154 lines
17 KiB
XML
154 lines
17 KiB
XML
<document ID-CLB-Dataset="269604" ID-DOI="http://dx.doi.org/10.3897/vz.73.e101372" ID-GBIF-Dataset="e5fee896-cf8b-412f-a286-2bd90981a414" ID-Pensoft-Pub="2625-8498-73-717" ID-Pensoft-UUID="318B657A15AB5708B3C35FC1A82B4945" ID-ZooBank="8F3D5EDA2F184E5CA53E2F7741FF1339" ModsDocID="2625-8498-73-717" checkinTime="1695893367890" checkinUser="pensoft" docAuthor="Szyndlar, Zbigniew & Georgalis, Georgios L." docDate="2023" docId="0FFDD39B00A2F26C42F3C6CE4079DFDA" docLanguage="en" docName="VertZool 73: 717-886" docOrigin="Vertebrate Zoology 73" docPubDate="2023-09-27" docSource="http://dx.doi.org/10.3897/vz.73.e101372" docTitle="Leiopython Hubrecht 1879" docType="treatment" docVersion="4" id="318B657A15AB5708B3C35FC1A82B4945" lastPageNumber="717" masterDocId="318B657A15AB5708B3C35FC1A82B4945" masterDocTitle="An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column" masterLastPageNumber="886" masterPageNumber="717" pageNumber="717" updateTime="1732872861233" updateUser="ExternalLinkService">
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<mods:mods id="51D1F0E7587568B4C869C1B4EC30A715" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8796FA488934A1BFA086CD4A404E4A49">
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<mods:title id="61F8F1EC9AB5CF4C02CD4FCD9A72D337">An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column</mods:title>
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<mods:name id="79A28C3699F714A1CC2F9117D60F0CAF" type="personal">
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<mods:namePart id="E70AA3EFEE7EF498F96BC34163E83F49">Szyndlar, Zbigniew</mods:namePart>
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<mods:affiliation id="482F02F763AA2854294964429CB6F38C">Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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<mods:roleTerm id="FB556A363D931380298D659567FACA5F">Author</mods:roleTerm>
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<mods:namePart id="CC628B81460A7CC01DB5A2D95124A6A3">Georgalis, Georgios L.</mods:namePart>
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<mods:nameIdentifier id="220EF9FD10B9B2F01661224F9DBD2BA5" type="ORCID">https://orcid.org/0000-0001-7759-6146</mods:nameIdentifier>
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<mods:affiliation id="788988DD8AC1CD4EBE0DE5A97D3772F3">Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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<mods:nameIdentifier id="C5D406E73106EEED4B8E83FFD250EDD5" type="email">dimetrodon82@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="7E67746C23A24BBD6ADC12462E1A6D15">text</mods:typeOfResource>
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<mods:titleInfo id="B5034A49D8552DAFE704C5A002FC608B">
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<mods:title id="7CCAD3273A058E009A751C08779653C0">Vertebrate Zoology</mods:title>
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<mods:part id="DEE1E9698282C865A621E76BAB356812">
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<mods:date id="0D96219247935B215C00A146A3EBB793">2023</mods:date>
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<mods:number id="F352FED8036DF6A37018157386B768A8">2023-09-27</mods:number>
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<mods:detail id="6BB6A93CD71AD9EC704D9AC15B67E8CE" type="volume">
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<mods:number id="615DA611C8A66D5721CA5E58D038E124">73</mods:number>
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<mods:extent id="E7EE1C46FE9D9D5171FCA3C11F6F17EE" unit="page">
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<mods:start id="B3EBD95AB868991E3E4A151E5F1BAE35">717</mods:start>
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<mods:end id="C1D6E4C44416A76EA9F7757E9073A773">886</mods:end>
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<mods:url id="61F24E944FDFBFFDB78523DDF9C1E607">http://dx.doi.org/10.3897/vz.73.e101372</mods:url>
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<mods:classification id="27FBA351A007ECA8650568E00BD84EE0">journal article</mods:classification>
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<mods:identifier id="6C07C6F74F3674BF24ED57B243675B10" type="DOI">http://dx.doi.org/10.3897/vz.73.e101372</mods:identifier>
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<mods:identifier id="9AEE57DA88F2E39E5C9276930D9F0C62" type="Pensoft-Pub">2625-8498-73-717</mods:identifier>
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</mods:mods>
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<treatment id="0FFDD39B00A2F26C42F3C6CE4079DFDA" ID-GBIF-Taxon="213623378" LSID="urn:lsid:plazi:treatment:0FFDD39B00A2F26C42F3C6CE4079DFDA" httpUri="http://treatment.plazi.org/id/0FFDD39B00A2F26C42F3C6CE4079DFDA" lastPageNumber="717" pageId="0" pageNumber="717" scope_family="Pythonidae" scope_infraOrder="Alethinophidia" scope_superFamily="Pythonoidea">
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<subSubSection id="ED19C6E06E2312117F330AEDCC19E9D9" pageId="0" pageNumber="717" type="nomenclature">
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<paragraph id="C05D348B7C47491EEB3B3380B5F8172F" pageId="0" pageNumber="717">
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<taxonomicName id="1B1A2FFCE83053BFB72F8185CCB43ECA" ID-CoL="87GSB" ID-ENA="3144915" authority="Hubrecht, 1879" authorityName="Hubrecht" authorityYear="1879" family="Pythonidae" genus="Leiopython" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leiopython" order="Squamata" pageId="0" pageNumber="717" rank="genus">Leiopython Hubrecht, 1879</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="E13C825DD3DC7BB82992A712963BB0F5" pageId="0" pageNumber="717" type="materials_examined">
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<paragraph id="518D05FB4EB471B22ED136598D686EC4" pageId="0" pageNumber="717">Material examined.</paragraph>
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<paragraph id="13DC9AF5BA9E29DE86FF1FE4D100A205" pageId="0" pageNumber="717">
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<taxonomicName id="9E34E45D65CE33DA6ED363B69CDAC0BC" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="24B4CAF0D2D765A0CE24237F10C78080" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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(Peters & Doria, 1878) (SMF
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<collectingCountry id="90D4E8A7424289EF3DC27BBD4181899E" name="Philippines">PH</collectingCountry>
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50).
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</paragraph>
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</subSubSection>
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<subSubSection id="0BEE0837D52D26FC866F01F79A0EC358" pageId="0" pageNumber="717" type="description">
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<paragraph id="3BA589D0472253025767592F4BCE4C86" pageId="0" pageNumber="717">Description (Figs 173-174).</paragraph>
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<paragraph id="1231F47F324DEFCE314CE09EDB7E6195" pageId="0" pageNumber="717">
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<emphasis id="FC31BBB6101B49DAB255D52C21B231F7" bold="true" pageId="0" pageNumber="717">
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<emphasis id="D87B28B3D34E3C1A8848688CF04AED70" italics="true" pageId="0" pageNumber="717">Trunk vertebrae</emphasis>
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.
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</emphasis>
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There is much similarity with
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<taxonomicName id="BDEDCFA726C81C85EA48974775FF09B6" authorityName="Gray" authorityYear="1849" family="Pythonidae" genus="Simalia" kingdom="Animalia" lsidName="Simalia" order="Squamata" pageId="0" pageNumber="717" rank="genus">
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<emphasis id="8F6DE0F7903B062118806CCB16CAEAFF" italics="true" pageId="0" pageNumber="717">Simalia</emphasis>
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</taxonomicName>
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described above. Centrum much shorter than wide; cotyle and condyle orbicular or moderately flattened; neural arch moderately vaulted; neural spine relatively short and its posterodorsal edge posteriorly inclined; hypapophyses disappear after V 50; haemal keel in succeeding vertebrae rather broad.
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</paragraph>
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<caption id="51EE1459FDF4B9AE97C348C3BBF6A6AA" doi="10.3897/vz.73.e101372.figure173" httpUri="https://binary.pensoft.net/fig/913064" pageId="0" pageNumber="717" start="Figure 173" startId="F173">
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<paragraph id="E83CF04E87297F2F52581A1DEEA041AB" pageId="0" pageNumber="717">
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<emphasis id="983109F1101B5E7DCE2B7DB1158ECE18" bold="true" pageId="0" pageNumber="717">Figure 173.</emphasis>
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<taxonomicName id="6304B7BD5F7DEE202D8397381257C69C" class="Reptilia" family="Pythonidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="family">Pythonidae</taxonomicName>
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:
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<taxonomicName id="C7BAECBDE1B341A3378CA3A3497DC9C6" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="BC1C50FA27BD7EB79FEC849C81AF6217" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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(SMF PH 50), trunk vertebra.
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</paragraph>
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</caption>
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<caption id="B4ACC968CDBAFEB90FB9D8D3D77DAD79" doi="10.3897/vz.73.e101372.figure174" httpUri="https://binary.pensoft.net/fig/913065" pageId="0" pageNumber="717" start="Figure 174" startId="F174">
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<paragraph id="F28255A8CAB360F4B280A68BD9373D15" pageId="0" pageNumber="717">
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<emphasis id="49093D09FD8CF9563B793E5AF5EF7C0B" bold="true" pageId="0" pageNumber="717">Figure 174.</emphasis>
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<taxonomicName id="87CFD278ADA3B032A93A4B1FC3D86943" class="Reptilia" family="Pythonidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="717" phylum="Chordata" rank="family">Pythonidae</taxonomicName>
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:
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<taxonomicName id="73E9ABF041CED9423D2C20616D2ED019" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="F7D757CF4E7052AB289F3189C09A7A12" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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(SMF PH 50), posteriormost trunk, cloacal, and anterior caudal vertebrae.
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</paragraph>
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</caption>
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<paragraph id="B7067DD0E48266DDD3BB843F4B385C06" pageId="0" pageNumber="717">
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<emphasis id="76E40DF598B0574E2ECBF6EBE9ECF9C6" bold="true" pageId="0" pageNumber="717">
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<emphasis id="8B970F6D1D04D079FFA9D4E148B0E614" italics="true" pageId="0" pageNumber="717">Trunk</emphasis>
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/
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<emphasis id="CE63CEE797FB47784C60843DA057C936" italics="true" pageId="0" pageNumber="717">caudal transition</emphasis>
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</emphasis>
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. The subcentral structures of this vertebral region are more or less similar to that of
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<taxonomicName id="D0C0912F40CA3CC1B657DE63F64A2D93" authorityName="Gray" authorityYear="1849" family="Pythonidae" genus="Simalia" kingdom="Animalia" lsidName="Simalia" order="Squamata" pageId="0" pageNumber="717" rank="genus">
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<emphasis id="EE45FFCD0A694A8135FB6046920627D2" italics="true" pageId="0" pageNumber="717">Simalia</emphasis>
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</taxonomicName>
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described above. The posteriormost trunk vertebrae develop a very short hypapophysis. Cloacal vertebrae possess a grooved hypapophysis. This grooved hypapophysis develops even more in anterior caudal vertebrae, being reminiscent of
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<normalizedToken id="26F2357BD020333352DAAFEC1E643CDD" originalValue="“quasi-haemapophyses”">"quasi-haemapophyses"</normalizedToken>
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, but normally shaped haemapophyses commence at the level of C 6, and subsequently continue throughout the tail. The posteriormost vertebrae are fused.
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</paragraph>
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<paragraph id="8593592650CE2565E167CAAB5D87372A" pageId="0" pageNumber="717">
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No vertebrae of
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<taxonomicName id="1E69297EDC33FF3DDEF1AD826D7958FE" authorityName="Hubrecht" authorityYear="1879" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython" order="Squamata" pageId="0" pageNumber="717" rank="genus">
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<emphasis id="262C9A2DDCC17CE5D8B2FD6817FEF770" italics="true" pageId="0" pageNumber="717">Leiopython</emphasis>
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</taxonomicName>
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had so far been figured but additional observations can be gleaned from the short description of the caudal vertebral patterns by
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<bibRefCitation id="3E207BFC6C0CB582563718E244061629" DOI="https://doi.org/10.2307/1562691" author="McDowell, SB" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="717" pagination="1 - 79" refId="B379" refString="McDowell, SB, 1975. A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae. Journal of Herpetology 9: 1 - 79, DOI: https://doi.org/10.2307/1562691" title="A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae." url="https://doi.org/10.2307/1562691" volume="9" year="1975">McDowell (1975)</bibRefCitation>
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, who personally studied a skeleton of
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<taxonomicName id="1D7FA156DF2134507A0A1025F9F2FD36" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="172A03DFF7A408C8F778A486E15B68C3" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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. According to that author, the anteriormost three caudal vertebrae ("3 vertebrae with simple transverse processes" in his writings) possessed a haemal keel ("free, triangular haemal bones on the ventral surface of the centra" in his writings), while the succeeding 65 caudal vertebrae possessed typically paired haemapophyses, followed by six fused posteriormost caudal vertebrae (
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<bibRefCitation id="B9D059DA0902B0AC0B1C113E9198FD0F" DOI="https://doi.org/10.2307/1562691" author="McDowell, SB" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="717" pagination="1 - 79" refId="B379" refString="McDowell, SB, 1975. A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae. Journal of Herpetology 9: 1 - 79, DOI: https://doi.org/10.2307/1562691" title="A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae." url="https://doi.org/10.2307/1562691" volume="9" year="1975">McDowell 1975</bibRefCitation>
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: 43).
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</paragraph>
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<paragraph id="BBDB337B537E36065D85BB3A38B34ACE" pageId="0" pageNumber="717">
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<emphasis id="9CE0AF9BA3845A33F0FFF4B7F6FBB121" bold="true" pageId="0" pageNumber="717">
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<emphasis id="20188B4DCA606A19D1EF4E6AC701964B" italics="true" pageId="0" pageNumber="717">Number of vertebrae</emphasis>
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.
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</emphasis>
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<taxonomicName id="B41A2DC67187787BFEE9399A0176FE6C" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="441773A097FEAC2411884C8C0E631965" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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(SMF PH 50): 350 (279+5+66 [including a final fusion]).
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</paragraph>
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<paragraph id="93ADA0C48146762D12AF806B279E281D" pageId="0" pageNumber="717">
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Data from literature (all for
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<taxonomicName id="7D68316C70A07630778CE98631AC49C4" authorityName="Peters & Doria" authorityYear="1878" baseAuthorityName="W.C.H.Peters & Doria" baseAuthorityYear="1878" family="Pythonidae" genus="Leiopython" kingdom="Animalia" lsidName="Leiopython albertisii" order="Squamata" pageId="0" pageNumber="717" rank="species" species="albertisii">
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<emphasis id="372B1CE48D5F106FEB84D1F153EE7E7F" italics="true" pageId="0" pageNumber="717">Leiopython albertisii</emphasis>
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</taxonomicName>
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): 273 trunk vertebrae plus 5 cloacal vertebrae plus 77 caudal vertebrae (posteriormost 6 caudal vertebrae are fused) (
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<bibRefCitation id="22D3011CD9B0EAD69D9B059969FEB9EB" DOI="https://doi.org/10.2307/1562691" author="McDowell, SB" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="717" pagination="1 - 79" refId="B379" refString="McDowell, SB, 1975. A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae. Journal of Herpetology 9: 1 - 79, DOI: https://doi.org/10.2307/1562691" title="A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae." url="https://doi.org/10.2307/1562691" volume="9" year="1975">McDowell 1975</bibRefCitation>
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); 347 vertebrae in total (
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<bibRefCitation id="D8FB5B639C202F3A6ABE1A622B6881F1" author="Parmley, D" journalOrPublisher="Georgia Journal of Science" pageId="0" pageNumber="717" pagination="220 - 232" refId="B433" refString="Parmley, D, Reed, HW, 2003. Size and age class estimates of North American Eocene palaeopheid snakes. Georgia Journal of Science 61: 220 - 232" title="Size and age class estimates of North American Eocene palaeopheid snakes." volume="61" year="2003">Parmley and Reed 2003</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |