185 lines
18 KiB
XML
185 lines
18 KiB
XML
<document id="AA771EADEF76D0067E06290D7CDC9EDD" ID-DOI="10.11646/zootaxa.3866.1.5" ID-GBIF-Dataset="1c892016-2918-4b05-8eea-9032f9f7c6ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="227564" ID-ZooBank="A3620CFB-E223-479C-8A9B-FF84F30143CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460916005144" checkinUser="plazi" docAuthor="Nguyen, Tung T., Tran, Binh T. T. & Nguyen, Anh D." docDate="2014" docId="0F1A87E0FFF0FFB3A8CAFB6AF0F0F8C7" docLanguage="en" docName="zt03866p121.pdf" docOrigin="Zootaxa 3866 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polypheretima spiridonovi" docType="treatment" docVersion="10" lastPageNumber="116" masterDocId="F323FF98FFFBFFB8A85DFFF5F256FFBA" masterDocTitle="Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta" masterLastPageNumber="121" masterPageNumber="105" pageNumber="116" updateTime="1698361745542" updateUser="plazi">
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<mods:title id="BD59B06E1C159C4F33F39EBA970C1CAE">Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta</mods:title>
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<mods:namePart id="153749E9DB1721AC6047443CA0366173">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="2E425A5E733F3EACC49705C317DC96D7">Tran, Binh T. T.</mods:namePart>
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<mods:namePart id="834CB6C7A7B4A78A06049DBE966FD5C9">Nguyen, Anh D.</mods:namePart>
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<mods:title id="413EFA4EAB46BCF385E0890C44E0DC83">Zootaxa</mods:title>
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<treatment id="0F1A87E0FFF0FFB3A8CAFB6AF0F0F8C7" ID-DOI="http://doi.org/10.5281/zenodo.6144724" ID-GBIF-Taxon="127711698" ID-Zenodo-Dep="6144724" LSID="urn:lsid:plazi:treatment:0F1A87E0FFF0FFB3A8CAFB6AF0F0F8C7" httpUri="http://treatment.plazi.org/id/0F1A87E0FFF0FFB3A8CAFB6AF0F0F8C7" lastPageNumber="116" pageId="11" pageNumber="116">
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<subSubSection id="CFA9657DFFF0FFB3A8CAFB6AF031FA9A" pageId="11" pageNumber="116" type="nomenclature">
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<paragraph id="870C36F6FFF0FFB3A8CAFB6AF025FB00" blockId="11.[151,627,1183,1243]" box="[151,627,1183,1210]" pageId="11" pageNumber="116">
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<heading id="DC44819AFFF0FFB3A8CAFB6AF025FB00" bold="true" box="[151,627,1183,1210]" fontSize="11" level="1" pageId="11" pageNumber="116" reason="1">
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<taxonomicName id="40B34D75FFF0FFB3A8CAFB6AF025FB00" authority="(Thai, 1996)" baseAuthorityName="Thai" baseAuthorityYear="1996" box="[151,627,1183,1210]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="116" phylum="Annelida" rank="species" species="spiridonovi">
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<emphasis id="B5C7EAE4FFF0FFB3A8CAFB6AF025FB00" bold="true" box="[151,627,1183,1210]" pageId="11" pageNumber="116">
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<emphasis id="B5C7EAE4FFF0FFB3A8CAFB6AF383FB03" bold="true" box="[151,469,1183,1209]" italics="true" pageId="11" pageNumber="116">Polypheretima spiridonovi</emphasis>
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(
|
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<bibRefCitation id="E3224B07FFF0FFB3A9B8FB55F03DFB00" author="Thai" box="[485,619,1184,1210]" pageId="11" pageNumber="116" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
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||
<collectingCountry id="FFA47666FFF0FFB3A9B8FB55F074FB00" box="[485,546,1184,1210]" name="Thailand" pageId="11" pageNumber="116">Thai</collectingCountry>
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, 1996
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</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="870C36F6FFF0FFB3A8CAFB37F2BDFB61" blockId="11.[151,627,1183,1243]" box="[151,235,1218,1243]" pageId="11" pageNumber="116">
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(
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<figureCitation id="1F882A73FFF0FFB3A8C2FB37F2B4FB61" box="[159,226,1218,1243]" captionStart="FIGURE 3" captionStartId="8.[151,250,1852,1874]" captionTargetBox="[151,1436,241,1824]" captionTargetId="figure@8.[151,1436,195,1830]" captionTargetPageId="8" captionText="FIGURE 3. Distribution of species of the genus Polypheretima in Vietnam. The exotic Po. elongata with many records all over the country is omitted." httpUri="https://zenodo.org/record/227567/files/figure.png" pageId="11" pageNumber="116">Fig. 3</figureCitation>
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)
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</paragraph>
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<paragraph id="870C36F6FFF0FFB3A8CAFAFFF031FA9A" blockId="11.[151,861,1290,1343]" box="[151,615,1290,1312]" pageId="11" pageNumber="116">
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<treatmentCitationGroup id="A7A311D8FFF0FFB3A8CAFAFFF031FA9A" box="[151,615,1290,1312]" pageId="11" pageNumber="116">
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<taxonomicName id="40B34D75FFF0FFB3A8CAFAFFF05EFA9A" authority="Thai, 1996: 3" authorityName="Thai" authorityPageNumber="3" authorityYear="1996" box="[151,520,1290,1312]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="116" phylum="Annelida" rank="species" species="spiridonovi">
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<emphasis id="B5C7EAE4FFF0FFB3A8CAFAFFF32FFA9A" box="[151,377,1290,1312]" italics="true" pageId="11" pageNumber="116">Pheretima spiridonovi</emphasis>
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<treatmentCitation id="061210E7FFF0FFB3A922FAFFF05EFA9A" author="Thai" box="[383,520,1290,1312]" page="3" pageId="11" pageNumber="116" year="1996">
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<bibRefCitation id="E3224B07FFF0FFB3A922FAFFF3A6FA9A" author="Thai" box="[383,496,1290,1312]" pageId="11" pageNumber="116" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
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<collectingCountry id="FFA47666FFF0FFB3A922FAFFF3E6FA9A" box="[383,432,1290,1312]" name="Thailand" pageId="11" pageNumber="116">Thai</collectingCountry>
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, 1996
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</bibRefCitation>
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||
: 3
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</treatmentCitation>
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</taxonomicName>
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,
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<figureCitation id="1F882A73FFF0FFB3AA4EFAFFF006FA9A" box="[531,592,1290,1312]" captionStart="FIGURE 2" captionStartId="5.[151,250,1690,1712]" captionTargetBox="[151,1436,241,1658]" captionTargetId="figure@5.[151,1436,195,1668]" captionTargetPageId="5" captionText="FIGURE 2. Distribution of species of the genus Planapheretima in Vietnam." httpUri="https://zenodo.org/record/227566/files/figure.png" pageId="11" pageNumber="116">Fig. 2</figureCitation>
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C.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="CFA9657DFFF0FFB3A8CAFADCF10BFA85" box="[151,861,1321,1343]" pageId="11" pageNumber="116" type="reference_group">
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<paragraph id="870C36F6FFF0FFB3A8CAFADCF10BFA85" blockId="11.[151,861,1290,1343]" box="[151,861,1321,1343]" pageId="11" pageNumber="116">
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<treatmentCitationGroup id="A7A311D8FFF0FFB3A8CAFADCF10BFA85" box="[151,861,1321,1343]" pageId="11" pageNumber="116">
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<taxonomicName id="40B34D75FFF0FFB3A8CAFADCF3F2FA85" box="[151,420,1321,1343]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="116" phylum="Annelida" rank="species" species="spiridonovi">
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<emphasis id="B5C7EAE4FFF0FFB3A8CAFADCF3F2FA85" box="[151,420,1321,1343]" italics="true" pageId="11" pageNumber="116">Polypheretima spiridonovi</emphasis>
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</taxonomicName>
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||
—
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<treatmentCitation id="061210E7FFF0FFB3A9E3FADCF02AFA85" author="Nakamura" box="[446,636,1321,1343]" page="7" pageId="11" pageNumber="116" year="1999">
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||
<bibRefCitation id="E3224B07FFF0FFB3A9E3FADCF032FA85" author="Nakamura" box="[446,612,1321,1343]" pageId="11" pageNumber="116" refString="Nakamura, Y. (1999) Checklist of earthworms of Pheretima genus group (Megascolecidae: Oligochaeta) of the world. Edaphologia, 64, 1 - 78." type="journal article" year="1999">Nakamura 1999</bibRefCitation>
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: 7
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</treatmentCitation>
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||
;
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<treatmentCitation id="061210E7FFF0FFB3AAD5FADCF10FFA85" author="Blakemore" box="[648,857,1321,1343]" page="93" pageId="11" pageNumber="116" year="2007">
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<bibRefCitation id="E3224B07FFF0FFB3AAD5FADCF165FA85" author="Blakemore" box="[648,819,1321,1343]" pageId="11" pageNumber="116" refString="Blakemore, R. J. (2007) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. Available from: www. annelida. net / earthworm / Pheretimoids. pdf (accessed 26 August 2014)" type="journal article" year="2007">Blakemore 2007</bibRefCitation>
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: 93
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFA9657DFFF0FFB3A8CAFA99F685FA12" pageId="11" pageNumber="116" type="materials_examined">
|
||
<paragraph id="870C36F6FFF0FFB3A8CAFA99F685FA12" blockId="11.[151,1437,1388,1917]" pageId="11" pageNumber="116">
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||
<emphasis id="B5C7EAE4FFF0FFB3A8CAFA99F31AFA3F" bold="true" box="[151,332,1388,1413]" pageId="11" pageNumber="116">
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<typeStatus id="58088854FFF0FFB3A8CAFA99F284FA3F" box="[151,210,1388,1413]" pageId="11" pageNumber="116">Type</typeStatus>
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material.
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||
</emphasis>
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<typeStatus id="58088854FFF0FFB3A906FA99F39EFA3F" box="[347,456,1388,1413]" pageId="11" pageNumber="116" type="holotype">Holotype</typeStatus>
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: 1C (SORC-HV.226) Khanh Hoa Province,
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<collectingCountry id="FFA47666FFF0FFB3ABBFFA98F646FA3E" box="[994,1040,1389,1412]" name="Germany" pageId="11" pageNumber="116">Duc</collectingCountry>
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Mu pass,
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<date id="F30D1036FFF0FFB3ACC6FA99F70EFA3E" box="[1179,1368,1388,1413]" pageId="11" pageNumber="116" value="1989-01-09">9 January 1989</date>
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, coll. Spiridonov S.E.
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<typeStatus id="58088854FFF0FFB3A90DFA65F3E0FA12" box="[336,438,1424,1448]" pageId="11" pageNumber="116" type="paratype">Paratype</typeStatus>
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: 1C (SORC-PV.226), same data as sample SORC-HV.226 (
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<bibRefCitation id="E3224B07FFF0FFB3AC12FA7AF690FA12" author="Thai" box="[1103,1222,1423,1448]" pageId="11" pageNumber="116" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
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<collectingCountry id="FFA47666FFF0FFB3AC12FA7AF6D2FA12" box="[1103,1156,1423,1448]" name="Thailand" pageId="11" pageNumber="116">Thai</collectingCountry>
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||
1996
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CFA9657DFFF0FFB3A89AFA41F1D3F956" pageId="11" pageNumber="116" type="description">
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<paragraph id="870C36F6FFF0FFB3A89AFA41F10DF972" blockId="11.[151,1437,1388,1917]" pageId="11" pageNumber="116">
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<emphasis id="B5C7EAE4FFF0FFB3A89AFA41F30FFA77" bold="true" box="[199,345,1460,1485]" pageId="11" pageNumber="116">Description.</emphasis>
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Body cylindrical. Colour light brown. Prostomium epilobous. Clitellum XIV–XVI. First dorsal pore in 12/13. Setae numerous, regularly distributed on segments,
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<quantity id="404B9B13FFF0FFB3AB23FA2DF192FA4A" box="[894,964,1495,1520]" metricMagnitude="0" metricUnit="m" metricValue="2.8701999999999996" pageId="11" pageNumber="116" unit="in" value="113.0">113 in</quantity>
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IV,
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<quantity id="404B9B13FFF0FFB3ABAFFA22F66FFA4A" box="[1010,1081,1495,1520]" metricMagnitude="0" metricUnit="m" metricValue="3.4543999999999997" pageId="11" pageNumber="116" unit="in" value="136.0">136 in</quantity>
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||
XIII,
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<quantity id="404B9B13FFF0FFB3AC23FA2DF693FA4A" box="[1150,1221,1495,1520]" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="11" pageNumber="116" unit="in" value="120.0">120 in</quantity>
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||
XXV, aa=1.5–2ab, zz=2–5zy.
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||
<collectingCountry id="FFA47666FFF0FFB3A949FA09F318F9AF" box="[276,334,1532,1557]" name="Malta" pageId="11" pageNumber="116">Male</collectingCountry>
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||
porophore on the bottom of a cone-like depression, about 0.25 body circumference apart. No genital markings in spermathecal and male regions. Numerous spermathecal pores in paired batteries, 6 pairs in 4/ 5, 6 pairs in 6/7 and 7 pairs and a single battery in 5/6. Ampulla heart-shaped, diverticula simple, attaching to duct. Holandric. Testes sacs ventral. Lymph glands absent. Ovaries large on septum 12/13. Intestine beginning at XV, without caeca. Last hearts in XII. Typhlosole simple, lamelliform, T=0.22.
