURE 1. A" captionStart-1="FIGURE 2. A" captionStart-2="FIGURE 3. A" captionStartId-0="3.[151,259,1759,1783]" captionStartId-1="4.[151,256,979,1003]" captionStartId-2="5.[151,256,1011,1035]" captionTargetBox-0="[188,1398,185,1695]" captionTargetBox-1="[190,1372,321,943]" captionTargetBox-2="[249,1301,369,970]" captionTargetId-0="figure@3.[188,1398,185,1735]" captionTargetId-1="figure@4.[177,1410,309,955]" captionTargetId-2="figure@5.[234,1353,346,986]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. A) Pristimantis muchimuk, sp. nov, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of Pristimantis muchimuk, sp. nov. Scale equals 5 mm." captionText-1="FIGURE 2. A) dorsal view and B) lateral view of the head of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." captionText-2="FIGURE 3. A) palm of right hand and B) sole of right foot, of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri-0="https://zenodo.org/record/195475/files/figure.png" httpUri-1="https://zenodo.org/record/195476/files/figure.png" httpUri-2="https://zenodo.org/record/195477/files/figure.png" pageId="1" pageNumber="36">Figures 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF80FFCAFF46FB5AFB24F066" box="[151,1268,1264,1290]" pageId="1" pageNumber="36" type="description">
<paragraph id="983C2D67FF80FFCAFF46FB5AFB24F066" blockId="1.[151,1437,1264,2036]" box="[151,1268,1264,1290]" pageId="1" pageNumber="36">
Common name in English: 
<taxonomicName id="5F8356E4FF80FFCAFE08FB5AFD8AF066" box="[473,602,1264,1290]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="42" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">Muchimuk</taxonomicName>
rain frog. Common name in Spanish: Ranita 
<taxonomicName id="5F8356E4FF80FFCAFBBFFB5AFB20F066" box="[1134,1264,1264,1290]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="muchimuk">muchimuk</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF80FFCAFF17FABFFB97F016" pageId="1" pageNumber="36" type="materials_examined">
<paragraph id="983C2D67FF80FFCAFF17FABFFB97F016" blockId="1.[151,1437,1264,2036]" pageId="1" pageNumber="36">
<typeStatus id="473893C5FF80FFCAFF17FABFFEE8F043" box="[198,312,1301,1327]" pageId="1" pageNumber="36" type="holotype">
<emphasis id="AAF7F175FF80FFCAFF17FABFFEE8F043" bold="true" box="[198,312,1301,1327]" pageId="1" pageNumber="36">Holotype</emphasis>
</typeStatus>
. 
<collectionCode id="FE92B5A2FF80FFCAFE9DFABFFE63F043" box="[332,435,1301,1327]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="1" pageNumber="36">MHNLS</collectionCode>
19652, an adult female from the base camp of the Muchimuk-Expedition 2009, collected by Igor Elorza, Javier Mesa and Charles Brewer-Carías on a step of the northern face of Churi-tepui, 
<geoCoordinate id="FDB74BA0FF80FFCAFF46FACAFE91F016" box="[151,321,1376,1402]" direction="north" orientation="latitude" pageId="1" pageNumber="36" precision="15" value="5.2791667">05º 16’ 45’’ N</geoCoordinate>
, 
<geoCoordinate id="FDB74BA0FF80FFCAFE9CFACAFE2BF016" box="[333,507,1376,1402]" direction="west" orientation="longitude" pageId="1" pageNumber="36" precision="15" value="-62.015556">62º 00´56’’ W</geoCoordinate>
, 2325 masl, Estado Bolívar, 
<collectingCountry id="E0946DF7FF80FFCAFC9BFACAFC13F016" box="[842,963,1376,1402]" name="Venezuela" pageId="1" pageNumber="36">Venezuela</collectingCountry>
(
<figureCitation id="00B831E2FF80FFCAFC02FACAFBEAF016" box="[979,1082,1376,1402]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,255,1070,1094]" captionStartId-1="7.[151,256,1819,1843]" captionTargetBox-0="[151,1435,197,1045]" captionTargetBox-1="[151,1434,645,1798]" captionTargetId-0="figure@6.[151,1436,197,1047]" captionTargetId-1="figure@7.[151,1436,645,1798]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Aerial view of the Churi tepui, picture taken from the northwest. The type locality appears as a white spot pointed by a white arrow. Photo: Charles Brewer-Carías." captionText-1="FIGURE 5. Distribution of Pristimantis muchimuk sp. nov. and relatives from the Guyanan Venezuela. White circles with numbers correspond to: 1: Pristimantis auricarens: Auyan-tepui. 2: P. a viu s and P. memorans: Tamacuari. 3: P. cantitans, P. pruinatus and P. yaviensis: Cerro Yavi. 4: P. guaiquinimensis, P. stegolepis and P. tepuiensis: Guaiquinimatepui. 5: P. marahuaka: Cerro Marahuaka. 6: P. marmoratus: base of Roraima-tepui. 7: P. m u c h i m u k: Churí-tepui in the Chimanta massif. 8: P. pulvinatus: Sierra de Lema. 9: P. sarisarinama: Sarisariñama-tepui. 10: P. yuruaniensis: Yuruanitepui. 11. P. zeuctotylus: base of Cerro Neblina. The distribution of P. v i l a r s i is in Barrio-Amorós &amp; Molina (2006)." httpUri-0="https://zenodo.org/record/195478/files/figure.png" httpUri-1="https://zenodo.org/record/195479/files/figure.png" pageId="1" pageNumber="36">Fig. 4–5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF80FFC9FF17FA2FFE77F2DE" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36" type="diagnosis">
<paragraph id="983C2D67FF80FFCAFF17FA2FFA87F2C6" blockId="1.[151,1437,1264,2036]" pageId="1" pageNumber="36">
<emphasis id="AAF7F175FF80FFCAFF17FA2FFE96F0F3" bold="true" box="[198,326,1413,1439]" pageId="1" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="5F8356E4FF80FFCAFE85FA2FFDBAF0F2" box="[340,618,1413,1438]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="muchimuk">
<emphasis id="AAF7F175FF80FFCAFE85FA2FFDBAF0F2" box="[340,618,1413,1438]" italics="true" pageId="1" pageNumber="36">Pristimantis muchimuk</emphasis>
</taxonomicName>
is a small (SVL 25.2 mm) species that we place in the diverse 
<taxonomicName id="5F8356E4FF80FFCAFF46FA01FE6EF0A8" box="[151,446,1451,1476]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="AAF7F175FF80FFCAFF46FA01FE6EF0A8" box="[151,446,1451,1476]" italics="true" pageId="1" pageNumber="36">Pristimantis unistrigatus</emphasis>
</taxonomicName>
group 
<emphasis id="AAF7F175FF80FFCAFDC5FA01FD86F0A8" box="[532,598,1451,1476]" italics="true" pageId="1" pageNumber="36">sensu</emphasis>
<bibRefCitation id="FC125096FF80FFCAFD8FFA00FC84F0A8" box="[606,852,1450,1476]" pageId="1" pageNumber="43" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New world direct-developing frogs (Anura: Terrarana). Molecular phylogeny, classification, biogeography and conservation. Zootaxa, 1737, 1 - 182." type="journal article">Hedges et al. (2008)</bibRefCitation>
, distinguished by Finger I shorter than Finger II, Toe V longer than Toe III, extending to the distal subarticular tubercle on Toe IV, cranial crests absent, and vomerine teeth present. 
<taxonomicName id="5F8356E4FF80FFCAFE6AFA5FFD03F362" box="[443,723,1525,1550]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="muchimuk">
<emphasis id="AAF7F175FF80FFCAFE6AFA5FFD03F362" box="[443,723,1525,1550]" italics="true" pageId="1" pageNumber="36">Pristimantis muchimuk</emphasis>
</taxonomicName>
is characterized by: (1) dorsal skin smooth to warty, warts smooth, low and flat (smooth in preservative), with a barely distinguishable middorsal raphe; ventral skin smooth; (2) tympanic annulus and membrane absent; (3) snout rounded in dorsal view, nearly truncate in profile; canthus rostralis rounded; (4) upper eyelid smooth, with one barely discernible tubercle on each eyelid; (5) choanae small, oval; vomerine dentigerous processes horizontal, small, posterior and medial to choanae, each bearing 3 teeth; tongue rounded posteriorly; (6) presence of vocal slits and nuptial pads not known; (7) Finger I shorter than Finger II; (8) fingers with lateral keels; (9) ulnar tubercles absent; (10) tarsal tubercles and calcars absent; (11) inner metatarsal tubercle oval, three times the size of the round outer metatarsal tubercle; (12) toes with lateral fringes; basal webbing between Toes III-IV-V; Toe V longer than Toe III; (13) in preservative, dorsal surfaces of body and limbs dark brown, almost black, with many small whithish spots; ventral surfaces dirty white, with brown melanophores concentrated on chin and throat. In life: dorsum black with many small yellow spots; flanks dark brown with pinkish white flecks; venter white.
</paragraph>
<paragraph id="983C2D67FF80FFC9FF17F81FFDCFF270" blockId="1.[151,1437,1264,2036]" lastBlockId="2.[151,1437,152,1970]" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
<emphasis id="AAF7F175FF80FFCAFF17F81FFD9EF2A3" bold="true" box="[198,590,1973,1999]" pageId="1" pageNumber="36">Comparison with other species.</emphasis>
<taxonomicName id="5F8356E4FF80FFCAFD84F81FFCB4F2A2" box="[597,868,1973,1998]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="36" phylum="Chordata" rank="species" species="muchimuk">
<emphasis id="AAF7F175FF80FFCAFD84F81FFCB4F2A2" box="[597,868,1973,1998]" italics="true" pageId="1" pageNumber="36">Pristimantis muchimuk</emphasis>
</taxonomicName>
(
<figureCitation id="00B831E2FF80FFCAFCA2F81FFC1EF2A3" box="[883,974,1973,1999]" captionStart="FIGURE 1. A" captionStartId="3.[151,259,1759,1783]" captionTargetBox="[188,1398,185,1695]" captionTargetId="figure@3.[188,1398,185,1735]" captionTargetPageId="3" captionText="FIGURE 1. A) Pristimantis muchimuk, sp. nov, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of Pristimantis muchimuk, sp. nov. Scale equals 5 mm." httpUri="https://zenodo.org/record/195475/files/figure.png" pageId="1" pageNumber="36">Fig. 1A</figureCitation>
) is unique among congeners inhabiting the highlands and mountain slopes in the Western 
<collectingCountry id="E0946DF7FF80FFCAFD25F870FC9AF298" box="[756,842,2010,2036]" name="Guyana" pageId="1" pageNumber="36">Guiana</collectingCountry>
Shield (
<collectingCountry id="E0946DF7FF80FFCAFC7FF870FBFBF298" box="[942,1067,2010,2036]" name="Venezuela" pageId="1" pageNumber="36">Venezuela</collectingCountry>
and 
<collectingCountry id="E0946DF7FF80FFCAFBB6F870FB15F298" box="[1127,1221,2010,2036]" name="Guyana" pageId="1" pageNumber="36">Guyana</collectingCountry>
) by the following combination of characters: tympanum absent, skin on dorsum and venter smooth, only one ill-defined tubercle on each upper eyelid, fingers and toes with well-developed lateral fringes, basal webbing between Toes III–V, and dorsum dark brown with small yellow and pink spots (in life), which become white in preservative. The following Guayanan 
<taxonomicName id="5F8356E4FF83FFC9FE76FEA2FD95F44D" box="[423,581,264,289]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="AAF7F175FF83FFC9FE76FEA2FD95F44D" box="[423,581,264,289]" italics="true" pageId="2" pageNumber="37">Pristimantis</emphasis>
</taxonomicName>
are compared with 
<emphasis id="AAF7F175FF83FFC9FC96FEA2FC36F44D" box="[839,998,264,289]" italics="true" pageId="2" pageNumber="37">P. m u c h i m u k</emphasis>
(characters of the new species in parentheses), as all are endemics from granitic or sandstone uplands or highlands (tepuis). 
<taxonomicName id="5F8356E4FF83FFC9FAD4FE87FDC5F400" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="auricarens">
<emphasis id="AAF7F175FF83FFC9FAD4FE87FEC7F407" italics="true" pageId="2" pageNumber="37">Pristimantis auricarens</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Auyantepui has tubercular dorsal skin (slightly warty to smooth) and areolate ventral skin (smooth), snout truncate to acutely rounded dorsally (rounded), rounded in profile (nearly truncate). 
<taxonomicName id="5F8356E4FF83FFC9FEB9FE37FCE5F4DB" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" box="[360,821,413,439]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="avius">
<emphasis id="AAF7F175FF83FFC9FEB9FE37FDEDF4DA" box="[360,573,413,438]" italics="true" pageId="2" pageNumber="37">Pristimantis avius</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Tamacuari, has tubercular dorsal skin (slightly warty to smooth), distinct tympanum (absent), no lateral fringes on fingers (present), and females are larger, up to 
<quantity id="5F7B8082FF83FFC9FF0AFE42FEFEF76E" box="[219,302,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="37" unit="mm" value="33.0">33 mm</quantity>
(
<quantity id="5F7B8082FF83FFC9FEEEFE42FE48F76E" box="[319,408,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="37" unit="mm" value="25.0">25 mm</quantity>
). 
<taxonomicName id="5F8356E4FF83FFC9FE7DFE42FC7AF76E" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" box="[428,938,488,514]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="cantitans">
<emphasis id="AAF7F175FF83FFC9FE7DFE42FD60F76D" box="[428,688,488,513]" italics="true" pageId="2" pageNumber="37">Pristimantis cantitans</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Cerro Yaví, has a visible tympanum (absent), upper eyelid with small warts (smooth, with only one barely distinguishable tubercle), small, low nonconical calcar tubercles (absent), lip bars present (absent), larger size, females up to 
<quantity id="5F7B8082FF83FFC9FB73FD98FB25F720" box="[1186,1269,562,588]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="37" unit="mm" value="45.0">45 mm</quantity>
(
<quantity id="5F7B8082FF83FFC9FAD4FD98FA8DF720" box="[1285,1373,562,588]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="37" unit="mm" value="25.0">25 mm</quantity>
), and an axillary tubercle (absent). 
<taxonomicName id="5F8356E4FF83FFC9FE27FDF2FBF0F71E" authority="Means &amp; Savage" authorityName="Means &amp; Savage" box="[502,1056,600,626]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="dendrobatoides">
<emphasis id="AAF7F175FF83FFC9FE27FDF2FC98F71D" box="[502,840,600,625]" italics="true" pageId="2" pageNumber="37">Pristimantis dendrobatoides</emphasis>
Means &amp; Savage
</taxonomicName>
, from Wokomung, 
<collectingCountry id="E0946DF7FF83FFC9FAD9FDF2FAB7F71E" box="[1288,1383,600,626]" name="Guyana" pageId="2" pageNumber="37">Guyana</collectingCountry>
, has verrucose dorsal skin (slightly warty to smooth), distinct tympanum (absent), black dorsum with several large red spots, ventral parts of hind limbs bright red in life (no red color). 
<taxonomicName id="5F8356E4FF83FFC9FC6FFD08FAE1F7D0" authority="Parker" authorityName="Parker" box="[958,1329,674,700]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="inguinalis">
<emphasis id="AAF7F175FF83FFC9FC6FFD08FB1AF7D7" box="[958,1226,674,699]" italics="true" pageId="2" pageNumber="37">Pristimantis inguinalis</emphasis>
(Parker)
</taxonomicName>
from the Guianas, has granular dorsal skin (slightly warty to smooth), tubercles on eyelids (absent), and a yelloworange ocellus in cloacal region (absent). 
<taxonomicName id="5F8356E4FF83FFC9FDAEFD47FBF8F66B" authority="Means &amp; Savage" authorityName="Means &amp; Savage" box="[639,1064,749,775]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="jester">
<emphasis id="AAF7F175FF83FFC9FDAEFD47FC87F66A" box="[639,855,749,774]" italics="true" pageId="2" pageNumber="37">Pristimantis jester</emphasis>
Means &amp; Savage
</taxonomicName>
, from Wokomung, 
<collectingCountry id="E0946DF7FF83FFC9FADAFD47FAB9F66B" box="[1291,1385,749,775]" name="Guyana" pageId="2" pageNumber="37">Guyana</collectingCountry>
, has a snout dorsally subovoid (rounded), and laterally rounded (nearly truncate); a concave canthus rostralis (rounded), no fringes on fingers or toes (present), the coloration include red on the flanks (no red). 
<taxonomicName id="5F8356E4FF83FFC9FF46FCF7FCD4F61B" authority="Fuentes &amp; Barrio-Amoros" authorityName="Fuentes &amp; Barrio-Amoros" box="[151,772,861,887]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="marahuaka">
<emphasis id="AAF7F175FF83FFC9FF46FCF7FE64F61A" box="[151,436,861,886]" italics="true" pageId="2" pageNumber="37">Pristimantis marahuaka</emphasis>
(Fuentes &amp; Barrio-Amorós)
</taxonomicName>
from Cerro 
<taxonomicName id="5F8356E4FF83FFC9FC49FCF7FBCCF61B" box="[920,1052,861,887]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="44" phylum="Chordata" rank="species" species="marahuaka">Marahuaka</taxonomicName>
is most similar morphologically (see 
<bibRefCitation id="FC125096FF83FFC9FF1FFC28FD9BF6F0" author="Fuentes" box="[206,587,898,924]" pageId="2" pageNumber="43" refString="Fuentes, O. &amp; Barrio-Amoros, C. L. (2004) A new Eleutherodactylus (Anura, Leptodactylidae) from Marahuaka tepui, Amazonas, Venezuela. Revista de la Academia Colombiana de Ciencias, 28, 285 - 290." type="journal article" year="2004">Fuentes &amp; Barrio-Amorós 2004</bibRefCitation>
); though it lacks webbing between toes (basal webbing present between toes III-V), subarticular tubercles are prominent (little notable), ventral skin areolate to granular (smooth), disc on finger II 2.4 times wider than adjacent phalanx (2.0), and different color, dorsum pale brown to yellowish brown in preservative, garnet brown in life with or without small silvery spots (dark brown with white spots); venter is dirty white or reticulated with brown (white). 
<taxonomicName id="5F8356E4FF83FFC9FC07FBB2FA4CF15E" authority="Boulenger" authorityName="Boulenger" box="[982,1436,1048,1074]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="AAF7F175FF83FFC9FC07FBB2FAD3F15D" box="[982,1283,1048,1073]" italics="true" pageId="2" pageNumber="37">Pristimantis marmoratus</emphasis>
(Boulenger)
</taxonomicName>
from lowlands, uplands and highlands of the 
<collectingCountry id="E0946DF7FF83FFC9FD6AFB97FCC1F13B" box="[699,785,1085,1111]" name="Guyana" pageId="2" pageNumber="37">Guiana</collectingCountry>
Shield, has a tubercular dorsal skin (slightly warty to smooth), tympanum distinct (absent), limbs barred (without bars), and a different habitat, rain to cloud forests in lowland and uplands up to 
<quantity id="5F7B8082FF83FFC9FE27FB22FD80F1CE" box="[502,592,1160,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="37" unit="m" value="1400.0">1400 m</quantity>
of the 
<collectingCountry id="E0946DF7FF83FFC9FD75FB22FD29F1CE" box="[676,761,1160,1186]" name="Guyana" pageId="2" pageNumber="37">Guiana</collectingCountry>
Shield (vs. non-gramineous tubiform meadows on rocky summit on a single tepui at 
<quantity id="5F7B8082FF83FFC9FE24FB07FD89F1AB" box="[501,601,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="2.325" pageId="2" pageNumber="37" unit="m" value="2325.0">2325 m</quantity>
). 
<taxonomicName id="5F8356E4FF83FFC9FDBEFB07FB4AF1AB" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" box="[623,1178,1197,1223]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="memorans">
<emphasis id="AAF7F175FF83FFC9FDBEFB07FC40F1AA" box="[623,912,1197,1222]" italics="true" pageId="2" pageNumber="37">Pristimantis memorans</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Tamacuari has tubercular dorsal skin (slightly warty to smooth) and areolate venter (smooth), tympanum distinct (absent), dark lip bars (absent), truncate finger discs (rounded). 
<taxonomicName id="5F8356E4FF83FFC9FCC2FB52FAC2F07E" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" box="[787,1298,1272,1298]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="pruinatus">
<emphasis id="AAF7F175FF83FFC9FCC2FB52FBCAF07D" box="[787,1050,1272,1297]" italics="true" pageId="2" pageNumber="37">Pristimantis pruinatus</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Cerro Yaví, has granular dorsal skin (slightly warty to smooth) and areolate venter (smooth), tympanum small but visible (absent), lateral keels on fingers absent (present), and small, non-conical calcar tubercles (absent). 
<taxonomicName id="5F8356E4FF83FFC9FF46FAC2FDFCF0EE" authority="Rivero" authorityName="Rivero" box="[151,556,1384,1410]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="pulvinatus">
<emphasis id="AAF7F175FF83FFC9FF46FAC2FE6AF0ED" box="[151,442,1384,1409]" italics="true" pageId="2" pageNumber="37">Pristimantis pulvinatus</emphasis>
(Rivero)
</taxonomicName>
from Sierra de Lema has many tubercles on dorsal surfaces (smooth), tympanum evident (absent), fingers and toes without lateral fringes (present), ulnar tubercles present (absent). 
<taxonomicName id="5F8356E4FF83FFC9FF46FA19FD56F0A0" authority="Means &amp; Savage" authorityName="Means &amp; Savage" box="[151,646,1458,1484]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="saltissimus">
<emphasis id="AAF7F175FF83FFC9FF46FA19FE60F0A0" box="[151,432,1459,1484]" italics="true" pageId="2" pageNumber="37">Pristimantis saltissimus</emphasis>
Means &amp; Savage
</taxonomicName>
, from Wokomung, 
<collectingCountry id="E0946DF7FF83FFC9FCBDFA18FC1BF0A0" box="[876,971,1458,1484]" name="Guyana" pageId="2" pageNumber="37">Guyana</collectingCountry>
, has snout profile subelliptical dorsally (rounded) and acuminate laterally (nearly truncate); canthus rostralis concave (rounded); fringes on fingers and toes absent (present) and no webbing between toes (present). 
<taxonomicName id="5F8356E4FF83FFC9FC42FA57FE94F350" authority="Barrio-Amoros &amp; Brewer-Carias" authorityName="Barrio-Amoros &amp; Brewer-Carias" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="sarisarinama">
<emphasis id="AAF7F175FF83FFC9FC42FA57FB15F37A" box="[915,1221,1533,1558]" italics="true" pageId="2" pageNumber="37">Pristimantis sarisarinama</emphasis>
Barrio-Amorós &amp; Brewer-Carías
</taxonomicName>
from Sarisariñama-tepui, has dorsal skin shagreneed (slightly warty to smooth), ventral skin areolate (smooth), evident tympanum (absent), fingers without lateral keels (present), toes without webbing (present) and keels (present). 
<taxonomicName id="5F8356E4FF83FFC9FE26F9C7FC28F3EB" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" box="[503,1016,1645,1671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="yaviensis">
<emphasis id="AAF7F175FF83FFC9FE26F9C7FD2CF3EA" box="[503,764,1645,1670]" italics="true" pageId="2" pageNumber="37">Pristimantis yaviensis</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
from Cerro Yaví, has flat tubercles on the upper eyelid (absent), and scattered warts on the dorsal skin (slightly warty to smooth, warts absent), fingers and toes lacking fringes (present). 
<taxonomicName id="5F8356E4FF83FFC9FD47F912FB7DF3BE" authority="Rodder &amp; Jungfer" authorityName="Rodder &amp; Jungfer" box="[662,1197,1720,1746]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="yuruaniensis">
<emphasis id="AAF7F175FF83FFC9FD47F912FC1BF3BD" box="[662,971,1720,1745]" italics="true" pageId="2" pageNumber="37">Pristimantis yuruaniensis</emphasis>
Rödder &amp; Jungfer
</taxonomicName>
from Yuruani-tepui lacks vomerine dentigerous processes (present), has a small but distinct tympanum (absent), and fingers and toes lack lateral fringes (present).
</paragraph>
<paragraph id="983C2D67FF83FFC9FF17F882FE77F2DE" blockId="2.[151,1437,152,1970]" pageId="2" pageNumber="37">
The other species of 
<taxonomicName id="5F8356E4FF83FFC9FE17F882FD85F22D" box="[454,597,1832,1857]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="AAF7F175FF83FFC9FE17F882FD85F22D" box="[454,597,1832,1857]" italics="true" pageId="2" pageNumber="37">Pristimantis</emphasis>
</taxonomicName>
in the 
<collectingCountry id="E0946DF7FF83FFC9FD7FF882FCD3F22E" box="[686,771,1832,1858]" name="Guyana" pageId="2" pageNumber="37">Guiana</collectingCountry>
region belong to the 
<taxonomicName id="5F8356E4FF83FFC9FBDAF882FA97F22D" box="[1035,1351,1832,1857]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="AAF7F175FF83FFC9FBDAF882FA97F22D" box="[1035,1351,1832,1857]" italics="true" pageId="2" pageNumber="37">Pristimantis conspicillatus</emphasis>
</taxonomicName>
Group (
<emphasis id="AAF7F175FF83FFC9FF70F8E7FF36F20A" box="[161,230,1869,1894]" italics="true" pageId="2" pageNumber="37">sensu</emphasis>
<bibRefCitation id="FC125096FF83FFC9FF3FF8E7FDF7F20B" author="Lynch" box="[238,551,1869,1895]" lastPageId="2" lastPageNumber="44" pageId="2" pageNumber="43" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in Western Ecuador: Systematics, Ecology and Biogeography. University of Kansas Natural History Museum, Special Publications, 23, 1 - 236." type="journal article" year="1997">Lynch &amp; Duellman 1997</bibRefCitation>
; 
<bibRefCitation id="FC125096FF83FFC9FDE4F8E7FCF3F20B" author="Hedges" box="[565,803,1869,1895]" pageId="2" pageNumber="43" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New world direct-developing frogs (Anura: Terrarana). Molecular phylogeny, classification, biogeography and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">Hedges et al. 2008</bibRefCitation>
). These are 
<emphasis id="AAF7F175FF83FFC9FC68F8E7FAB7F20A" box="[953,1383,1869,1894]" italics="true" pageId="2" pageNumber="37">
<taxonomicName id="5F8356E4FF83FFC9FC68F8E7FBA9F20A" box="[953,1145,1869,1894]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="chiastonotus">P. chiastonotus</taxonomicName>
, gutturalis, 
<taxonomicName id="5F8356E4FF83FFC9FAC1F8E7FAB3F20A" box="[1296,1379,1869,1894]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="vilarsi">vilarsi</taxonomicName>
,
</emphasis>
and 
<emphasis id="AAF7F175FF83FFC9FF46F8D9FEF1F2E0" box="[151,289,1907,1932]" italics="true" pageId="2" pageNumber="37">
<taxonomicName id="5F8356E4FF83FFC9FF46F8D9FECEF2E0" box="[151,286,1907,1932]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="37" phylum="Chordata" rank="species" species="zeuctotylus">zeuctotylus</taxonomicName>
,
</emphasis>
which are widely distributed throughout the lowlands. All have a long snout and the first finger longer than the second.
</paragraph>
</subSubSection>
<caption id="CCFC7DEFFF82FFC8FF46F975FC62F27A" httpUri="https://zenodo.org/record/195475/files/figure.png" pageId="3" pageNumber="38" targetBox="[188,1398,185,1695]" targetPageId="3">
<paragraph id="983C2D67FF82FFC8FF46F975FC62F27A" blockId="3.[151,1437,1759,1814]" pageId="3" pageNumber="38">
<emphasis id="AAF7F175FF82FFC8FF46F975FEF0F39B" bold="true" box="[151,288,1759,1783]" pageId="3" pageNumber="38">FIGURE 1.</emphasis>
A) 
<taxonomicName id="5F8356E4FF82FFC8FE9DF975FD80F39A" box="[332,592,1759,1782]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF82FFC8FE9DF975FD80F39A" box="[332,592,1759,1782]" italics="true" pageId="3" pageNumber="38">Pristimantis muchimuk</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B1C44C0EFF82FFC8FDB1F975FD63F39A" box="[608,691,1759,1782]" pageId="3" pageNumber="38" rank="species">sp. nov</taxonomicNameLabel>
, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of 
<taxonomicName id="5F8356E4FF82FFC8FE57F955FDADF27A" box="[390,637,1791,1814]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="38" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF82FFC8FE57F955FDADF27A" box="[390,637,1791,1814]" italics="true" pageId="3" pageNumber="38">Pristimantis muchimuk</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B1C44C0EFF82FFC8FD5BF955FD0CF27A" box="[650,732,1791,1814]" pageId="3" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
Scale equals 5 mm.
</paragraph>
</caption>
<subSubSection id="D0997EECFF82FFCFFF17F895FC89F29F" lastPageId="4" lastPageNumber="39" pageId="3" pageNumber="38" type="description">
<paragraph id="983C2D67FF82FFCFFF17F895FDCFF590" blockId="3.[151,1437,1855,2030]" lastBlockId="4.[151,1437,152,252]" lastPageId="4" lastPageNumber="39" pageId="3" pageNumber="38">
<emphasis id="AAF7F175FF82FFC8FF17F895FE8FF235" bold="true" box="[198,351,1855,1881]" pageId="3" pageNumber="38">Description.</emphasis>
Body slender (
<figureCitation id="00B831E2FF82FFC8FDC1F895FD83F235" box="[528,595,1855,1881]" captionStart="FIGURE 1. A" captionStartId="3.[151,259,1759,1783]" captionTargetBox="[188,1398,185,1695]" captionTargetId="figure@3.[188,1398,185,1735]" captionTargetPageId="3" captionText="FIGURE 1. A) Pristimantis muchimuk, sp. nov, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of Pristimantis muchimuk, sp. nov. Scale equals 5 mm." httpUri="https://zenodo.org/record/195475/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
B). Head slightly wider than body, slightly longer than wide; HL 39.2% of the SVL; rounded in dorsal view (
<figureCitation id="00B831E2FF82FFC8FD8DF8CEFD6FF212" box="[604,703,1892,1918]" captionStart="FIGURE 2. A" captionStartId="4.[151,256,979,1003]" captionTargetBox="[190,1372,321,943]" captionTargetId="figure@4.[177,1410,309,955]" captionTargetPageId="4" captionText="FIGURE 2. A) dorsal view and B) lateral view of the head of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri="https://zenodo.org/record/195476/files/figure.png" pageId="3" pageNumber="38">Fig. 2A</figureCitation>
), nearly truncate in profile (
<figureCitation id="00B831E2FF82FFC8FBC9F8CEFBB3F212" box="[1048,1123,1892,1918]" captionStart="FIGURE 2. A" captionStartId="4.[151,256,979,1003]" captionTargetBox="[190,1372,321,943]" captionTargetId="figure@4.[177,1410,309,955]" captionTargetPageId="4" captionText="FIGURE 2. A) dorsal view and B) lateral view of the head of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri="https://zenodo.org/record/195476/files/figure.png" pageId="3" pageNumber="38">Fig. 2</figureCitation>
B); nares not protuberant, directed laterally; canthus rostralis rounded, soft; loreal region concave; lips not protruding; one barely discernible tubercle on each eyelid, remainder of head without tubercles; interocular region barely wider tan upper eyelid width; temporal region almost vertical; supratympanic fold well defined, tympanic annulus and membrane absent. Choanae small, oval, not concealed by palatal shelf of maxillary arch, vomerine dentigerous processes barely distinguishable, transverse, posterior to level of choanae, bearing three teeth each; tongue rounded posteriorly.
</paragraph>
<caption id="CCFC7DEFFF85FFCFFF46FC79FEF6F166" httpUri="https://zenodo.org/record/195476/files/figure.png" pageId="4" pageNumber="39" targetBox="[190,1372,321,943]" targetPageId="4">
<paragraph id="983C2D67FF85FFCFFF46FC79FEF6F166" blockId="4.[151,1436,979,1034]" pageId="4" pageNumber="39">
<emphasis id="AAF7F175FF85FFCFFF46FC79FECDF687" bold="true" box="[151,285,979,1003]" pageId="4" pageNumber="39">FIGURE 2.</emphasis>
A) dorsal view and B) lateral view of the head of the holotype of 
<taxonomicName id="5F8356E4FF85FFCFFC28FC79FB26F686" box="[1017,1270,979,1002]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF85FFCFFC28FC79FB26F686" box="[1017,1270,979,1002]" italics="true" pageId="4" pageNumber="39">Pristimantis muchimuk</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B1C44C0EFF85FFCFFAD5FC79FA8AF686" box="[1284,1370,979,1002]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
Scale equals 2 mm.
</paragraph>
</caption>
<paragraph id="983C2D67FF85FFCFFF15FB99FD88F1F4" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">Skin on dorsum smooth, without tubercles; dorsal surfaces of limbs smooth, middorsal raphe barely distinguishable, anal sheath and tubercles in cloacal region absent; skin on flanks, throat, chest, belly and ventral surfaces of hind limbs smooth.</paragraph>
<paragraph id="983C2D67FF85FFCFFF15FB09FE13F014" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">
Hand (
<figureCitation id="00B831E2FF85FFCFFEC9FB09FEA9F1D1" box="[280,377,1187,1213]" captionStart="FIGURE 3. A" captionStartId="5.[151,256,1011,1035]" captionTargetBox="[249,1301,369,970]" captionTargetId="figure@5.[234,1353,346,986]" captionTargetPageId="5" captionText="FIGURE 3. A) palm of right hand and B) sole of right foot, of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri="https://zenodo.org/record/195477/files/figure.png" pageId="4" pageNumber="39">Fig. 3A</figureCitation>
) with a thenar tubercle ovoid; palmar tubercle deeply bifid; supernumerary tubercles present, large, low, non-protruding; subarticular tubercles round, little notable; fringes well developed on fingers; Finger I shorter than II (its length 87% of II); relative length of fingers III&gt;IV&gt;II&gt;I; finger discs expanded, rounded, on Fingers III and IV twice width of adjacent phalange; on Finger II 1.5 the width of adjacent phalange, and on Finger I slightly wider than adjacent phalange; all discs with ventral pads. Ulnar fold and tubercles absent.
</paragraph>
<paragraph id="983C2D67FF85FFCFFF15FA29FDD4F311" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">
Foot length 40.4% of SVL (
<figureCitation id="00B831E2FF85FFCFFDCFFA29FDB9F0F1" box="[542,617,1411,1437]" captionStart="FIGURE 3. A" captionStartId="5.[151,256,1011,1035]" captionTargetBox="[249,1301,369,970]" captionTargetId="figure@5.[234,1353,346,986]" captionTargetPageId="5" captionText="FIGURE 3. A) palm of right hand and B) sole of right foot, of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri="https://zenodo.org/record/195477/files/figure.png" pageId="4" pageNumber="39">Fig. 3</figureCitation>
B); calcar, tarsal tubercles, and tarsal fold absent. Inner metatarsal tubercle oval, 3 times size of round outer metatarsal tubercle; supernumerary tubercles present, numerous, small; subarticular tubercles round, little notable; toes with well-developed dermal fringes; basal webbing between Toes III-IV-V; relative length of the toes IV&gt;V&gt;III&gt;II&gt;I; discs on toes round, smaller than those on fingers; disc on Toe IV slightly smaller than disc on Finger III. When adpressed, tip of disc on Toe V slightly surpasses distal subarticular tubercle of Toe IV; tip of disc on Toe III slightly surpasses penultimate subarticular tubercle of Toe IV.
</paragraph>
<paragraph id="983C2D67FF85FFCFFF15F923FBABF254" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">
Color in preservative: dorsal surfaces of body and limbs uniform dark brown with profusely scattered small white spots; palmar surfaces of Fingers I and II dirty white, those of Fingers III and IV with many dark brown flecks; plantar surfaces of Toes I–III white, rest of plantar surfaces with dark brown flecks, darkest laterally; ventral surfaces dirty white with few brown flecks on belly and chest, and more profuse on throat. Iris gray; palpebral membrane with a profusion of melanophores inferiorly (
<figureCitation id="00B831E2FF85FFCFFBC0F8B4FB88F254" box="[1041,1112,1822,1848]" captionStart="FIGURE 2. A" captionStartId="4.[151,256,979,1003]" captionTargetBox="[190,1372,321,943]" captionTargetId="figure@4.[177,1410,309,955]" captionTargetPageId="4" captionText="FIGURE 2. A) dorsal view and B) lateral view of the head of the holotype of Pristimantis muchimuk, sp. nov. Scale equals 2 mm." httpUri="https://zenodo.org/record/195476/files/figure.png" pageId="4" pageNumber="39">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="983C2D67FF85FFCFFF15F8E9FD45F2C4" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">
Color in life (from a color photograph; 
<figureCitation id="00B831E2FF85FFCFFD48F8E9FD27F231" box="[665,759,1859,1885]" captionStart="FIGURE 1. A" captionStartId="3.[151,259,1759,1783]" captionTargetBox="[188,1398,185,1695]" captionTargetId="figure@3.[188,1398,185,1735]" captionTargetPageId="3" captionText="FIGURE 1. A) Pristimantis muchimuk, sp. nov, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of Pristimantis muchimuk, sp. nov. Scale equals 5 mm." httpUri="https://zenodo.org/record/195475/files/figure.png" pageId="4" pageNumber="39">Fig. 1A</figureCitation>
): dorsal ground color uniform reddish brown, with many small yellow, mostly round, spots dorsally and many small pinkish white spots on flanks and limbs. Venter white. Iris gray with fine black reticulation.
</paragraph>
<paragraph id="983C2D67FF85FFCFFF17F819FC89F29F" blockId="4.[151,1437,1075,2035]" pageId="4" pageNumber="39">
<emphasis id="AAF7F175FF85FFCFFF17F819FDEDF2A1" bold="true" box="[198,573,1971,1997]" pageId="4" pageNumber="39">
Measurements of the 
<typeStatus id="473893C5FF85FFCFFE03F819FDEDF2A1" box="[466,573,1971,1997]" pageId="4" pageNumber="39" type="holotype">holotype</typeStatus>
</emphasis>
. SVL: 25.2; ShL: 12.4; FL: 10.2; HW: 9.1; HL: 9.9; UEW: 2.4; IOD: 2.6; ED: 3.0.7; FD: 1.2; T4D: 1.1; ETS: 4.1; 1FiL: 2.9; 2Fil: 3.3.
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF84FFCEFF17FF3DFED4F44E" pageId="5" pageNumber="40" type="discussion">
<paragraph id="983C2D67FF84FFCEFF17FF3DFED4F44E" blockId="5.[151,1436,151,290]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF17FF3DFE90F5DD" bold="true" box="[198,320,151,177]" pageId="5" pageNumber="40">Remarks.</emphasis>
The 
<typeStatus id="473893C5FF84FFCEFE52FF32FE3AF5DE" box="[387,490,152,178]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
is slightly dehydrated (
<figureCitation id="00B831E2FF84FFCEFCDCFF32FC82F5DE" box="[781,850,152,178]" captionStart="FIGURE 1. A" captionStartId="3.[151,259,1759,1783]" captionTargetBox="[188,1398,185,1695]" captionTargetId="figure@3.[188,1398,185,1735]" captionTargetPageId="3" captionText="FIGURE 1. A) Pristimantis muchimuk, sp. nov, holotype in life. Photo: Javier Mesa. B) Dorsolateral view of the preserved holotype of Pristimantis muchimuk, sp. nov. Scale equals 5 mm." httpUri="https://zenodo.org/record/195475/files/figure.png" pageId="5" pageNumber="40">Fig. 1</figureCitation>
B) because it was preserved directly in alcohol, without fixing it in formalin. Some characters are still easily observed (fringes on fingers and toes, folds on forearms and tarsi), but should be compared with future living conspecific individuals or specimens fixed in formalin.
</paragraph>
</subSubSection>
<caption id="CCFC7DEFFF84FFCEFF46FC59FEF6F140" httpUri="https://zenodo.org/record/195477/files/figure.png" pageId="5" pageNumber="40" targetBox="[249,1301,369,970]" targetPageId="5">
<paragraph id="983C2D67FF84FFCEFF46FC59FEF6F140" blockId="5.[151,1436,1011,1068]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF46FC59FECCF167" bold="true" box="[151,284,1011,1035]" pageId="5" pageNumber="40">FIGURE 3.</emphasis>
A) palm of right hand and B) sole of right foot, of the holotype of 
<taxonomicName id="5F8356E4FF84FFCEFC2DFC5EFB27F167" box="[1020,1271,1012,1035]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF84FFCEFC2DFC5EFB27F167" box="[1020,1271,1012,1035]" italics="true" pageId="5" pageNumber="40">Pristimantis muchimuk</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B1C44C0EFF84FFCEFAD7FC5EFA8BF167" box="[1286,1371,1012,1035]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
Scale equals 2 mm.
</paragraph>
</caption>
<subSubSection id="D0997EECFF84FFCEFF17FBFFFE93F37C" pageId="5" pageNumber="40" type="distribution">
<paragraph id="983C2D67FF84FFCEFF17FBFFFEEFF016" blockId="5.[151,1437,1109,1962]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF17FBFFFEB7F103" bold="true" box="[198,359,1109,1135]" pageId="5" pageNumber="40">Distribution.</emphasis>
The species is known only from Churi-tepui (
<figureCitation id="00B831E2FF84FFCEFC74FBFFFC20F103" box="[933,1008,1109,1135]" captionStart="FIGURE 4" captionStartId="6.[151,255,1070,1094]" captionTargetBox="[151,1435,197,1045]" captionTargetId="figure@6.[151,1436,197,1047]" captionTargetPageId="6" captionText="FIGURE 4. Aerial view of the Churi tepui, picture taken from the northwest. The type locality appears as a white spot pointed by a white arrow. Photo: Charles Brewer-Carías." httpUri="https://zenodo.org/record/195478/files/figure.png" pageId="5" pageNumber="40">Fig. 4</figureCitation>
), one of the 12 tepuis forming the 
<taxonomicName id="5F8356E4FF84FFCEFF46FBD7FEDAF1FB" box="[151,266,1149,1175]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimanta</taxonomicName>
massif. The summit area of Churi-tepui is 47.5 km2, whereas the total summit area of 
<taxonomicName id="5F8356E4FF84FFCEFADFFBD7FA50F1FB" box="[1294,1408,1149,1175]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimanta</taxonomicName>
is 
<quantity id="5F7B8082FF84FFCEFF46FB0CFF3FF1AC" box="[151,239,1190,1216]" metricMagnitude="5" metricUnit="m" metricValue="6.2299999999999995" pageId="5" pageNumber="40" unit="km" value="623.0">623 km</quantity>
2, and the slope area is 
<quantity id="5F7B8082FF84FFCEFE2AFB0CFD83F1AC" box="[507,595,1190,1216]" metricMagnitude="5" metricUnit="m" metricValue="9.15" pageId="5" pageNumber="40" unit="km" value="915.0">915 km</quantity>
2 (
<bibRefCitation id="FC125096FF84FFCEFDBAFB0CFC1EF1AC" author="McDiarmid" box="[619,974,1190,1216]" pageId="5" pageNumber="44" refString="McDiarmid, R. W. &amp; Donnelly, A. (2005) The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World, 461 - 560. In: Donnelly, M. A., Crother, B. I., Guyer, C., Wake, M. H. &amp; White, M. E. (Eds.), Ecology and Evolution in the Tropics: A Herpetological Perspective, University of Chicago Press, Chicago, Illinois." type="book chapter" year="2005">McDiarmid &amp; Donnelly 2005</bibRefCitation>
). To date, 
<taxonomicName id="5F8356E4FF84FFCEFB96FB0CFA85F1D3" box="[1095,1365,1190,1215]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="muchimuk">
<emphasis id="AAF7F175FF84FFCEFB96FB0CFA85F1D3" box="[1095,1365,1190,1215]" italics="true" pageId="5" pageNumber="40">Pristimantis muchimuk</emphasis>
</taxonomicName>
is the only species of the genus described for the 
<taxonomicName id="5F8356E4FF84FFCEFD4AFB61FCDEF189" box="[667,782,1227,1253]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimanta</taxonomicName>
massif. The species 
<taxonomicName id="5F8356E4FF84FFCEFBD7FB61FB0CF188" box="[1030,1244,1227,1252]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="AAF7F175FF84FFCEFBD7FB61FB0CF188" box="[1030,1244,1227,1252]" italics="true" pageId="5" pageNumber="40">Eleutherodactylus</emphasis>
</taxonomicName>
sp. I mentioned by 
<bibRefCitation id="FC125096FF84FFCEFF6DFB5AFDFDF066" author="McDiarmid" box="[188,557,1264,1290]" pageId="5" pageNumber="44" refString="McDiarmid, R. W. &amp; Donnelly, A. (2005) The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World, 461 - 560. In: Donnelly, M. A., Crother, B. I., Guyer, C., Wake, M. H. &amp; White, M. E. (Eds.), Ecology and Evolution in the Tropics: A Herpetological Perspective, University of Chicago Press, Chicago, Illinois." type="book chapter" year="2005">McDiarmid &amp; Donnelly (2005)</bibRefCitation>
from Murey (= Eruoda) tepui in the 
<taxonomicName id="5F8356E4FF84FFCEFC0EFB5AFB81F066" box="[991,1105,1264,1290]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimanta</taxonomicName>
massif probably represent a different undescribed species (see Discussion). 
<figureCitation id="00B831E2FF84FFCEFD01FABCFCC6F05C" box="[720,790,1302,1328]" captionStart="FIGURE 5" captionStartId="7.[151,256,1819,1843]" captionTargetBox="[151,1434,645,1798]" captionTargetId="figure@7.[151,1436,645,1798]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Pristimantis muchimuk sp. nov. and relatives from the Guyanan Venezuela. White circles with numbers correspond to: 1: Pristimantis auricarens: Auyan-tepui. 2: P. a viu s and P. memorans: Tamacuari. 3: P. cantitans, P. pruinatus and P. yaviensis: Cerro Yavi. 4: P. guaiquinimensis, P. stegolepis and P. tepuiensis: Guaiquinimatepui. 5: P. marahuaka: Cerro Marahuaka. 6: P. marmoratus: base of Roraima-tepui. 7: P. m u c h i m u k: Churí-tepui in the Chimanta massif. 8: P. pulvinatus: Sierra de Lema. 9: P. sarisarinama: Sarisariñama-tepui. 10: P. yuruaniensis: Yuruanitepui. 11. P. zeuctotylus: base of Cerro Neblina. The distribution of P. v i l a r s i is in Barrio-Amorós &amp; Molina (2006)." httpUri="https://zenodo.org/record/195479/files/figure.png" pageId="5" pageNumber="40">Fig. 5</figureCitation>
shows the distribution of 
<emphasis id="AAF7F175FF84FFCEFB81FABCFB35F043" box="[1104,1253,1302,1327]" italics="true" pageId="5" pageNumber="40">P. m u c h i m u k</emphasis>
and the rest of Venezuelan species south of the Orinoco river except 
<taxonomicName id="5F8356E4FF84FFCEFCF6FA91FC42F038" box="[807,914,1339,1364]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="vilarsi">
<emphasis id="AAF7F175FF84FFCEFCF6FA91FC42F038" box="[807,914,1339,1364]" italics="true" pageId="5" pageNumber="40">P. vilarsi</emphasis>
</taxonomicName>
(see distribution map in 
<bibRefCitation id="FC125096FF84FFCEFB14FA91FEE3F016" author="Barrio-Amoros" pageId="5" pageNumber="43" refString="Barrio-Amoros, C. L., Rivas, G. &amp; Kaiser, H. (2006) New species of Colostethus (Anura: Dendrobatidae) from the Peninsula de Paria, Venezuela. Journal of Herpetology, 40, 371 - 377." type="journal article" year="2006">Barrio-Amorós &amp; Molina 2006</bibRefCitation>
).
</paragraph>
<paragraph id="983C2D67FF84FFCEFF17FA2FFE93F37C" blockId="5.[151,1437,1109,1962]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF17FA2FFEA5F0F3" bold="true" box="[198,373,1413,1439]" pageId="5" pageNumber="40">Conservation.</emphasis>
We recommend that the UICN status of 
<emphasis id="AAF7F175FF84FFCEFC54FA2CFBCDF0F3" box="[901,1053,1414,1439]" italics="true" pageId="5" pageNumber="40">P. m u c h i m u k</emphasis>
must be Data Deficient (DD) (according to 
<bibRefCitation id="FC125096FF84FFCEFEEEFA01FDC4F0A9" author="Stuart" box="[319,532,1451,1477]" pageId="5" pageNumber="44" refString="Stuart, S. N., Hoffman, M., Chanson, J., Cox, N., Berridge, R., Ramani, P. &amp; Young, B. (Eds.) (2008) Threatened Amphibians of the World. Lynx Editions, Barcelona, Spain; IUCN, Gland. Switzerland; Conservation International, Arlington, Virginia, U. S. A." type="book" year="2008">Stuart et al. 2008</bibRefCitation>
). Much research about the population status must be done in the 
<taxonomicName id="5F8356E4FF84FFCEFAF9FA01FA4DF0A9" box="[1320,1437,1451,1477]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimantá</taxonomicName>
massif, but the new species seems to be rare, as we only found one specimen in four visits and Gorzula (1992) never found it.
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF84FFCEFF17F9B0FDEDF279" pageId="5" pageNumber="40" type="description">
<paragraph id="983C2D67FF84FFCEFF17F9B0FDEDF279" blockId="5.[151,1437,1109,1962]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF17F9B0FDE7F358" bold="true" box="[198,567,1562,1588]" pageId="5" pageNumber="40">Habitat and Natural History.</emphasis>
The single specimen was found after rain, during the day, on a leaf of a fallen 
<taxonomicName id="5F8356E4FF84FFCEFF34F9EBFD19F336" box="[229,713,1600,1626]" class="Amphibia" family="Strabomantidae" genus="Brochinnia" kingdom="Animalia" order="Anura" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="hechtioides">
<emphasis id="AAF7F175FF84FFCEFF34F9EBFDD1F336" box="[229,513,1601,1626]" italics="true" pageId="5" pageNumber="40">Brochinnia hechtioides</emphasis>
(Bromeliaceae)
</taxonomicName>
. The frog was apparently disturbed by human activity, and consequently visible during daylight. The habitat is known as “Non-Gramineous tubiform meadows” (sensu 
<bibRefCitation id="FC125096FF84FFCEFF46F921FEFBF3C9" author="Huber" box="[151,299,1675,1701]" pageId="5" pageNumber="43" refString="Huber, O. (1995) Vegetation, 97 - 160. In: Berry, P. E. Holst, B. K. &amp; Yatskievych, K. (Eds). Flora of the Venezuelan Guayana. Vol. 1. Timber Press, Portland, Oregon." type="book chapter" year="1995">Huber 1995</bibRefCitation>
) on a flat swampy surface, with a height of no more than one meter, and predominance of 
<emphasis id="AAF7F175FF84FFCEFF46F91BFAD6F3A6" box="[151,1286,1713,1738]" italics="true" pageId="5" pageNumber="40">
<taxonomicName id="5F8356E4FF84FFCEFF46F91BFD90F3A6" authority="Orectanthe" authorityName="Orectanthe" box="[151,576,1713,1738]" class="Gastropoda" family="Cancellariidae" genus="Brocchinia" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="hechtioides">Brocchinia hechtioides, Orectanthe</taxonomicName>
ptaritepuyana, Heliamphora heterodoxa, Pterozonium sp.,
</emphasis>
and 
<emphasis id="AAF7F175FF84FFCEFAEBF91BFF68F383" italics="true" pageId="5" pageNumber="40">
<taxonomicName id="5F8356E4FF84FFCEFAEBF91BFA4DF3A6" box="[1338,1437,1713,1738]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="genus">Drosera</taxonomicName>
sp.
</emphasis>
In the vertical crevices nearby there is a dwarf forest of the dwarf tree 
<taxonomicName id="5F8356E4FF84FFCEFC25F97CFB01F383" box="[1012,1233,1750,1775]" class="Magnoliopsida" family="Bonnetiaceae" genus="Bonnetia" kingdom="Animalia" order="Malpighiales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="roraimae">
<emphasis id="AAF7F175FF84FFCEFC25F97CFB01F383" box="[1012,1233,1750,1775]" italics="true" pageId="5" pageNumber="40">Bonnetia roraimae</emphasis>
</taxonomicName>
with intermittent streams running in the rainy season.
</paragraph>
</subSubSection>
<subSubSection id="D0997EECFF84FFCEFF17F88AFA8FF2C6" pageId="5" pageNumber="40" type="etymology">
<paragraph id="983C2D67FF84FFCEFF17F88AFA8FF2C6" blockId="5.[151,1437,1109,1962]" pageId="5" pageNumber="40">
<emphasis id="AAF7F175FF84FFCEFF17F88AFE9BF256" bold="true" box="[198,331,1824,1850]" pageId="5" pageNumber="40">Etymology</emphasis>
. 
<taxonomicName id="5F8356E4FF84FFCEFE8AF88BFE06F256" box="[347,470,1825,1850]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">
<emphasis id="AAF7F175FF84FFCEFE8AF88BFE06F256" box="[347,470,1825,1850]" italics="true" pageId="5" pageNumber="40">Muchimuk</emphasis>
</taxonomicName>
refers to a demon in the mythology of the indigenous Pemon people. The demon has avian form, like a giant raptor, and takes humans and other beasts for food. It inhabits the summits of the 
<taxonomicName id="5F8356E4FF84FFCEFF46F8C1FEDCF2E9" box="[151,268,1899,1925]" class="Reptilia" family="Colubridae" genus="Thamnodynastes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="35" phylum="Chordata" rank="species" species="chimanta">Chimantá</taxonomicName>
and Tramén tepuis. 
<taxonomicName id="5F8356E4FF84FFCEFDD5F8C1FD57F2E9" box="[516,647,1899,1925]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="muchimuk" status="sp. nov.">Muchimuk</taxonomicName>
is also the name of the Expedition made in 
<date id="EC3D0BA7FF84FFCEFB4EF8C1FACAF2E9" box="[1183,1306,1899,1925]" pageId="5" pageNumber="40" value="2009-05">May 2009</date>
to explore different galleries of the Charles Brewer cave system. The specific name is used as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>