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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.509.9506" ID-GBIF-Dataset="66954afc-cfb8-414b-8682-210fe363f991" ID-PMC="PMC4493346" ID-Pensoft-Pub="1313-2970-509-123" ID-PubMed="26175606" ID-ZBK="2664655ECBF6456BBC3DFE805E29D745" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-509-123" ModsDocOrigin="ZooKeys 509" ModsDocTitle="The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O.morio (Coleoptera, Carabidae, Omphreini)" checkinTime="1451244225243" checkinUser="pensoft" docAuthor="Curcic, Srecko, Sciaky, Riccardo, Antic, Dragan &amp; Vesovic, Nikola" docDate="2015" docId="0FAC35B0F4124D67B40D6339914A1764" docLanguage="en" docName="ZooKeys 509: 123-139" docOrigin="ZooKeys 509" docSource="http://dx.doi.org/10.3897/zookeys.509.9506" docTitle="Omphreus (Omphreus) morio subsp. durmitorensis Curcic &amp; Sciaky, ssp. n." docType="treatment" docUuid="BA83B21E-917D-4674-ACC1-B40D65FE121E" docUuidSource="ZooBank" docVersion="5" lastPageNumber="131" masterDocId="FFB2FFED4E115B1FFFA4042CFFE98E09" masterDocTitle="The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O. morio (Coleoptera, Carabidae, Omphreini)" masterLastPageNumber="139" masterPageNumber="123" pageNumber="128" updateTime="1668160421974" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O. morio (Coleoptera, Carabidae, Omphreini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Curcic, Srecko</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sciaky, Riccardo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Antic, Dragan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vesovic, Nikola</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>509</mods:number>
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<mods:start>123</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.509.9506</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.509.9506</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-509-123</mods:identifier>
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<treatment ID-GBIF-Taxon="152060424" LSID="urn:lsid:zoobank.org:act:BA83B21E-917D-4674-ACC1-B40D65FE121E" httpUri="http://treatment.plazi.org/id/0FAC35B0F4124D67B40D6339914A1764" lastPageId="8" lastPageNumber="131" pageId="5" pageNumber="128">
<subSubSection pageId="5" pageNumber="128" type="multiple">
<paragraph pageId="5" pageNumber="128">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="128" type="nomenclature">
<paragraph pageId="5" pageNumber="128">
<taxonomicName LSID="http://zoobank.org/BA83B21E-917D-4674-ACC1-B40D65FE121E" authority="Curcic &amp; Sciaky" class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="5" pageNumber="128" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">
Omphreus (Omphreus) morio durmitorensis
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; Sciaky
</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="128">ssp. n.</taxonomicNameLabel>
Figures 9-13
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="129" type="material examined">
<paragraph pageId="6" pageNumber="129">
<pageBreakToken pageId="6" pageNumber="129" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="6" pageNumber="129">
Holotype male labeled as follows: &quot;northwestern Montenegro, Mt. Durmitor, Sedlo pass, 2,100-2,200 m a.s.l., near
<normalizedToken originalValue="Žabljak">Zabljak</normalizedToken>
, 28.
<normalizedToken originalValue="VI">VI-</normalizedToken>
17.VII.2014, from pitfall traps, leg. S.
<normalizedToken originalValue="Ćurčić”">Curcic&quot;</normalizedToken>
(white label, printed) / Holotypus
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; R. Sciaky det. 2014&quot; (red label, printed) (IZFB). Paratypes: one female, same data as for holotype (IZFB); one male labeled as follows: &quot;northwestern Montenegro, Mt. Durmitor, Sedlo pass, 2,100 m a.s.l., 28-29.VI.2014, from pitfall traps, leg. S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; D.
<normalizedToken originalValue="Antić”">Antic&quot;</normalizedToken>
(IZFB); one male labeled as follows: &quot;northwestern Montenegro, Mt. Durmitor,
<normalizedToken originalValue="Žabljak">Zabljak</normalizedToken>
, 1,950 m a.s.l., VIII.2006, leg. P. Zanandrea&quot; (CRS); one female labeled as follows: &quot;northwestern Montenegro, Mt. Durmitor, Sedlo pass, 2,200 m a.s.l., 07.VIII.2007, leg. F. Sandel&quot; (CRS). All paratypes are labeled with white, printed locality labels and with red printed labels &quot;Paratypus
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; R. Sciaky det. 2014&quot;.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="130" pageId="6" pageNumber="129" type="description">
<paragraph pageId="6" pageNumber="129">Description.</paragraph>
<paragraph pageId="6" pageNumber="129">Size large: TL: R 17.73-18.91 mm (M 18.13 mm). Body elongate; elytra ovate (Figure 9). Body color black, mouthparts, apical antennomeres, and tarsi black-brownish. Tegument shiny, except slightly matt elytra.</paragraph>
<caption pageId="6" pageNumber="129">
<paragraph pageId="6" pageNumber="129">
Figures 9-13.
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. from northwestern Montenegro, Mt. Durmitor, Sedlo pass, 2,100-2,200 m a.s.l., near
<normalizedToken originalValue="Žabljak">Zabljak</normalizedToken>
. 9 holotype male, habitus (dorsal view) 10 holotype male, aedeagus (dorsal view) 11 holotype male, aedeagus (lateral view) 12 holotype male, abdominal sternite IX (urite) 13 paratype female, genitalia. Scale bars 5 mm (9) and 1 mm (10-13).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="129">Head rounded, somewhat elongated [HW/HL: M 0.95 (R 0.93-0.99)], shorter and narrower (HW/PW: M 0.695) than pronotum (Figure 9). Head beyond eye level somewhat constricted. Labrum broad, medially rounded, carrying four setae. Epistome huge, concave anteriorly, with two setae. Both vertex and occiput wrinkled. Frontal foveae deepened and long. Gula bisetose. Mandibles elongated, sickle-formed, broadened basally. Labial palpomere 1 short, without setae. Labial palpomeres 2 and 3 longer. Both the labial palpomere 3 and maxillar palpomere 3 broadened distally and densely pubescent. AL: M 10.38 mm (R 10.07-10.64 mm). Antennae pubescent from antennomere 4. AL/TL: M 0.57 (R 0.54-0.60). Antennomere 1 club-like, sharply widened distally, with a few long setae distally, somewhat shorter than the following three antennomeres combined. Antennomere 2 slightly shorter than antennomere 3.</paragraph>
<paragraph pageId="6" pageNumber="129">Pronotum sub-campaniform, elongate, PW/PL: M 0.91 (R 0.90-0.92). Fore angles somewhat prominent, rounded, hind angles obtuse, somewhat rounded (Figure 9). Lateral margins well developed, thickened, arcuate anteriorly, then sinuate and narrowing posteriorly. Anterior pronotal margin somewhat concave, while the base strongly concave. Pronotum widest in front of the fore third. Lateral furrows narrow and shallow, with four anterior setae, one median seta, and one posterior seta each. Median furrow long and deep. Basal foveae deep and long, slightly shorter than half of pronotum length. Pronotal disc somewhat convex proximally.</paragraph>
<paragraph pageId="7" pageNumber="130">
<pageBreakToken pageId="7" pageNumber="130" start="start">Elytra</pageBreakToken>
ovate, relatively long, arcuate laterally, EW/EL: M 0.56 (R 0.53-0.58) (Figure 9). EL/EW: M 1.79. EW: M 5.94 mm in males, 6.12 mm in females. Elytral striae shallow, weakly punctate, points somewhat deeper basally; elytral intervals flattened. Scutellum large, triangular; scutellar stria present, but without scutellar punctures. Elytra widest slightly below the middle; one setiferous puncture in interval 7 basally (close to stria 6), four or five setiferous punctures in interval 7 medially (close to stria 7), and two setiferous punctures situated on stria 7 apically on each elytron; the punctures deep and large. Shoulders obtusely rounded. Umbilicate series regular, with the setae densely distributed. Elytral disc somewhat convex distally.
</paragraph>
<paragraph pageId="7" pageNumber="130">Protarsomeres 1 and 2 widened in males. Metacoxae long and rounded. Tarsal claws elongated, glabrous, without teeth (Figure 9).</paragraph>
<paragraph pageId="7" pageNumber="130">Aedeagus long, median lobe strongly widened sub-apically in dorsal view (Figure 10), while arcuate, moderately widened and with a shallow concavity in the sub-apical part in lateral view (Figure 11), with a straight short rounded triangular apex (Figures 10 and 11). Parameres wide, the right being much huger (Figure 10). Basal bulb narrow and elongated (Figures 10 and 11).</paragraph>
<paragraph pageId="7" pageNumber="130">Male abdominal sternite IX (urite) large, sub-triangular (Figure 12).</paragraph>
<paragraph pageId="7" pageNumber="130">Both gonocoxites and gonosubcoxites IX as presented in Figure 13. Gonocoxites IX wide and elongated, somewhat curved, gradually narrowing distally, each with a pointed apex, basally joined with massive rounded gonosubcoxites IX.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="131" pageId="7" pageNumber="130" type="differential diagnosis and remarks">
<paragraph pageId="7" pageNumber="130">Differential diagnosis and remarks.</paragraph>
<paragraph pageId="7" pageNumber="130">
The new subspecies is compared here with the morphologically nearest subspecies of
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="species" species="morio" subGenus="Omphreus">Omphreus (Omphreus) morio</taxonomicName>
. These are
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
, and
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. In all these subspecies the first antennomere is club-like distally, somewhat shorter than the following three antennomeres combined (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
,
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
).
</paragraph>
<paragraph pageId="7" pageNumber="130">
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
in the shape of the elytra (arcuate laterally vs. somewhat rounded laterally), the elytra length/width ratio (M 1.79 vs. around 1.66), and the body length (R 17.73-18.91 mm vs. 16-18 mm) (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
,
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
). The former taxon is distributed on Mt. Durmitor (northwestern Montenegro), while the latter inhabits Mt.
<normalizedToken originalValue="Visočica">Visocica</normalizedToken>
(central Bosnia and Herzegovina) (the type locality) (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
).
<bibRefCitation author="Apfelbeck, V" journalOrPublisher="R. Friedlaender und Sohn, Berlin" pageId="12" pageNumber="135" title="Die Kaeferfauna der Balkanhalbinsel, mit Beruecksichtigung Klein-Asiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea" year="1904">Apfelbeck (1904)</bibRefCitation>
and
<bibRefCitation author="Drovenik, B" editor="Nonveiller, G" journalOrPublisher="The Montenegrin Academy of Sciences and Arts, Titograd" pageId="12" pageNumber="135" pagination="185 - 227" title="Cicindelidae in Carabidae (Insecta, Coleoptera)." volumeTitle="The Fauna of Durmitor" year="1984">Drovenik (1984)</bibRefCitation>
recorded the presence of an
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="genus">Omphreus</taxonomicName>
on Mt. Durmitor reported as
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
. While the former author recorded it for the mountain without precise localities, the latter author found it in several different sites (Crno Jezero Lake surroundings, forests below Mali
<normalizedToken originalValue="Štulac">Stulac</normalizedToken>
, Zminje Jezero Lake surroundings, Veliki
<normalizedToken originalValue="Štulac">Stulac</normalizedToken>
,
<normalizedToken originalValue="Ališnica">Alisnica</normalizedToken>
,
<normalizedToken originalValue="Međed">Međed</normalizedToken>
, Velika Karlica, Savin Kuk, Lokvice, Bobotov Kuk, Dobri Do, Sedlo, Todorov Do,
<normalizedToken originalValue="Prutaš">Prutas</normalizedToken>
,
<normalizedToken originalValue="Škrčka">Skrcka</normalizedToken>
Jezera Lakes surroundings, Bolj, forest below
<normalizedToken originalValue="Ćurevac">Curevac</normalizedToken>
, Nagorje, and
<normalizedToken originalValue="Šljeme">Sljeme</normalizedToken>
) (
<bibRefCitation author="Apfelbeck, V" journalOrPublisher="R. Friedlaender und Sohn, Berlin" pageId="12" pageNumber="135" title="Die Kaeferfauna der Balkanhalbinsel, mit Beruecksichtigung Klein-Asiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea" year="1904">Apfelbeck 1904</bibRefCitation>
,
<bibRefCitation author="Drovenik, B" editor="Nonveiller, G" journalOrPublisher="The Montenegrin Academy of Sciences and Arts, Titograd" pageId="12" pageNumber="135" pagination="185 - 227" title="Cicindelidae in Carabidae (Insecta, Coleoptera)." volumeTitle="The Fauna of Durmitor" year="1984">Drovenik 1984</bibRefCitation>
). On the basis of the material collected from Mt. Durmitor (at Sedlo and
<normalizedToken originalValue="Žabljak">Zabljak</normalizedToken>
sites) loaned by a few colleagues, as well as the one collected by the first and third author (S.
<normalizedToken originalValue="Ć">C</normalizedToken>
. and D.A.), we have concluded that the taxon from Mt. Durmitor actually represents a new subspecies,
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n.
</paragraph>
<paragraph pageId="8" pageNumber="131">
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">
<pageBreakToken pageId="8" pageNumber="131" start="start">Omphreus</pageBreakToken>
(Omphreus) morio durmitorensis
</taxonomicName>
ssp. n. differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
in the shape of the elytra (elongate, arcuate laterally vs. short, rounded laterally), the elytra length/width ratio (M 1.79 vs. around 1.50), and the body length (R 17.73-18.91 mm vs. 15-16 mm) (
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
). Furthermore, they inhabit quite distant areas:
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. is found in northwestern Montenegro (Mt. Durmitor), while
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
lives in southwestern Serbia (Mt. Murtenica and Mt. Zlatibor) (
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
).
</paragraph>
<paragraph pageId="8" pageNumber="131">
The diagnostic differences between
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. and
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. are presented in the differential diagnosis of the latter.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="131" type="etymology">
<paragraph pageId="8" pageNumber="131">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="131">The new subspecies is named after Mt. Durmitor, its terra typica.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="131" type="distribution">
<paragraph pageId="8" pageNumber="131">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="131">So far known only from the type locality, Mt. Durmitor, northwestern Montenegro.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="131" type="habitat">
<paragraph pageId="8" pageNumber="131">Habitat.</paragraph>
<paragraph pageId="8" pageNumber="131">The subspecies prefers high-altitude habitats (1,950-2,200 m a.s.l.) on Mt. Durmitor in northwestern Montenegro. The type series of the new subspecies was collected by pitfall traps filled with alcoholic vinegar, placed at different sites on the border between alpine meadows and rocks, up to a timberline on Mt. Durmitor.</paragraph>
</subSubSection>
</treatment>
</document>