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<collectingCountry id="FFA47666FFF0FFB3ABBBF979F672F91E" box="[998,1060,1676,1700]" name="Spain" pageId="11" pageNumber="116">Septa</collectingCountry>
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||
<date id="F30D1036FFF0FFB3AC71F979F625F91E" box="[1068,1139,1676,1700]" pageId="11" pageNumber="116" value="2010-09-08">8/9/10</date>
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absent, 5/6/7/8 thickened. Pharyngeal micronephridia in 6/7. (Data from
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<bibRefCitation id="E3224B07FFF0FFB3AAC2F95AF143F972" author="Thai" box="[671,789,1711,1736]" pageId="11" pageNumber="116" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
|
||
<collectingCountry id="FFA47666FFF0FFB3AAC2F95AF084F972" box="[671,722,1711,1736]" name="Thailand" pageId="11" pageNumber="116">Thai</collectingCountry>
|
||
1996
|
||
</bibRefCitation>
|
||
: 3–4.)
|
||
</paragraph>
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||
<paragraph id="870C36F6FFF0FFB3A89AF921F1D3F956" blockId="11.[151,1437,1388,1917]" box="[199,901,1748,1773]" pageId="11" pageNumber="116">
|
||
<emphasis id="B5C7EAE4FFF0FFB3A89AF921F388F957" bold="true" box="[199,478,1748,1773]" pageId="11" pageNumber="116">
|
||
Records from
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||
<collectingCountry id="FFA47666FFF0FFB3A92CF921F38FF957" box="[369,473,1748,1773]" name="Vietnam" pageId="11" pageNumber="116">Vietnam</collectingCountry>
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||
.
|
||
</emphasis>
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||
Khanh Hoa Province (
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<collectingCountry id="FFA47666FFF0FFB3AA82F921F159F956" box="[735,783,1748,1772]" name="Germany" pageId="11" pageNumber="116">Duc</collectingCountry>
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||
Mu pass).
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="CFA9657DFFF0FFB3A89AF902F0CBF8AA" box="[199,669,1783,1808]" pageId="11" pageNumber="116" type="distribution">
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||
<paragraph id="870C36F6FFF0FFB3A89AF902F0CBF8AA" blockId="11.[151,1437,1388,1917]" box="[199,669,1783,1808]" pageId="11" pageNumber="116">
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||
<emphasis id="B5C7EAE4FFF0FFB3A89AF902F336F8AA" bold="true" box="[199,352,1783,1808]" pageId="11" pageNumber="116">Distribution.</emphasis>
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||
Only known from
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||
<collectingCountry id="FFA47666FFF0FFB3AA6AF902F0CCF8AA" box="[567,666,1783,1808]" name="Vietnam" pageId="11" pageNumber="116">Vietnam</collectingCountry>
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||
.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFA9657DFFF0FFB3A89AF8E9F0F0F8C7" pageId="11" pageNumber="116" type="discussion">
|
||
<paragraph id="870C36F6FFF0FFB3A89AF8E9F0F0F8C7" blockId="11.[151,1437,1388,1917]" pageId="11" pageNumber="116">
|
||
<emphasis id="B5C7EAE4FFF0FFB3A89AF8E9F36DF88F" bold="true" box="[199,315,1820,1845]" pageId="11" pageNumber="116">Remarks.</emphasis>
|
||
The species is fairly similar to
|
||
<taxonomicName id="40B34D75FFF0FFB3AAE6F8E9F6C9F88E" authority="Beddard, 1900" authorityName="Beddard" authorityYear="1900" box="[699,1183,1820,1845]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="116" phylum="Annelida" rank="species" species="polytheca">
|
||
<emphasis id="B5C7EAE4FFF0FFB3AAE6F8E9F18EF88E" box="[699,984,1820,1844]" italics="true" pageId="11" pageNumber="116">Polypheretima polytheca</emphasis>
|
||
(Beddard, 1900)
|
||
</taxonomicName>
|
||
but differs in having spermathecal pores in 4/5–6/7, testes sacs in ventral position, absence of a copulatory pouch, and absence of genital markings.
|
||
<typeStatus id="58088854FFF0FFB3A950F890F319F8C7" box="[269,335,1893,1917]" pageId="11" pageNumber="116">Types</typeStatus>
|
||
were rechecked for this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |