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<document ID-CLB-Dataset="86636" ID-DOI="http://dx.doi.org/10.3897/vz.72.e80019" ID-GBIF-Dataset="574d4a22-42bb-4f6c-8e2d-6d49f9a2766d" ID-Pensoft-Pub="2625-8498-72-495" ID-Pensoft-UUID="218D013821E75A06B266110EB2A38790" ID-ZooBank="0B2305D45FE6489CAE3647BC0D25147B" ModsDocID="2625-8498-72-495" checkinTime="1656747396186" checkinUser="pensoft" docAuthor="Lyu, Zhi-Tong, Wang, Jian, Zeng, Zhao-Chi, Luo, Lin, Zhang, Yan-Wu, Guo, Chun-Peng, Ren, Jin-Long, Qi, Shuo, Mo, Yun-Ming &amp; Wang, Ying-Yong" docDate="2022" docId="0F6484DEEBB85FEAA4E6B11DFA8FD882" docLanguage="en" docName="VertZool 72: 495-512" docOrigin="Vertebrate Zoology 72" docPubDate="2022-07-01" docSource="http://dx.doi.org/10.3897/vz.72.e80019" docTitle="Occidozyga lingnanica Lyu &amp; Wang 2022, sp. nov." docType="treatment" docUuid="064C120B-3811-42D8-9550-0231528D723C" docUuidSource="ZooBank" docVersion="5" id="218D013821E75A06B266110EB2A38790" lastPageNumber="495" masterDocId="218D013821E75A06B266110EB2A38790" masterDocTitle="Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China" masterLastPageNumber="512" masterPageNumber="495" pageNumber="495" updateTime="1732809209568" updateUser="ExternalLinkService">
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<mods:title id="AC7A0CE2A23FD427F372225ED3E7AFAC">Taxonomic clarifications on the floating frogs (Anura: Dicroglossidae: Occidozyga sensu lato) in southeastern China</mods:title>
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<mods:namePart id="1669B176B8FECE89D01D65B05138160A">Lyu, Zhi-Tong</mods:namePart>
<mods:nameIdentifier id="FAEC52F088C871896874BE4E3C000113" type="ORCID">https://orcid.org/0000-0002-4891-3859</mods:nameIdentifier>
<mods:affiliation id="36C13CC524C9A476E6BA958F2273DC05">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
<mods:nameIdentifier id="69E01CCA62167B14CE281489FB880C5E" type="email">lvzht@foxmail.com</mods:nameIdentifier>
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<mods:namePart id="2E108F58B260C0F9BD9A3FC0D0B97A4B">Wang, Jian</mods:namePart>
<mods:nameIdentifier id="ED5A3E53742D6BBC8B5A19710F86E0EB" type="ORCID">https://orcid.org/0000-0003-4249-7767</mods:nameIdentifier>
<mods:affiliation id="0F3D40C5529507CAEE37B6B8FE4ECD1A">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
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<mods:namePart id="EE5122E703C87F4E5056CD3AC81105A6">Zeng, Zhao-Chi</mods:namePart>
<mods:affiliation id="2E8EB1332A16FA3719EA22B7BE093368">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
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<mods:namePart id="EC17E9F22AF67CB4E470C4E5FF7475CB">Luo, Lin</mods:namePart>
<mods:affiliation id="489FC272109E9044CE8698610D7E9173">Shenzhen Administrative Bureau for Wildlife and Forest Plants Protection, Shenzhen 518048, PR China</mods:affiliation>
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<mods:namePart id="25FA2411814E0FA974F896B1C0E43F85">Zhang, Yan-Wu</mods:namePart>
<mods:affiliation id="E298299BD9B8AA13C3F4C9B29D3EA4E4">Shenzhen Administrative Bureau for Wildlife and Forest Plants Protection, Shenzhen 518048, PR China</mods:affiliation>
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<mods:namePart id="0DB660032DCBAED068BD7202C3BAE588">Guo, Chun-Peng</mods:namePart>
<mods:affiliation id="76DD9F2664EDB798596429E623AFF235">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, PR China</mods:affiliation>
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<mods:roleTerm id="F84501AC6C754156FE22E593B4539FB4">Author</mods:roleTerm>
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<mods:namePart id="AB5B659DFD5555940AD638C6AB633AE7">Ren, Jin-Long</mods:namePart>
<mods:affiliation id="F3DC53B5199DD6E698A3BDE1BCE7EEA1">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, PR China</mods:affiliation>
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<mods:namePart id="F25383DE4BDC939EFF138D679D7C4603">Qi, Shuo</mods:namePart>
<mods:nameIdentifier id="4B8AAE3435AA170458C6E3211435C4B7" type="ORCID">https://orcid.org/0000-0002-2924-6093</mods:nameIdentifier>
<mods:affiliation id="5C82F292C08DDD0149D9DA65FFDD8BD5">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
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<mods:namePart id="86E6E5F11C650A8D9E0605DDC0370217">Mo, Yun-Ming</mods:namePart>
<mods:affiliation id="C4643F708665EE8914943ABFBA1803B4">Natural History Museum of Guangxi, Nanning 530012, PR China</mods:affiliation>
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<mods:namePart id="3CF6A253B9475EB844FA0A3A44C959F9">Wang, Ying-Yong</mods:namePart>
<mods:affiliation id="7941AC285F95F867A6164B00943AFB67">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
<mods:nameIdentifier id="57796F942306BEDE2980B5A1D16956E2" type="email">wangyy@mail.sysu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="BF15C8B30CDDED3A6693CFE2389BD5CB">text</mods:typeOfResource>
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<mods:title id="FDDD09AAA575E5500D067E20EABA0412">Vertebrate Zoology</mods:title>
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<mods:part id="D756D55D336BDB99A7EF054CF5901050">
<mods:date id="3B43207CCFCBB0C24A3BA342469CD756">2022</mods:date>
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<mods:number id="98D7CCA98C3AD39C6AC910F3A38A77CC">2022-07-01</mods:number>
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<mods:number id="6979F3671A089D50A77A3CC97F0A3633">72</mods:number>
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<mods:start id="B62FD0888333AE65111B12DB2C5F670C">495</mods:start>
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<mods:url id="18BC7758889C4BA6711433F446023355">http://dx.doi.org/10.3897/vz.72.e80019</mods:url>
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<mods:classification id="B62F149A1F1C721FA0A5488600CF5385">journal article</mods:classification>
<mods:identifier id="CF7AA6E7C78EE7527628933CBA27001F" type="DOI">http://dx.doi.org/10.3897/vz.72.e80019</mods:identifier>
<mods:identifier id="B712E72D5D7C168A8FFB9A7B5BFE3EAE" type="Pensoft-Pub">2625-8498-72-495</mods:identifier>
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<subSection id="956937621F3082BEF3D1BB1562CEE1BA" pageId="0" pageNumber="495" type="discussion">
<treatment id="0F6484DEEBB85FEAA4E6B11DFA8FD882" ID-GBIF-Taxon="196375048" LSID="urn:lsid:zoobank.org:act:064C120B-3811-42D8-9550-0231528D723C" httpUri="http://treatment.plazi.org/id/0F6484DEEBB85FEAA4E6B11DFA8FD882" lastPageNumber="495" pageId="0" pageNumber="495" scope_family="Dicroglossidae" scope_order="Anura">
<subSubSection id="0713A5A62DD06911BDB53C00AC7729AF" pageId="0" pageNumber="495" type="nomenclature">
<paragraph id="4DE6D0789472B120AEB727B0552207A1" pageId="0" pageNumber="495">
<taxonomicName id="A131A289EA5A8860B8976CAA4EC628C8" LSID="https://zoobank.org/064C120B-3811-42D8-9550-0231528D723C" authority="Lyu &amp; Wang" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica" status="sp. nov.">Occidozyga lingnanica Lyu &amp; Wang</taxonomicName>
<taxonomicNameLabel id="7B7189F034F94CC2C8E401C8BFAEF12D" pageId="0" pageNumber="495">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DF2C23A292751F3AA2AD994A6B0E8CCA" pageId="0" pageNumber="495" type="reference_group">
<paragraph id="F5175AC8806BCE3240EE40345F618B3D" pageId="0" pageNumber="495">
<taxonomicName id="EDD22F048B37960C2966BAB3A1B1F5C2" authority="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang, 2022" authorityName="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Ooeidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ooeidozyga laevis subsp. martensi" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="subSpecies" species="laevis" status="sp. nov." subSpecies="martensi">Ooeidozyga laevis martensi</taxonomicName>
-
<bibRefCitation id="C954D5A21FAA142729A1643B037877F6" author="Pope, CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="495" pagination="397 - 611" refId="B38" refString="Pope, CH, 1931. Notes on amphibians from Fukien, Hainan and other parts of China. Bulletin of the American Museum of Natural History 61: 397 - 611" title="Notes on amphibians from Fukien, Hainan and other parts of China." volume="61" year="1931">Pope 1931</bibRefCitation>
; Liu &amp; Hu 1961
</paragraph>
<paragraph id="30DF9CE592B61A8321389516884AB8BA" pageId="0" pageNumber="495">
<taxonomicName id="B4BB86122AD6AAC947811124B1207F15" authority="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang, 2022" authorityName="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga martensii" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="martensii" status="sp. nov.">Occidozyga martensii</taxonomicName>
-
<bibRefCitation id="E9F56B2C1FE94097CE663EFBE312B074" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B12" refString="Fei, L, Ye, CY, Huang, YZ, 1990. Key to Chinese Amphibians. Publishing House for Scientific and Technological Literature, Chongqing, China." title="Key to Chinese Amphibians. Publishing House for Scientific and Technological Literature, Chongqing, China." year="1990">Fei et al. 1990</bibRefCitation>
; Zhao &amp; Adler 1993;
<bibRefCitation id="4DB7BC37AF7BBB6CC7F71AD7C466D685" author="Li, ZC" journalOrPublisher="Algemeene Konst-en Letter-Bode" pageId="0" pageNumber="495" refId="B29" refString="Li, ZC, Xiao, Z, Liu, SR, 2011. Amphibians and reptiles of Guangdong. Guangdong Science and Technology Press, Guangzhou, China." title="Amphibians and reptiles of Guangdong. Guangdong Science and Technology Press, Guangzhou, China." year="2011">Li et al. 2011</bibRefCitation>
</paragraph>
<paragraph id="0A7DF80B06D4667E9B58785A681177E0" pageId="0" pageNumber="495">
<taxonomicName id="3E7D24BF297D23E66E462988AC9BDFCA" authority="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang, 2022" authorityName="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Phrynoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phrynoglossus martensii" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="martensii" status="sp. nov.">Phrynoglossus martensii</taxonomicName>
-
<bibRefCitation id="07962BB0AE1C0BC8C52252019ABCE02A" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B11" refString="Fei, L, Hu, SQ, Ye, CY, Huang, YZ, 2009. Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." title="Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." year="2009">Fei et al. 2009</bibRefCitation>
,
<bibRefCitation id="031636918B5A05B0DF1E4895CD14B030" author="Fei, L" journalOrPublisher="Herpetologica Sinica" pageId="0" pageNumber="495" pagination="1 - 43" refId="B13" refString="Fei, L, Ye, CY, Jiang, JP, 2010. Phylogenetic systematics of Ranidae. Herpetologica Sinica 12: 1 - 43" title="Phylogenetic systematics of Ranidae." volume="12" year="2010">2010</bibRefCitation>
,
<bibRefCitation id="0FBBD9C0CFE5E7FBEC00F355A9317EF9" author="Fei, L" journalOrPublisher="Herpetologica Sinica" pageId="0" pageNumber="495" refId="B14" refString="Fei, L, Ye, CY, Jiang, JP, 2012. Colored Atlas of Chinese Amphibians and Their Distributions. Sichuan Publishing House of Science &amp; Technology, Chengdu, China." title="Colored Atlas of Chinese Amphibians and Their Distributions. Sichuan Publishing House of Science &amp; Technology, Chengdu, China." year="2012">2012</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="71D23158A2114433CBBFBB087F8B2D0B" pageId="0" pageNumber="495" type="type material">
<paragraph id="8EA996401D13091325072F0648F29A8E" pageId="0" pageNumber="495">Type materials.</paragraph>
<paragraph id="5F3D7EF31E1E1D7749031E3F2486D8BE" pageId="0" pageNumber="495">
<emphasis id="79291B6A9FB1F44E7D867C16B8229A01" bold="true" pageId="0" pageNumber="495">Holotype.</emphasis>
SYS a008846, adult male, collected on 18 July 2021 by Zhi-Tong Lyu from Mt Wugongling (
<geoCoordinate id="351478C14291A90024C9BB03A47A9FAA" degrees="22.5914" direction="north" orientation="latitude" precision="5" value="22.5914">22.5914 N</geoCoordinate>
,
<geoCoordinate id="1103B8E7621B50E24B695CB503BDBC9A" degrees="114.4927" direction="east" orientation="longitude" precision="5" value="114.4927">114.4927 E</geoCoordinate>
), Shenzhen City, Guangdong.
<emphasis id="1CA342802523BC4BD1EA758A2E3AA9BE" bold="true" pageId="0" pageNumber="495">Paratypes.</emphasis>
Seven adult males and three adult females. Male SYS a008847 CIB 118529, collected at the same time as the holotype; males SYS a005589-90, collected on 15 November 2016 by Zhi-Tong Lyu and Jian Wang from Mt Wugongling; males SYS a007372-7373, and females SYS a07374-7375, collected on 5 September 2018 by Jian Wang and Hong-Hui Chen from
<normalizedToken id="9AAA4797A993293E7C1894198F0A7C9A" originalValue="Qi’ao">Qi'ao</normalizedToken>
Island (
<geoCoordinate id="16BEEBFF03192D16A99F82FCA6756651" degrees="22.4140" direction="north" orientation="latitude" precision="5" value="22.414">22.4140 N</geoCoordinate>
,
<geoCoordinate id="9EA2669E1CDE4217F7589B2F12991267" degrees="113.6446" direction="east" orientation="longitude" precision="5" value="113.6446">113.6446 E</geoCoordinate>
), Zhuhai City, Guangdong Province, China; female SYS a004650, collected on 14 April 2016 by Jian Wang from Mt Yunkai (
<geoCoordinate id="CC5CEA26BF7FC80688D1C8C298D78433" degrees="22.2786" direction="north" orientation="latitude" precision="5" value="22.2786">22.2786 N</geoCoordinate>
,
<geoCoordinate id="70CD33CBEEAB3D0F23663822ACD4BBCF" degrees="111.1886" direction="east" orientation="longitude" precision="5" value="111.1886">111.1886 E</geoCoordinate>
), Xinyi City, Guangdong; males SYS a007657-7658, collected on 8 April 2019 by Jian Wang from Mt Yunkai.
</paragraph>
</subSubSection>
<subSubSection id="58FA78BF373BD1314978D712915EEC33" pageId="0" pageNumber="495" type="other examined specimens">
<paragraph id="714ABE63FB94BD45FFAFE85F8BCD6338" pageId="0" pageNumber="495">Other examined specimens.</paragraph>
<paragraph id="CA3205EFD17F8EC689D31D7DA5AE98E9" pageId="0" pageNumber="495">
Five adult males and five adult females. Male SYS a005543 and female SYS a005542, from Mt Jiaoyiling (
<geoCoordinate id="9FE13D809AF7F5B60C9A4D66F06F83F8" degrees="21.1592" direction="north" orientation="latitude" precision="5" value="21.1592">21.1592 N</geoCoordinate>
,
<geoCoordinate id="73951DFC09BD6EB7DEDC1394F12D3311" degrees="110.3093" direction="east" orientation="longitude" precision="5" value="110.3093">110.3093 E</geoCoordinate>
), Zhanjiang City, Guangdong; female SYS a005267, from Mt. Diaoluo (18.6408 N, 109.9317 E), Lingshui Li Autonomous County, Hainan; males SYS a005436-5437, 5440-5441 and females SYS a005438-5439, 5442, from Mt Bawangling (1 9.0944 N,
<geoCoordinate id="3FEC9B5AF66A144A3C54B4F407A66B55" degrees="109.0491" direction="east" orientation="longitude" precision="5" value="109.0491">109.0491 E</geoCoordinate>
), Changjiang Li Autonomous County, Hainan.
</paragraph>
</subSubSection>
<subSubSection id="017E0D28AA9D2E6913A160BE58C7E08B" pageId="0" pageNumber="495" type="etymology">
<paragraph id="3A6CFF08065842035C3CBDAD1823CC15" pageId="0" pageNumber="495">Etymology.</paragraph>
<paragraph id="7DFFD77763264663C2BC18F06E8625E7" pageId="0" pageNumber="495">
The specific name
<taxonomicName id="B27DAE737376AA5A91D2ED01735ADDA1" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="4A19B60B346FFBDD356597DDCDCB2A1D" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
refers to the lingnan region, a geographic area covering Guangdong, Guangxi, and Hainan in southeastern China, where this new frog species occurs in. This specific name is also dedicated to the former Lingnan University (1888-1952) that was incorporated into Sun Yat-sen University after 1953.
</paragraph>
</subSubSection>
<subSubSection id="FE312C34A66471D38778D866595856C9" pageId="0" pageNumber="495" type="common name">
<paragraph id="5E243A68A16D4EA6637DD46252FF840C" pageId="0" pageNumber="495">Common name.</paragraph>
<paragraph id="368A948701D01B708A0B07BE62895EE3" pageId="0" pageNumber="495">
&quot;Lingnan floating frog&quot; in English / &quot;岭南浮蛙 (lǐng
<normalizedToken id="E251D82A5413C1FF6D59AA6997C30F71" originalValue="nán">nan</normalizedToken>
<normalizedToken id="50F3B5272F766D7D38E83040A808AC01" originalValue="fú">fu</normalizedToken>
<normalizedToken id="D278AA9337B9818877C1E9D76FB22151" originalValue="wā)”">wa)&quot;</normalizedToken>
in Chinese.
</paragraph>
</subSubSection>
<subSubSection id="0223EDF776DC93307BE2BBEEC03D2F49" pageId="0" pageNumber="495" type="diagnosis">
<paragraph id="8F5E8F477573F75ECD30DE0E4BD2BE05" pageId="0" pageNumber="495">Diagnosis.</paragraph>
<paragraph id="A03012C34DF3636AF667FFADBBABA34F" pageId="0" pageNumber="495">(1) Body stocky, size small, SVL 19.9-22.1 mm in males (n=8) and 26.8-28.8 mm in females (n=3); (2) snout short triangular shaped; (3) nostrils located laterally; (4) eye orientation laterally; (5) canthus rostralis visible, rounded; (6) loreal region vertical, not concave/convex; (7) tongue wide and swollen, unnotched, rounded distally, lingual papilla absent; (8) vomerine ridge and vomerine teeth absent; (9) supratympanic fold distinct and raised, slightly curved on the temporal region; (10) tympanum hidden, edge invisible; (11) relative finger lengths II=I&lt;IV&lt;III, relative toe lengths I&lt;II&lt;V&lt;III&lt;IV; (12) fingers without webs, toes with two third webs; (13) heels not meeting, tibio-tarsal articulation reaching at the posterior margin of supratympanic fold; (14) tarsal fold absent.</paragraph>
</subSubSection>
<subSubSection id="5EE1E02F51B5A232470192C027CC277C" pageId="0" pageNumber="495" type="comparisons">
<paragraph id="7855FB23A16C796310DD8FD88CECBE3B" pageId="0" pageNumber="495">Comparisons.</paragraph>
<paragraph id="7A920880560131F1950EDAC04069EC39" pageId="0" pageNumber="495">
<taxonomicName id="7236678FCF950FF6EF2D130CC47ACF75" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="6C81A8F295B21FFEB5444701F1FDD627" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. has been long-term misidentified as
<taxonomicName id="4C47A7B8429419CC997A778132A52602" lsidName="O. martensii" pageId="0" pageNumber="495" rank="species" species="martensii">
<emphasis id="EE5CD934BE9D0CA159152486F07214D4" italics="true" pageId="0" pageNumber="495">O. martensii</emphasis>
</taxonomicName>
, however, it differs from the latter by a combination of the following morphological characters: tympanum hidden, edge invisible (vs. tympanum edge raised), relative finger lengths II=I&lt;IV (vs. II=IV&lt;I), relative toe lengths V&lt;III (vs. III&lt;V), tarsal fold absent (vs. present), tibio-tarsal articulation reaching the posterior margin of supratympanic fold (vs. reaching the region of eye).
</paragraph>
<paragraph id="30ADDAAACF01811F938F4C2B1D973466" pageId="0" pageNumber="495">
<taxonomicName id="88240673A28E1FC842A5DF9EAD04CAD4" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="38F33EF328A2AB87A503357879246EFF" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. can be distinguished from the remaining three known congeners in Clade VIII (Fig.
<figureCitation id="0BB9BB156EC70EA186BA62C05704CE7D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic trees of the genus Occidozyga sensu lato based on mitochondrial genes 16 S and CO 1. Bayesian posterior probabilities (BPP) and bootstrap supports (BS) are labeled above and below the nodes respectively. Numbers at the terminal of the lineages correspond to the IDs in Table 1." figureDoi="10.3897/vz.72.e80019.figure1" httpUri="https://binary.pensoft.net/fig/709314" pageId="0" pageNumber="495">1</figureCitation>
) by the canthus rostralis rounded (vs. absent in
<taxonomicName id="549C353EA157D2ED35813068B33D4935" lsidName="O. magnapustulosa" pageId="0" pageNumber="495" rank="species" species="magnapustulosa">
<emphasis id="DE6E590EC0D02FCF03E8E65680CF1FF0" italics="true" pageId="0" pageNumber="495">O. magnapustulosa</emphasis>
</taxonomicName>
), loreal region oblique, not concave or convex (vs. slight convex in
<taxonomicName id="C440AF642B151590BAF9ABDFCEFDA138" lsidName="O. magnapustulosa" pageId="0" pageNumber="495" rank="species" species="magnapustulosa">
<emphasis id="1A5EB212C06DCFAE4AD3614CE59E0A03" italics="true" pageId="0" pageNumber="495">O. magnapustulosa</emphasis>
</taxonomicName>
and
<taxonomicName id="A15A820C400F3269A44382408E0C5949" lsidName="O. swanbornorum" pageId="0" pageNumber="495" rank="species" species="swanbornorum">
<emphasis id="03A0AFCAD1115CD2AD39D2A3506D6206" italics="true" pageId="0" pageNumber="495">O. swanbornorum</emphasis>
</taxonomicName>
comb. nov.), relative finger lengths II=I&lt;IV (vs. II&lt;IV&lt;I in
<taxonomicName id="BA2E4EB8962074F12C00AEB1E9C967B9" lsidName="O. myanhessei" pageId="0" pageNumber="495" rank="species" species="myanhessei">
<emphasis id="160FDFB868F52D14ADB95AEABA8D976D" italics="true" pageId="0" pageNumber="495">O. myanhessei</emphasis>
</taxonomicName>
comb. nov., IV&lt;II&lt;I in
<taxonomicName id="69CAAEEF0A0A9049315EC79557A72FD8" lsidName="O. swanbornorum" pageId="0" pageNumber="495" rank="species" species="swanbornorum">
<emphasis id="6B79923AB9715CE43F9AEC528D4DF6F1" italics="true" pageId="0" pageNumber="495">O. swanbornorum</emphasis>
</taxonomicName>
comb. nov.), tarsal fold absent (vs. present in
<taxonomicName id="610A9617254F45A64F3A63FF9A24E503" lsidName="O. magnapustulosa" pageId="0" pageNumber="495" rank="species" species="magnapustulosa">
<emphasis id="AE60831AD48DFA70E3B4A2EA4EB51E5E" italics="true" pageId="0" pageNumber="495">O. magnapustulosa</emphasis>
</taxonomicName>
and
<taxonomicName id="1896A84DF4FF87B82C3FC633F2B73AA7" lsidName="O. myanhessei" pageId="0" pageNumber="495" rank="species" species="myanhessei">
<emphasis id="95BBD72E03930DD2DE4FC1D98F311D54" italics="true" pageId="0" pageNumber="495">O. myanhessei</emphasis>
</taxonomicName>
comb. nov.).
</paragraph>
<paragraph id="69BAEEEECEA3FE390605D5969846C31B" pageId="0" pageNumber="495">
For the remaining 13 congeners,
<taxonomicName id="0EFF66959B01DFBD9A1AF1C8A479AB14" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="6BE90642B18B9B17840F4B1D34C6E151" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. distinctly differs from
<taxonomicName id="B469DDF380F0EDB3FE9969BFB9F5890C" lsidName="O. lima" pageId="0" pageNumber="495" rank="species" species="lima">
<emphasis id="DD4E0E13B797B3AB695C1EC9399650B5" italics="true" pageId="0" pageNumber="495">O. lima</emphasis>
</taxonomicName>
,
<taxonomicName id="A6304EAC4A6C332B949B3A6DB71BC67B" lsidName="O. obscura" pageId="0" pageNumber="495" rank="species" species="obscura">
<emphasis id="02E6912351CC8F174112B5DF353E5EBA" italics="true" pageId="0" pageNumber="495">O. obscura</emphasis>
</taxonomicName>
comb. nov., and
<taxonomicName id="FB30B6A5358A1674ABFE23A08272D6AA" lsidName="O. berbeza" pageId="0" pageNumber="495" rank="species" species="berbeza">
<emphasis id="9CA7FEF8D3AA8C24F6758C1D6C972383" italics="true" pageId="0" pageNumber="495">O. berbeza</emphasis>
</taxonomicName>
by the tongue wide and swollen (vs. narrow and slender); from
<taxonomicName id="284416FB97A40B0F574ABA5A0A1A3F71" lsidName="O. rhacoda" pageId="0" pageNumber="495" rank="species" species="rhacoda">
<emphasis id="D362050F855BBF26E842A7D4CC2FAA77" italics="true" pageId="0" pageNumber="495">O. rhacoda</emphasis>
</taxonomicName>
by the dorsolateral fold absent (vs. present); from
<taxonomicName id="268B0A1F226B26567413068E87FD8E85" lsidName="O. shiwandashanensis" pageId="0" pageNumber="495" rank="species" species="shiwandashanensis">
<emphasis id="20BF23BCE081836F0ED56259E03DF050" italics="true" pageId="0" pageNumber="495">O. shiwandashanensis</emphasis>
</taxonomicName>
by the tarsal fold absent (vs. present); from
<taxonomicName id="E2A86EEE19FC512D8C5F85DCC706BBBF" lsidName="O. celebensis" pageId="0" pageNumber="495" rank="species" species="celebensis">
<emphasis id="6044B8019E0F09AEDB46F589AD90AE20" italics="true" pageId="0" pageNumber="495">O. celebensis</emphasis>
</taxonomicName>
,
<taxonomicName id="7504A8CDFBEB96A111031344E94D7864" lsidName="O. laevis" pageId="0" pageNumber="495" rank="species" species="laevis">
<emphasis id="8956FA69151563634F914E3EEB82715C" italics="true" pageId="0" pageNumber="495">O. laevis</emphasis>
</taxonomicName>
, and
<taxonomicName id="A315F4C2175E0F9E6D0733E82F4EEB75" lsidName="O. sumatrana" pageId="0" pageNumber="495" rank="species" species="sumatrana">
<emphasis id="923972E16A938B43CFB1A3CA1EEC80EA" italics="true" pageId="0" pageNumber="495">O. sumatrana</emphasis>
</taxonomicName>
by the eye orientation laterally (vs. vertically); from
<taxonomicName id="2997538E9619EB8F720A48C140165AF4" lsidName="O. baluensis" pageId="0" pageNumber="495" rank="species" species="baluensis">
<emphasis id="FC166193124EEAA32BC8546F16140F25" italics="true" pageId="0" pageNumber="495">O. baluensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4FD588C67690160F9DA02559727C422D" lsidName="O. floresiana" pageId="0" pageNumber="495" rank="species" species="floresiana">
<emphasis id="87F79CBE209FDBE5633E86DD45A238EB" italics="true" pageId="0" pageNumber="495">O. floresiana</emphasis>
</taxonomicName>
, and
<taxonomicName id="43EAAEF1F6FCF98FB7B78B89974A5507" lsidName="O. semipalmata" pageId="0" pageNumber="495" rank="species" species="semipalmata">
<emphasis id="94CE6394ED46A9DEA0E310A9869251A7" italics="true" pageId="0" pageNumber="495">O. semipalmata</emphasis>
</taxonomicName>
by the canthus rostralis rounded (vs. absent); from
<taxonomicName id="C000E09CF16CDF0049F90C2EC477F046" lsidName="O. diminutiva" pageId="0" pageNumber="495" rank="species" species="diminutiva">
<emphasis id="A675A611873D8C42F01321C7B526FC59" italics="true" pageId="0" pageNumber="495">O. diminutiva</emphasis>
</taxonomicName>
and
<taxonomicName id="78957678A8D66A81702CCCE6A15576D9" lsidName="O. tompotika" pageId="0" pageNumber="495" rank="species" species="tompotika">
<emphasis id="95B241D37987210C2AD73508B0D591A0" italics="true" pageId="0" pageNumber="495">O. tompotika</emphasis>
</taxonomicName>
by the supratympanic fold curved (vs. straight).
</paragraph>
</subSubSection>
<subSubSection id="E936B3FB6107E2EEA103C58C3D4052C7" pageId="0" pageNumber="495" type="description">
<paragraph id="D3BB5A748728B5038196B2CAB3C31C25" pageId="0" pageNumber="495">Description of holotype.</paragraph>
<paragraph id="872FB7240878E3BEA9F8AEC31D2F33F2" pageId="0" pageNumber="495">
SYS a008846, adult male (Fig.
<figureCitation id="141AA70F336015AFADBDE63C54C986CF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the adult male holotype SYS a 008846 of Occidozyga lingnanica sp. nov. in life. A dorsolateral view B ventral view C left hand D right foot E dorsal view of right hand showing the nuptial pad F calling in situ showing the subgular vocal sac." figureDoi="10.3897/vz.72.e80019.figure5" httpUri="https://binary.pensoft.net/fig/709318" pageId="0" pageNumber="495">5</figureCitation>
). Body stocky, small-sized, SVL 21.4 mm. Head longer than wide (HDW/HDL 0.97), plat above; pineal ocellus distinct; snout short triangular shaped, distinctly protruding beyond lower jaw, tip of snout rounded in dorsal view and profile; canthus rostralis rounded, loreal region oblique, not concave or convex; nostril rounded, laterally, not raised, at the middle between tip of snout and eye; eye orientation laterally, pupil diamond shaped; interorbital space distinctly narrower than internarial distance; tympanum hidden, edge invisible; vomerine ridge and teeth absent; tongue wide and swollen, unnotched, rounded distally, lingual papilla absent.
</paragraph>
<caption id="695BEFD54C5E82F9135E72262E609B4D" doi="10.3897/vz.72.e80019.figure5" httpUri="https://binary.pensoft.net/fig/709318" pageId="0" pageNumber="495" start="Figure 5" startId="F5">
<paragraph id="D79FBFFED34B1CB3F86049444424E9CC" pageId="0" pageNumber="495">
<emphasis id="55863BA8A9D8DB3C2323B78853A772B3" bold="true" pageId="0" pageNumber="495">Figure 5.</emphasis>
Morphological features of the adult male holotype SYS a008846 of
<taxonomicName id="727F8B871666D9ADC5C8F069F84CB669" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="06EBC8CA8377B71774669FBA8476A64E" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. in life.
<emphasis id="D33F953D0B4D679E55E8C21B6B2C5390" bold="true" pageId="0" pageNumber="495">A</emphasis>
dorsolateral view
<emphasis id="7A56AF4D29BC53215F48A9F559806825" bold="true" pageId="0" pageNumber="495">B</emphasis>
ventral view
<emphasis id="1D719E2C6B052781BF51FBDFE290812B" bold="true" pageId="0" pageNumber="495">C</emphasis>
left hand
<emphasis id="0DFA00EF1EBBB42D1BED7B7375A55A2E" bold="true" pageId="0" pageNumber="495">D</emphasis>
right foot
<emphasis id="38C94F47807874EB779B224DFB15B149" bold="true" pageId="0" pageNumber="495">E</emphasis>
dorsal view of right hand showing the nuptial pad
<emphasis id="5B9DBCEB9FD65AAC6747ED1D997B6553" bold="true" pageId="0" pageNumber="495">F</emphasis>
calling in situ showing the subgular vocal sac.
</paragraph>
</caption>
<paragraph id="E53E2B49C848626B2CA14E652EE26C79" pageId="0" pageNumber="495">Forelimbs short, lower arm 15% of SVL and hand 25% of SVL; fingers relatively thin and long, relative finger lengths II=I&lt;IV&lt;III; tips of fingers rounded, not dilated and without disks; fingers without webs and fringes; subarticular tubercles present at the bases of each finger, prominent and rounded; supernumerary tubercles absent; inner and outer palmar tubercles prominent and rounded.</paragraph>
<paragraph id="188749A1700428B84F5E0EA2A41ED297" pageId="0" pageNumber="495">Hind limbs robust, tibia 44% of SVL and foot 63% of SVL; heels not meeting when hind limbs flexed at right angles to the axis of the body; tibio-tarsal articulation reaching at the posterior margin of supratympanic fold when hind limb is stretched along the side of the body; toes distinctly long and thin, relative lengths I &lt;II &lt;V &lt;III &lt;IV; tips of toes rounded, dilated into rounded disks; toes with two third webs, metatarsal web present, distinct lateral fringes on lateral edges of toes I and V; subarticular tubercles rounded, prominent; inner metatarsal tubercle large and long-elliptic, distinctly raised, length triple the width; outer metatarsal tubercle absent; tarsal fold absent.</paragraph>
<paragraph id="10DA2CDDC53A13D979ECDEDC14A9CDCB" pageId="0" pageNumber="495">Dorsal surface relatively rough, large tubercles scattering on skin of dorsum, flanks, and dorsal limbs, not bearing spinules on the dorsal skin; a faint fold across head between orbits; supratympanic fold distinct, raised, extending from the posterior corner of the eye, slightly curved on the temporal region, to the previous shoulder; dorsolateral fold absent. Ventral surface with large flattened tubercles; a fold across breast; dense granules on the ventral tarsi.</paragraph>
</subSubSection>
<subSubSection id="D4BFB9C0C93441B290A1C7CDE1DA7F25" pageId="0" pageNumber="495" type="coloration of holotype">
<paragraph id="B9B73550893521D1345EED2A2C472465" pageId="0" pageNumber="495">Coloration of holotype.</paragraph>
<paragraph id="FA7930836D1687582427D1B8DAA15721" pageId="0" pageNumber="495">
In life (Fig.
<figureCitation id="8722CD9CC40E099E3B44662E496174D7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the adult male holotype SYS a 008846 of Occidozyga lingnanica sp. nov. in life. A dorsolateral view B ventral view C left hand D right foot E dorsal view of right hand showing the nuptial pad F calling in situ showing the subgular vocal sac." figureDoi="10.3897/vz.72.e80019.figure5" httpUri="https://binary.pensoft.net/fig/709318" pageId="0" pageNumber="495">5</figureCitation>
), dorsal surface grayish brown with irregular black speckles; dorsal limbs with dark brown transverse bars; mid-dorsal stripe yellowish brown but indistinct; a narrow transverse bar between orbits; supratympanic fold dark brown. Pupil bordered with yellow; iris brown with irregular dark or light speckles. Skin of throat dark with white mottling; skin of chest and belly uniform creamy white; ventral surface of limbs grayish pink with dark speckles. Nuptial pad light yellow, slightly transparent.
</paragraph>
<paragraph id="C5230EC4B020BDC41BCFB220FB1072B4" pageId="0" pageNumber="495">In preservative, dorsum light gray; black speckles on dorsum and transverse bars on limbs light brown; mid-dorsal stripe grayish white and more distinct; nuptial pad light gray, slightly transparent; ventral surface grayish white; mottling on throat gray white.</paragraph>
</subSubSection>
<subSubSection id="2AD876BF510EAE97EFD0D366250A2F1F" pageId="0" pageNumber="495" type="variation">
<paragraph id="6122C5DC97DEFB6D0D4B65C4C0F6CA4B" pageId="0" pageNumber="495">Variation.</paragraph>
<paragraph id="DB76066B7C4231F5B3E821BDDF68AE61" pageId="0" pageNumber="495">
The measurements of the type series are given in Table
<tableCitation id="A5286D63657E4A0D8D07A6DC5458B708" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Measurements (in mm) of the examined specimens of Occidozyga lingnanica sp. nov." httpUri="http://table.plazi.org/id/E42770B6E64558992AD2C93C99E1709F" pageId="0" pageNumber="495" tableUuid="E42770B6E64558992AD2C93C99E1709F">4</tableCitation>
. All individuals were similar in morphology. Dorsal coloration varies in life, from light brown, yellowish brown, to dark brown (Figs
<figureCitation id="78FDCD09788EA61A116FF2E47A635753" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the adult male holotype SYS a 008846 of Occidozyga lingnanica sp. nov. in life. A dorsolateral view B ventral view C left hand D right foot E dorsal view of right hand showing the nuptial pad F calling in situ showing the subgular vocal sac." figureDoi="10.3897/vz.72.e80019.figure5" httpUri="https://binary.pensoft.net/fig/709318" pageId="0" pageNumber="495">5</figureCitation>
,
<figureCitation id="163278B525AB9D8679FBFCA0B606B468" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation of Occidozyga lingnanica sp. nov. in life. A adult male paratype SYS a 008847 B adult female paratype SYS a 004650 C adult male paratype SYS a 007373 D adult female paratype SYS a 007375 E adult male SYS a 005437 F adult male SYS a 005440." figureDoi="10.3897/vz.72.e80019.figure6" httpUri="https://binary.pensoft.net/fig/709319" pageId="0" pageNumber="495">6</figureCitation>
); mid-dorsal stripe varies among individuals, distinct (Figs
<figureCitation id="2C8B820EBF82624B6ADB464F96506A5D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of Occidozyga obscura comb. nov. and Occidozyga lingnanica sp. nov. in preservative. Occidozyga obscura comb. nov.: A adult male SYS a 008121 B adult female SYS a 008122; Occidozyga lingnanica sp. nov.: C adult male paratype SYS a 007373 D adult female paratype SYS a 007375. 1 dorsal view 2 ventral view." figureDoi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495">4C</figureCitation>
,
<figureCitation id="272B29591EA085B91B5C00EAECA61820" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation of Occidozyga lingnanica sp. nov. in life. A adult male paratype SYS a 008847 B adult female paratype SYS a 004650 C adult male paratype SYS a 007373 D adult female paratype SYS a 007375 E adult male SYS a 005437 F adult male SYS a 005440." figureDoi="10.3897/vz.72.e80019.figure6" httpUri="https://binary.pensoft.net/fig/709319" pageId="0" pageNumber="495">6C</figureCitation>
), indistinct (Fig.
<figureCitation id="942833AB53F7FBE0564286B6F9A8DE6F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological features of the adult male holotype SYS a 008846 of Occidozyga lingnanica sp. nov. in life. A dorsolateral view B ventral view C left hand D right foot E dorsal view of right hand showing the nuptial pad F calling in situ showing the subgular vocal sac." figureDoi="10.3897/vz.72.e80019.figure5" httpUri="https://binary.pensoft.net/fig/709318" pageId="0" pageNumber="495">5</figureCitation>
), or absent (Figs
<figureCitation id="E5329B3135864049513588E4D94DB52A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of Occidozyga obscura comb. nov. and Occidozyga lingnanica sp. nov. in preservative. Occidozyga obscura comb. nov.: A adult male SYS a 008121 B adult female SYS a 008122; Occidozyga lingnanica sp. nov.: C adult male paratype SYS a 007373 D adult female paratype SYS a 007375. 1 dorsal view 2 ventral view." figureDoi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495">4D</figureCitation>
,
<figureCitation id="CBC0B6C5206D29E703C04933ACBED989" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation of Occidozyga lingnanica sp. nov. in life. A adult male paratype SYS a 008847 B adult female paratype SYS a 004650 C adult male paratype SYS a 007373 D adult female paratype SYS a 007375 E adult male SYS a 005437 F adult male SYS a 005440." figureDoi="10.3897/vz.72.e80019.figure6" httpUri="https://binary.pensoft.net/fig/709319" pageId="0" pageNumber="495">6A, B, D-F</figureCitation>
); an irregular orange marking on the occipital region in some individuals (Fig.
<figureCitation id="81868789A4150F6258E19DD19C979F04" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation of Occidozyga lingnanica sp. nov. in life. A adult male paratype SYS a 008847 B adult female paratype SYS a 004650 C adult male paratype SYS a 007373 D adult female paratype SYS a 007375 E adult male SYS a 005437 F adult male SYS a 005440." figureDoi="10.3897/vz.72.e80019.figure6" httpUri="https://binary.pensoft.net/fig/709319" pageId="0" pageNumber="495">6F</figureCitation>
).
</paragraph>
<caption id="6A0E1BEB977BFA5FE66C72E3B548822C" doi="10.3897/vz.72.e80019.figure6" httpUri="https://binary.pensoft.net/fig/709319" pageId="0" pageNumber="495" start="Figure 6" startId="F6">
<paragraph id="E441544DEA2A183C50414AF3B27E9BF3" pageId="0" pageNumber="495">
<emphasis id="99D8B0E39225EF1365BB7A78A724F335" bold="true" pageId="0" pageNumber="495">Figure 6.</emphasis>
Color variation of
<taxonomicName id="5A592BD59CB193FA09B71971CE0C89B5" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="E19481D991805D7993D0C52F416946C7" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. in life.
<emphasis id="87B2800EE66413AE70A2553648E6FE93" bold="true" pageId="0" pageNumber="495">A</emphasis>
adult male paratype SYS a008847
<emphasis id="4AED448540DA110B01A46B7FE72D3D2D" bold="true" pageId="0" pageNumber="495">B</emphasis>
adult female paratype SYS a004650
<emphasis id="4CECCBE7AA07C9A4C1972EE63E1D9729" bold="true" pageId="0" pageNumber="495">C</emphasis>
adult male paratype SYS a007373
<emphasis id="7E5AA7A1EB92785A8B608E84A5AF41F2" bold="true" pageId="0" pageNumber="495">D</emphasis>
adult female paratype SYS a007375
<emphasis id="0C4385A067FC555F3D3AB4211E5A78B7" bold="true" pageId="0" pageNumber="495">E</emphasis>
adult male SYS a005437
<emphasis id="9127051429C1B02C011F409D63A3CDC1" bold="true" pageId="0" pageNumber="495">F</emphasis>
adult male SYS a005440.
</paragraph>
</caption>
<caption id="E42770B6E64558992AD2C93C99E1709F" ID-Table-UUID="E42770B6E64558992AD2C93C99E1709F" httpUri="http://table.plazi.org/id/E42770B6E64558992AD2C93C99E1709F" pageId="0" pageNumber="495" start="Table 4" startId="T4">
<paragraph id="F53250D35FC48F65045AC101E30C60C0" pageId="0" pageNumber="495">
<emphasis id="4DAF5499540694FB7D0306B4F88746AC" bold="true" pageId="0" pageNumber="495">Table 4.</emphasis>
Measurements (in mm) of the examined specimens of
<taxonomicName id="E3E0C503F7900C7136F63DFAECC0D1E6" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="7B1E94CAA368A39049B8EA374E45A80E" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph id="FF1BD4955EC65E8E9CDB3C613236CE2F" pageId="0" pageNumber="495">
<table id="BB5B40A5376E972EAB952471FC7A3ED9" pageId="0" pageNumber="495">
<tr id="11E393D92508AA9A474AFC5009F34305" pageId="0" pageNumber="495">
<td id="B9DAED284ED8E2479C31A38E7713050B" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="E274297427588680C99A8F9D522F99FD" bold="true" pageId="0" pageNumber="495">Specimen</emphasis>
</td>
<td id="943B24B9D0171AC15F2035A66F6048F6" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="4BAEE0B19784119FEF7C51C633742292" bold="true" pageId="0" pageNumber="495">SYS a007372</emphasis>
</td>
<td id="35CCF21376B5A74F03019F4DBE296E9D" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="B1E95F19CF8CE6B2514C8F46DDF24D62" bold="true" pageId="0" pageNumber="495">SYS a007373</emphasis>
</td>
<td id="A34CB4AE4E934EE6B2E17E4453B0371C" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="2B5CA8C72C24A26AED894FAE89160B55" bold="true" pageId="0" pageNumber="495">SYS a008846</emphasis>
</td>
<td id="FB5919DB28A2A45C389421FEC0A31140" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="8AE99C8587E0EB63187004BDF26DA155" bold="true" pageId="0" pageNumber="495">SYS a008847</emphasis>
</td>
<td id="26F411A8000C7D39FB20361EFD222FB7" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="FA94F0BE08999907D184738E8F051639" bold="true" pageId="0" pageNumber="495">SYS a005589</emphasis>
</td>
<td id="EA210B1A361EF1EDBB9755932EBECBF5" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="56938E4358179E6ABFF184D918502FE2" bold="true" pageId="0" pageNumber="495">SYS a005590</emphasis>
</td>
<td id="A5B6F4BFDDF72C8D714DD34236822ACA" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="BA0D65E4A1079E1656D8E94D5D8F3490" bold="true" pageId="0" pageNumber="495">SYS a007657</emphasis>
</td>
<td id="5B40BE9190F98AF3E67A45450FFC37B1" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="EE781DDCE92C4BC8E27EBEF182C2E565" bold="true" pageId="0" pageNumber="495">SYS a007658</emphasis>
</td>
<td id="C4165A278503D95EF6339FC14A53CBCF" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="AE963412A1D9012F7F62A4AE16146721" bold="true" pageId="0" pageNumber="495">SYS a007374</emphasis>
</td>
<td id="8C8E7648DD66AA7D8CD6EC4E2453F5A6" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="EE21722C87AB2A8B3C30E8D4A4D60E0F" bold="true" pageId="0" pageNumber="495">SYS a007375</emphasis>
</td>
<td id="045372C36FAA11F7CBD0EAEF148AD680" colspan="1" pageId="0" pageNumber="495" rowspan="1">
<emphasis id="9711D07C2A8E7051E79CBFCAB8F5422B" bold="true" pageId="0" pageNumber="495">SYS a004650</emphasis>
</td>
</tr>
<tr id="D4E249A7750A4660F1CC29A6C661EA8A" pageId="0" pageNumber="495">
<td id="6D8D24BBB6FEC2BB4B26AFC267F9F073" colspan="1" pageId="0" pageNumber="495" rowspan="1">Sex</td>
<td id="8F2B6AC5E1C0EFA3E8415F95D53DD0A3" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="0B7A3996D991AC4AD3B89E8ADC893E76" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="2154C68D5B0E26E4976A18BACEA0E6E5" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="24B42C19D5C29043E92E2E6A836A750F" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="85A18541050B6CD1619CFF34445450D6" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="84E72BE32905BA1E75C55720C50F5759" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="3B80276EA1361B6B67084F29F7933C5A" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="780BFB38531862DF5E6367E274A35AD5" colspan="1" pageId="0" pageNumber="495" rowspan="1">Male</td>
<td id="5BBA0CCEB6CE184172D64237663B7A53" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
<td id="8E8ACBDA9C9D1175ECD0DF835D9E75C5" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
<td id="58C59E6ABDEDCC1DB2B07F7B42FCBFF1" colspan="1" pageId="0" pageNumber="495" rowspan="1">Female</td>
</tr>
<tr id="EE0455C6F4F7FAFF6CCDC79F98D578B7" pageId="0" pageNumber="495">
<td id="BC40991E8662A52C99A5DD31A19625BF" colspan="1" pageId="0" pageNumber="495" rowspan="1">SVL</td>
<td id="9FB644B7406948067D0A7FE7921A284E" colspan="1" pageId="0" pageNumber="495" rowspan="1">22.1</td>
<td id="728B3568535AA8908F7B00DDE00DB95D" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.6</td>
<td id="8D8E5986AF7D63B5D5092673E7538C30" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.4</td>
<td id="7587299464C1D8704D5FAB1795F136F2" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.5</td>
<td id="1EDADC5AF8ABB9FB80E20A388A232837" colspan="1" pageId="0" pageNumber="495" rowspan="1">19.9</td>
<td id="E77FC331E771737408F8BBDA12993F4E" colspan="1" pageId="0" pageNumber="495" rowspan="1">20.2</td>
<td id="A59C3CAB8D4489A2D3E7D483CD0714CC" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.8</td>
<td id="EF2C94E3851E52BBCEE13C7933739FC3" colspan="1" pageId="0" pageNumber="495" rowspan="1">21.7</td>
<td id="0D8AFC1E22A7AF61932F569445082B7B" colspan="1" pageId="0" pageNumber="495" rowspan="1">26.8</td>
<td id="E5DF75870D08EB3DAD88FC169B4880B3" colspan="1" pageId="0" pageNumber="495" rowspan="1">28.5</td>
<td id="7DA5413028B18ED9190A2843A8E2BCB6" colspan="1" pageId="0" pageNumber="495" rowspan="1">28.8</td>
</tr>
<tr id="A8D7CDA6EE74B7AF916C8BBC93908FCE" pageId="0" pageNumber="495">
<td id="DD7FAEB706787E8F724D84C416BA83D6" colspan="1" pageId="0" pageNumber="495" rowspan="1">HDL</td>
<td id="1EB070447DE30FCBF3C4A70520C67B08" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.8</td>
<td id="6F225D3636465BD0B69012726951CE41" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.1</td>
<td id="41FF7BFA3E17B215BC89565512BC5142" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.5</td>
<td id="EE97130899DD2B911C4282B985BE0AD1" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.5</td>
<td id="4209016EA5CD8515F8E057AA19F89DA2" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.6</td>
<td id="E816185B9FF7ED4289AE370ECAAC366D" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.6</td>
<td id="8AAA6C171B4A54FF46EE9B022B1852A9" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.1</td>
<td id="35D9440014110491A3DD58010A1CDACB" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.8</td>
<td id="C452EA581EA492D41809DF4EB92DD9F4" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.0</td>
<td id="D68E1E12B178DF239C580B56B6AA9478" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.5</td>
<td id="1D3251EDDBCD59F52A59BD04902BDDB4" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.0</td>
</tr>
<tr id="8D763570D298BB433EDAB6452D565502" pageId="0" pageNumber="495">
<td id="6C853DB39D6D412CDACAAB39584EA0DB" colspan="1" pageId="0" pageNumber="495" rowspan="1">HDW</td>
<td id="FD1F0504D592B482FFDB78BD44D2D665" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.4</td>
<td id="010261BAD971BC223079CC5A0B90FB00" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.7</td>
<td id="2DCB05E90F147B4C0D054EC48B55F49F" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.2</td>
<td id="2B7052B2D06A1256C26E8F6EE6154CEB" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.2</td>
<td id="B070CB6164755C75A08884E8DBEEE284" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.5</td>
<td id="D65C3DA8F8A93079D8CC72F99AA902E6" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.5</td>
<td id="CF654596E6167DB018596CC6345F6EBD" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.6</td>
<td id="BBF94D1CAE25D94B6A5E7665D43879B4" colspan="1" pageId="0" pageNumber="495" rowspan="1">7.5</td>
<td id="7DB3884103077405924F7EA5F47ACE00" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.9</td>
<td id="0B1CB864E01DFAE2D783070A4D505B43" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.3</td>
<td id="F601B2132EEF99370E255520F0059F78" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.0</td>
</tr>
<tr id="8C60BA6AB5872D6C838A75C306A1C810" pageId="0" pageNumber="495">
<td id="3A2ED1FD133AC51756642FEBFC1B2066" colspan="1" pageId="0" pageNumber="495" rowspan="1">SNT</td>
<td id="9874ECB9613BA062AD63421E658AE23F" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.9</td>
<td id="E48CE748A4854FDB312BEAA6E119C3F2" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.7</td>
<td id="707EC3E09EE9FB2E28C4C816439E8F6A" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.5</td>
<td id="FBBA5984B55A825599E294CBDA0CA430" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.6</td>
<td id="C9B42A2A91CBF647D8A61D6D3839861C" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.3</td>
<td id="BA2A8546DF21D93104C617A80A9F815F" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.3</td>
<td id="9D0A75A7FE24B1C20C292B8FA511FC0B" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.9</td>
<td id="7A453ABCDA63C019ED6A97729495444F" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.9</td>
<td id="8C0B368D167C0E521EDF530F4ECF89FC" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.0</td>
<td id="38354B1B9F44A9D431087C6C35BD390E" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.5</td>
<td id="6BD99AE855A58DC3FD7A25BA8AA61A61" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.3</td>
</tr>
<tr id="5A5481D6E74FAA56F5C7511E977776B5" pageId="0" pageNumber="495">
<td id="C74664A41E7F7FE06FDC043549FBF7AC" colspan="1" pageId="0" pageNumber="495" rowspan="1">IND</td>
<td id="B13EA6B2E978F7C91FDCE2A7B8AE2977" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.9</td>
<td id="6921AEE41A8180DA4988AC3A8D014F4D" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.8</td>
<td id="E08765AC696CF7849591501C1B63B3DF" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
<td id="3F95FFB5900E8E13E18404620563BE70" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
<td id="A11339AC7322BBF4D32EF5FB9EB4DC40" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
<td id="29BC630E2981366D728EB283F1F2BD67" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.6</td>
<td id="8E17B7195BAB18868FCC62BD2E6B7B5E" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.7</td>
<td id="804B8C29E606AAFE44C149A87C50FECA" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.8</td>
<td id="6DBBD335E053B0777405C16BB10D6148" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.1</td>
<td id="6A683A2A2D38B547C1981C9321A8174D" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.2</td>
<td id="265D05537CF3A59B238D6755DD726564" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.3</td>
</tr>
<tr id="4FA25D213E78D7CFD35901DEEF620347" pageId="0" pageNumber="495">
<td id="D17E5EE3529CCC221AAA71D7E0DED6B4" colspan="1" pageId="0" pageNumber="495" rowspan="1">IOD</td>
<td id="F80BB85E75171E4130E196AA9CE05EED" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.5</td>
<td id="9D4FF749377488412098A132FEA0C14A" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.4</td>
<td id="2B7AFF131A3861F676D6CCE0CB406646" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.1</td>
<td id="6DE44890C8E0BADC4FC9DD7A922EBDF5" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.2</td>
<td id="E0F1A855BD7B07F02F39D20AF41B11CC" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.0</td>
<td id="6F0759F498F56220F50912981CD2411D" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.1</td>
<td id="4F1FF7644D6E7D5328659BC1D04FE4C4" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.3</td>
<td id="07B6D532D023D30EC691505F47CF9DD6" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.3</td>
<td id="6F13F55368BFF748BC65A6AB3FCB4250" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.7</td>
<td id="E03F63BBDC2DB0FA33EE5BB7040F3226" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.8</td>
<td id="C20858FEA1303B5DE9BBB50EFB5E5E26" colspan="1" pageId="0" pageNumber="495" rowspan="1">1.9</td>
</tr>
<tr id="EC02F0B3055F0808CFD91DDB30567182" pageId="0" pageNumber="495">
<td id="6B599A53A953754F5E8199F999891E0E" colspan="1" pageId="0" pageNumber="495" rowspan="1">ED</td>
<td id="A7D9ACC6030F2F8E14ED9EAA6EF7C277" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.6</td>
<td id="EF9AADF1A6095F50D6F45DFFAF5C8101" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.5</td>
<td id="FE1EF0D1E85659740641770D140C6491" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.5</td>
<td id="29D5261C1CC741BE70614A8A16486110" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.5</td>
<td id="815D5C5E021DF3856CE8E8705537D371" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.3</td>
<td id="47E9B60537958F51D175A8B8A008A4E8" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.3</td>
<td id="EEA6CCF37F603FB5B353065123DF5C12" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.5</td>
<td id="DF0E546257E222B348F540DBC1813621" colspan="1" pageId="0" pageNumber="495" rowspan="1">2.7</td>
<td id="287F0B3EFCBEC85B9F9A9EE25AFAD94F" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.3</td>
<td id="697D7A5C310C629B43D2E780FDA65130" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.4</td>
<td id="1B824B054C44955B6DA6CFBD2953CC91" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.0</td>
</tr>
<tr id="BE3C3BD8E3B3D153BBAAA50D196B0416" pageId="0" pageNumber="495">
<td id="3D9ADAF9128A474B0A3E73B1E15B47B5" colspan="1" pageId="0" pageNumber="495" rowspan="1">HND</td>
<td id="54103606D2D6CCAD7EAA44F961ADF0C0" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.4</td>
<td id="98E6A15212037D876FFED69B97775D27" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.3</td>
<td id="608D91512BA3837B71CD59958C2AA819" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.3</td>
<td id="933B42483DA02C38672B5C5F37D80B32" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.3</td>
<td id="2452939C591F656C2E8E0161FC7932D7" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.7</td>
<td id="E3530D5BEBAEC756297207E4FCD4D77F" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.6</td>
<td id="F5637FD6DC8492679CECFB0176039CF3" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.5</td>
<td id="649B0E6DC1F86796AA7F0F396EC38FD0" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.4</td>
<td id="D8ACBFDE43680E546AE1D4C2840298DA" colspan="1" pageId="0" pageNumber="495" rowspan="1">5.6</td>
<td id="BA9EDFB734364E4E868AE9B85660E4E3" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.8</td>
<td id="BB9E95C234A8BE5C823644AC208A8F54" colspan="1" pageId="0" pageNumber="495" rowspan="1">6.1</td>
</tr>
<tr id="F5251989ADB46D54D7B685504AB3D49A" pageId="0" pageNumber="495">
<td id="9EFFD148CD17749EDDBAF635638AB118" colspan="1" pageId="0" pageNumber="495" rowspan="1">RAD</td>
<td id="A5EA7CE7BF42D7C473638FD8C3721845" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.7</td>
<td id="43D446D9C4EF8F525EE15A7A42DADF1E" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.5</td>
<td id="18F935F5EA7040C750570176D8F80C2B" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.2</td>
<td id="2B6B04AF170BF06A84C54817AC4E7E9A" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.4</td>
<td id="A0B2D335E2FCED4616D255221AB1C547" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.5</td>
<td id="FB9A836622DC529E258B778F82C38DB3" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.7</td>
<td id="266CCEA05F97BD951B8B1C33250F4A33" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.4</td>
<td id="980C4ED8822E40EE8B292945473B8DE9" colspan="1" pageId="0" pageNumber="495" rowspan="1">3.3</td>
<td id="70C4C58AD2E3340B4683DA08E007244A" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.2</td>
<td id="81C4E4520BA4359C9499820E76FA1937" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.4</td>
<td id="D088BC64A5B69D9CA6CA54C38F0CBE02" colspan="1" pageId="0" pageNumber="495" rowspan="1">4.8</td>
</tr>
<tr id="4DBCC318BB05A598BC646B54E7A036DB" pageId="0" pageNumber="495">
<td id="254A0B49CBA3C156B41DC8101B934372" colspan="1" pageId="0" pageNumber="495" rowspan="1">FTL</td>
<td id="43A7EC417842FD5A259C03CEAA51C4E1" colspan="1" pageId="0" pageNumber="495" rowspan="1">14.1</td>
<td id="6A94D064CAC9BBDF7F74A70F4316FA15" colspan="1" pageId="0" pageNumber="495" rowspan="1">14.4</td>
<td id="75DA4EE6F647D43E1032EDCD3EDC8B59" colspan="1" pageId="0" pageNumber="495" rowspan="1">13.4</td>
<td id="3EC5052B8FE1EBE25364DA45919E44B6" colspan="1" pageId="0" pageNumber="495" rowspan="1">14.2</td>
<td id="E35ADDA7228ADB4155B3081A1FE5CE42" colspan="1" pageId="0" pageNumber="495" rowspan="1">12.9</td>
<td id="148F9366A792146F8578A6A751D2C89D" colspan="1" pageId="0" pageNumber="495" rowspan="1">13.8</td>
<td id="75084A7DDE674C47CF6C5E68F115732D" colspan="1" pageId="0" pageNumber="495" rowspan="1">14.7</td>
<td id="00160716B60E1A4F795BAAB8DB8768E4" colspan="1" pageId="0" pageNumber="495" rowspan="1">14.1</td>
<td id="E6FAA5E8FB36DEBFC6EB5E62D8500F90" colspan="1" pageId="0" pageNumber="495" rowspan="1">17.9</td>
<td id="50B8BA2845E8015216F49EBBE9D70D0A" colspan="1" pageId="0" pageNumber="495" rowspan="1">19.2</td>
<td id="EEED5E6DB22D86D2D1EC9E41CAE774CC" colspan="1" pageId="0" pageNumber="495" rowspan="1">19.5</td>
</tr>
<tr id="687896FC473B0E883463C4F9E1BA67B0" pageId="0" pageNumber="495">
<td id="30E95D6C10FF83F2BAA3C1FFD7DC8EEE" colspan="1" pageId="0" pageNumber="495" rowspan="1">TIB</td>
<td id="820BCEC1D80B3473B4DA371832223295" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.8</td>
<td id="D2F93E40144683E6556E4C19B817AB1B" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.7</td>
<td id="568B20AC2EEC2E2F5D647D8D369F94BB" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.4</td>
<td id="C115DCC068ABDF4E64EC66E72DB47AB8" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.8</td>
<td id="D3C98EF6C0F3EE87A08C1E1C4584A7CA" colspan="1" pageId="0" pageNumber="495" rowspan="1">8.6</td>
<td id="F226B8090BD43493ED73F389FB446840" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.2</td>
<td id="544D9EB7FBE54807E1E70E6DD28F519F" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.5</td>
<td id="3F34E6A8461DE64F98ED6B667C1A12CE" colspan="1" pageId="0" pageNumber="495" rowspan="1">9.5</td>
<td id="09F27F952DA7F2F9F28437B7CF18E713" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.7</td>
<td id="9DB79907B60342270C1F6141269B0565" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.4</td>
<td id="8C24B8427200565EC73E787DB0C24CF6" colspan="1" pageId="0" pageNumber="495" rowspan="1">11.8</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="CE0A6740666C551B4E45AEDE8192FEDB" pageId="0" pageNumber="495" type="description">
<paragraph id="BACE34D1BF8AB8D3A9F3D161C7B28C3E" pageId="0" pageNumber="495">Male secondary sexual characteristics.</paragraph>
<paragraph id="CF01E7382899B24CA4219C4155865F88" pageId="0" pageNumber="495">
Male with a single subgular vocal sac; in breeding season, a single, light yellow, swollen, and granular nuptial pad on the dorsal surface of finger I, nuptial spinules invisible. Males (SVL 19.9-22.1 mm) distinctly smaller than females (SVL 26.8-28.8 mm) (Fig.
<figureCitation id="66DCD3E26CFEBE7566A2AA08CD460E67" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of Occidozyga obscura comb. nov. and Occidozyga lingnanica sp. nov. in preservative. Occidozyga obscura comb. nov.: A adult male SYS a 008121 B adult female SYS a 008122; Occidozyga lingnanica sp. nov.: C adult male paratype SYS a 007373 D adult female paratype SYS a 007375. 1 dorsal view 2 ventral view." figureDoi="10.3897/vz.72.e80019.figure4" httpUri="https://binary.pensoft.net/fig/709317" pageId="0" pageNumber="495">4C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A7AA4812AAF2B1307F698D7B231991A0" pageId="0" pageNumber="495" type="ecology">
<paragraph id="716DCF9AC7AF58AB35645B15D455F70C" pageId="0" pageNumber="495">Ecology.</paragraph>
<paragraph id="5827EFF80587E83E6393041127628142" pageId="0" pageNumber="495">
This frog inhabits natural or artificial ponds and paddy fields in hilly regions. Males call in the water surface or waterside grass from dusk to dawn. The breeding season is from May to August (this study;
<bibRefCitation id="52A69F78945A8CEF62576A3703ED4FB0" author="Fei, L" journalOrPublisher="Current Herpetology" pageId="0" pageNumber="495" refId="B11" refString="Fei, L, Hu, SQ, Ye, CY, Huang, YZ, 2009. Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." title="Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, China." year="2009">Fei et al. 2009</bibRefCitation>
). In southeastern China,
<taxonomicName id="39262FF7932CB788F45D254AC59EFABA" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="02B85932B55EE90DE10D00F07756793F" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. shares a similar environment to that of
<taxonomicName id="121D5C62486B6ECF3202A3EFB83D1A89" authorityName="Lyu &amp; Wang &amp; Zeng &amp; Luo &amp; Zhang &amp; Guo &amp; Ren &amp; Qi &amp; Mo &amp; Wang" authorityYear="2022" baseAuthorityName="Gray" baseAuthorityYear="1831" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga obscura" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="obscura">
<emphasis id="C0AACF54F4C710DEF280A62F52CD5C85" italics="true" pageId="0" pageNumber="495">Occidozyga obscura</emphasis>
</taxonomicName>
comb. nov., but prefers to hilly regions compared with the latter in plain areas.
</paragraph>
</subSubSection>
<subSubSection id="650CD6EEA3BDBC20E7102827D50DE532" pageId="0" pageNumber="495" type="distribution">
<paragraph id="CEC39E1ED579F12F18B7B6625665D347" pageId="0" pageNumber="495">Distribution.</paragraph>
<paragraph id="E8351C291B71CCC1E737BA810FE5839A" pageId="0" pageNumber="495">
<taxonomicName id="D5C1BC48375C7454E86E8F7976E8B747" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="EABF0B032B774C3175735C5FDD4AFD16" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. can be recognized from multiple localities in Guangdong and Hainan of southeastern China based on the phylogenetic result in this work (Figs
<figureCitation id="02441DD7C73AE061E99359EB412C6D34" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic trees of the genus Occidozyga sensu lato based on mitochondrial genes 16 S and CO 1. Bayesian posterior probabilities (BPP) and bootstrap supports (BS) are labeled above and below the nodes respectively. Numbers at the terminal of the lineages correspond to the IDs in Table 1." figureDoi="10.3897/vz.72.e80019.figure1" httpUri="https://binary.pensoft.net/fig/709314" pageId="0" pageNumber="495">1</figureCitation>
,
<figureCitation id="6443289FBF5ECCA2320BF736199ED54C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the floating frogs Occidozyga obscura comb. nov., Occidozyga lingnanica sp. nov., and their closest congeners. Numbers correspond to the IDs in Table 1." figureDoi="10.3897/vz.72.e80019.figure2" httpUri="https://binary.pensoft.net/fig/709315" pageId="0" pageNumber="495">2</figureCitation>
). The records of
<taxonomicName id="90E20A8CDD868EC4F9E22B0E5FC64280" lsidName="O. martensi" pageId="0" pageNumber="495" rank="species" species="martensi">
<emphasis id="687071B52272E8D26BEA0041714CF6D8" italics="true" pageId="0" pageNumber="495">O. martensi</emphasis>
</taxonomicName>
from Yunnan, southwestern China require further clarifications with morphological examinations (see Discussion).
</paragraph>
</subSubSection>
<subSubSection id="8D6FC93DF39A240FB81EDD6920FB86F0" pageId="0" pageNumber="495" type="conservation status">
<paragraph id="AF0443130901208A846120855215C7B4" pageId="0" pageNumber="495">Conservation status.</paragraph>
<paragraph id="E885D48F3CB5FC0078E840322DDD3815" pageId="0" pageNumber="495">
<taxonomicName id="C00EED2512B5AB3123154CCD4BF9E893" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="71260D82C7ED9B7377EC8FB4530B2CA8" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. was previously reported as common and widespread species in southeastern China under the nomen
<taxonomicName id="43E1DCF39DDFAFFF84EF2DA6B8AF8E73" lsidName="O. martensii" pageId="0" pageNumber="495" rank="species" species="martensii">
<emphasis id="FDCCF524FBA3AF8F185833474D41C054" italics="true" pageId="0" pageNumber="495">O. martensii</emphasis>
</taxonomicName>
. Nevertheless, during our repeated surveys throughout southeastern China, the population quantity of this species is found rapidly declining due to the influence of human activities such as pesticide abuse and urban construction. We recommend
<taxonomicName id="FCD35E1175C51C0AB7D7E37AF8B8AF0B" authorityName="Lyu &amp; Wang" authorityYear="2022" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Occidozyga lingnanica" order="Anura" pageId="0" pageNumber="495" phylum="Chordata" rank="species" species="lingnanica">
<emphasis id="D64E4F23E12F11486F16E95C34B5182C" italics="true" pageId="0" pageNumber="495">Occidozyga lingnanica</emphasis>
</taxonomicName>
sp. nov. to be listed as Vulnerable (VU) [IUCN Red List criteria B1b(ii)(iii)].
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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