167 lines
16 KiB
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167 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.699.12031" ID-GBIF-Dataset="dbf6bb9f-7ea4-4fa3-83ee-f58263427e39" ID-PMC="PMC5673864" ID-Pensoft-Pub="1313-2970-699-1" ID-PubMed="29200927" ID-ZBK="0F92401F3F7C4896AD9D72BC84348C7D" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-699-1" ModsDocOrigin="ZooKeys 699" ModsDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" checkinTime="1505836577638" checkinUser="pensoft" docAuthor="Cherman, Mariana Alejandra, Mise, Kleber Makoto, Moron †, Miguel Angel, Vaz-de-Mello, Fernando Z. & Almeida, Lucia Massutti de" docDate="2017" docId="0F96384A1202DAF3A719B513E4D60478" docLanguage="en" docName="ZooKeys 699: 1-120" docOrigin="ZooKeys 699" docSource="http://dx.doi.org/10.3897/zookeys.699.12031" docTitle="Liogenys rufocastanea Moser 1921" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="FFD9636FB945C41DFF8AB133FFFA9D76" masterDocTitle="A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="51" updateTime="1668164813467" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cherman, Mariana Alejandra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mise, Kleber Makoto</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moron †, Miguel Angel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vaz-de-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Almeida, Lucia Massutti de</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>699</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>120</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.699.12031</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.699.12031</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-699-1</mods:identifier>
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<mods:identifier type="ZBK">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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<mods:identifier type="ZooBank">0F92401F3F7C4896AD9D72BC84348C7D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134311197" LSID="urn:lsid:plazi:treatment:0F96384A1202DAF3A719B513E4D60478" httpUri="http://treatment.plazi.org/id/0F96384A1202DAF3A719B513E4D60478" lastPageId="52" lastPageNumber="52" pageId="50" pageNumber="51">
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<subSubSection pageId="50" pageNumber="51" type="nomenclature">
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<paragraph pageId="50" pageNumber="51">
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<taxonomicName authority="Moser, 1921" authorityName="Moser" authorityYear="1921" class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastanea" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastanea">Liogenys rufocastanea Moser, 1921</taxonomicName>
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Fig. 70
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="reference_group">
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<paragraph pageId="50" pageNumber="51">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastaneus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastaneus">Liogenys rufocastaneus</taxonomicName>
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Moser, 1918: 103 (orig. desc.);
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<bibRefCitation author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="103" pageNumber="104" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 228 (check.);
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<bibRefCitation author="Frey, G" journalOrPublisher="Frey" pageId="105" pageNumber="106" pagination="36 - 64" title="Betimmumgstabelle und Revision der Gattung Liogenys Guerrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologischen Arbeiten aus dem Museum G." volume="20" year="1969">Frey 1969</bibRefCitation>
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: 42 (key);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" pagination="1 - 458" title="A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)." url="https://doi.org/10.11646/zootaxa.211.1.1" volume="211" year="2003">Evans 2003</bibRefCitation>
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: 214 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://digitalcommons.unl.edu/entomologypapers/2" year="2005">Evans and Smith 2005</bibRefCitation>
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: 178 (check.);
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<bibRefCitation author="Evans, AV" journalOrPublisher="Zootaxa" pageId="104" pageNumber="105" url="http://museum.unl.edu/research/entomology/SSSA/nwmelos.htm" year="2009">Evans and Smith 2009</bibRefCitation>
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: 182 (check.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="type material">
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<paragraph pageId="50" pageNumber="51">Type material.</paragraph>
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<paragraph pageId="50" pageNumber="51">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastaneus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastaneus">Liogenys rufocastaneus</taxonomicName>
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male syntype (ZMHB): [white printed]
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<normalizedToken originalValue="“Paraguay”">"Paraguay"</normalizedToken>
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, [white handwritten] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="rufocastaneus" pageId="50" pageNumber="51" rank="species" species="rufocastaneus">rufocastaneus</taxonomicName>
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/Mos/Type m#.", [light red printed]
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<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
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, [white printed] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="rufocastaneus" pageId="50" pageNumber="51" rank="species" species="rufocastaneus">rufocastaneus</taxonomicName>
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/Mos.". Genitalia mounted. This type is here designated the lectotype: [white, outlined in red, printed] "LECTOTYPE/
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastaneus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastaneus">Liogenys rufocastaneus</taxonomicName>
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/Moser, 1921/des. M. A. Cherman 2016". A female syntype (ZMHB): [white printed]
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<normalizedToken originalValue="“Brasilien”">"Brasilien"</normalizedToken>
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, [white handwritten] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="rufocastaneus" pageId="50" pageNumber="51" rank="species" species="rufocastaneus">rufocastaneus</taxonomicName>
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/Mos/Type f#." [light red printed]
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<normalizedToken originalValue="“Typus”">"Typus"</normalizedToken>
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, [white printed] "
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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/
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<taxonomicName lsidName="rufocastaneus" pageId="50" pageNumber="51" rank="species" species="rufocastaneus">rufocastaneus</taxonomicName>
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/Mos." These syntype are here designated as paralectotype, with the label: "PARALECTOTYPE/
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastaneus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastaneus">Liogenys rufocastaneus</taxonomicName>
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/ Moser, 1921/ des. M. A. Cherman 2016".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="type material">
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<paragraph pageId="50" pageNumber="51">Non-type material.</paragraph>
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<paragraph pageId="50" pageNumber="51">without locality and date, Kraatz, 1 ex. (SDEI); PARAGUAY. Without date, Fry, 1 ex. (NHMB); Without date, Kraatz, 9 ex. (SDEI). BRAZIL. Without date, Kraatz, 7 ex. (SDEI).</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="diagnosis">
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<paragraph pageId="50" pageNumber="51">Diagnosis.</paragraph>
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<paragraph pageId="50" pageNumber="51">Body, elytra and pronotum brown; elongate; clypeus weakly emarginate; clypeus and frons almost coplanar; outer sides of anterior teeth follow the lateral margin of clypeus; clypeal lateral margin straight; pronotal posterior corners rounded; mesotibia cylindrical in cross section; pygidium flat and wide; pygidial width exceeding distance between spiracles of propygidium; pygidial disc bristled only on apex, punctures very sparse; pygidial apex quadrate; males genitalia, basal region of parameres wider than the parameres together at its maximum width; apex ornamented and emarginate laterally forming two projections at each side (Fig. 70F), subapical angles sharp, truncated and slightly divergent apically.</paragraph>
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<caption pageId="50" pageNumber="51">
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<paragraph pageId="50" pageNumber="51">
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Figure 70.
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastanea" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rufocastanea">Liogenys rufocastanea</taxonomicName>
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Moser. A Dorsal view B Lateral view C Frontal view D Clypeus and pronotum E Pygidium F Parameres, dorsal view G Parameres, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="51" pageNumber="52" type="redescription">
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<paragraph pageId="51" pageNumber="52">
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<pageBreakToken pageId="51" pageNumber="52" start="start">Redescription</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="51" pageNumber="52">
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Length: 8.0-9.0 mm; width: 4.1-4.2 mm. Brown. Head: distance between eyes nearly twice the width of one eye in males, wider in females; frons shorter than clypeus; clypeus and frons almost coplanar; anterior margin of clypeus slightly emarginate, emargination shallow, rounded and wide; outer sides of anterior teeth follow the lateral margin of clypeus; clypeal lateral margin straight; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width twice width of apex; fovea deep, extending past the transverse midline of the palpomere; labium transversely carinated, as wide as it is long; antenna 10-articulated, lamellae lighter in color and longer than flagellum. Thorax: anterior margin of pronotum slightly produced medially; maximum length of pronotum exceeding the length of tarsomeres I, II and III together, disc glabrous, punctures very sparse and coarse; pronotal posterior corners rounded; proepisternum with short bristles; mesosternum scaly; sides of metasternum with sparse scales anteriorly, few long bristles on the anterior margin; distance between meso- and metacoxae up to twice longer than the metacoxa; scutellum ogival, sparse and coarse punctures at the sides of the base. Elytra: shiny, glabrous, uniform brown; elytra more than three times longer than the pronotum; elytral suture and elytron unicolored, distinctly elevated; all four elytral ridges barely noticeable. Legs: procoxa, sparse scales on infra-carinal, outer surface up to inner margins of femur, smooth at 12
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; three protibial teeth, middle and apical equal in size, the three teeth equally spaced; protibial inner apical spur present; mesofemural disc setose, with a row of long bristles on the anterior margin; mesotibia cylindrical in cross section, disc finely sculptured, two mesotibial transverse carinae, the apical one incomplete in males; basal apophysis of metacoxa produced beyond the outer margin of trochanter, disc scaly; inner margin of metatibia carinated towards apex, apical inner surface setose; metatibial disc finely sculptured; metatibial transverse carina present posteriorly; metatibial apical spurs of different lengths, the longest shorter than the diameter of the tibial apex; protarsomere II long; in males pro- and mesotarsomeres I to IV enlarged, protarsomeres wider than the mesotarsomeres, twice as wide as metatarsi; basal metatarsomere twice shorter and slightly wider than tarsomere II; claw bifid, symmetrical, superior tooth longer and narrower than the inferior; distance between teeth shorter than the inferior tooth. Abdomen: ventrites bristled sparsely on disc and sides; propygidium visible, glabrous or scarcely bristled; pygidium flat, sub-trapezoidal, wide, as longer as wide, pygidial width exceeding distance between spiracles of propygidium; pygidial disc bristled only on apex; pygidial apex quadrate. Parameres: basal region wide and somewhat swollen, very short, parameral split at 1/3; ventral sides of base projected laterally; parameral inner margins straight; apex ornated, emarginate laterally forming two projections at each side, the basal one sharp, apical one truncated and slightly divergent (Fig. 70F). In lateral view parameres straight almost coplanar with basal region (Fig. 70G).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="51" pageNumber="52" type="type locality">
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<paragraph pageId="51" pageNumber="52">Type locality.</paragraph>
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<paragraph pageId="51" pageNumber="52">PARAGUAY.</paragraph>
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</subSubSection>
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<subSubSection pageId="51" pageNumber="52" type="distribution">
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<paragraph pageId="51" pageNumber="52">Geographical distribution.</paragraph>
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<paragraph pageId="51" pageNumber="52">BRAZIL; PARAGUAY.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="remarks">
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<paragraph pageId="51" pageNumber="52">Remarks.</paragraph>
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<paragraph lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys rufocastanea" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufocastanea">Liogenys rufocastanea</taxonomicName>
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resembles
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Pacuvia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacuvia castanea" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="castanea">Pacuvia castanea</taxonomicName>
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Curtis -the species type of the Chilean genus- in the shape of the clypeus plus frons, which are almost
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<pageBreakToken pageId="52" pageNumber="53" start="start">coplanar</pageBreakToken>
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and in the clypeus slightly emarginate, being this one a non-common feature among
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Liogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogenys" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
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species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |