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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.171.2273" ID-PMC="PMC3288681" ID-Pensoft-Pub="1313-2970-171-17" ID-Pensoft-UUID="62463D177D1EFFB0FFEB073EFFA9FFF5" ID-PubMed="22423194" ID-Zenodo-Dep="576912" ID-ZooBank="91A0AA5641074E29B847BA6924FB7BC8" ModsDocID="1313-2970-171-17" checkinTime="1451249329658" checkinUser="pensoft" docAuthor="Guemues, Burcin Askim &amp; Neubert, Eike" docDate="2012" docId="0EAE017979EAF1E59D44B0EC71853538" docLanguage="en" docName="ZooKeys 171: 17-37" docOrigin="ZooKeys 171" docPubDate="2012-02-24" docSource="http://dx.doi.org/10.3897/zookeys.171.2273" docTitle="Meijeriella frivaldskyi" docType="treatment" docVersion="5" id="62463D177D1EFFB0FFEB073EFFA9FFF5" lastPageNumber="23" masterDocId="62463D177D1EFFB0FFEB073EFFA9FFF5" masterDocTitle="New taxa of terrestrial molluscs from Turkey (Gastropoda, Pristilomatidae, Enidae, Hygromiidae, Helicidae)" masterLastPageNumber="37" masterPageNumber="17" pageNumber="23" updateTime="1668153260529" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New taxa of terrestrial molluscs from Turkey (Gastropoda, Pristilomatidae, Enidae, Hygromiidae, Helicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guemues, Burcin Askim</mods:namePart>
<mods:affiliation>Burcin Askim Guemues, Gazi University, Science Faculty, Department of Biology (Zoology), Teknik Okullar, Besevler, 06500, Ankara, Tuerkiye</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubert, Eike</mods:namePart>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, Bernastr. 15, CH- 3005 Bern, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-02-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>171</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>37</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.171.2273</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.171.2273</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-171-17</mods:identifier>
<mods:identifier type="ZooBank">91A0AA5641074E29B847BA6924FB7BC8</mods:identifier>
<mods:identifier type="Pensoft-UUID">62463D177D1EFFB0FFEB073EFFA9FFF5</mods:identifier>
<mods:identifier type="Zenodo-Dep">576912</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033352" LSID="urn:lsid:plazi:treatment:0EAE017979EAF1E59D44B0EC71853538" httpUri="http://treatment.plazi.org/id/0EAE017979EAF1E59D44B0EC71853538" lastPageNumber="23" pageId="6" pageNumber="23">
<subSubSection pageId="6" pageNumber="23" type="nomenclature">
<paragraph pageId="6" pageNumber="23">
<taxonomicName LSID="0EAE0179-79EA-F1E5-9D44-B0EC71853538" authority="(L. Pfeiffer, 1847)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1847" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meijeriella frivaldskyi" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="frivaldskyi">
<pageBreakToken pageId="6" pageNumber="23" start="start">Meijeriella</pageBreakToken>
frivaldskyi (L. Pfeiffer, 1847)
</taxonomicName>
<figureCitation captionStart="Figure 23" captionStartId="F2" captionText="Figure 2 - 3. 2 Genital organs of Meijeriella canaliculata (Kaz Dagi) 3 Genital organs of Meijeriella frivaldskyi (ruins of Truva)." httpUri="https://binary.pensoft.net/fig/14966" pageId="6" pageNumber="23">Figs 3</figureCitation>
<figureCitation captionStart="Figures 412" captionStartId="F3" captionText="Figures 4 - 12. Enidae. 4 - 10 Meijeriella spp. 4 Meijeriella canaliculata, Vil. Balikesir, Edremit, Kaz Dagi, Camp Area, 39.67 ° N, 26.95 ° E, H = 14.65 mm. 5 - 8 Meijeriella frivaldskyi. 5 syntype Bulimus frivaldskyi, SMF 63750, H = 14.8 mm 6 syntype Bulimus raynevalianus, MHNG 12106, H = 15.8 mm 7 Meijeriella frivaldskyi, SMF 312495, Ruins of Truva, leg. H. P. M. G. Menkhorst, H = 11.45 mm 8 Meijeriella frivaldskyi, NMBE 28357, Ruins of Truva, leg. B. A. Guemues, 10.06.2008, H = 10.6 mm. 9 - 11 Turanena spp. 9. Turanena demirsoyi sp. n., holotype NMBE 32704, Erzincan, Kemaliye, Dilli Deresi, 870 m alt., leg. B. A. Guemues, H = 12.15 mm 10 Turanenea cochlicopoides, Paratype SMF 309776, Vil. Gaziantep, along the road Gaziantep → Bahce, ca. 12 km NW Gaziantep, 1000 m alt., leg. Menkhorst, 15.07.1986, H = 5.6 mm 11 Turanena cochlicopoides; NMBE 32713, between Arslanoba and Dolunay villages (the rocks around the Cahit Bilgin Park), 39.249 ° N, 38.599 ° E, 1560 m alt., leg. B. A. Guemues, 07.09.2007 12 Buliminus alepensis alepensis NMBE 32709, Cit Koeyue, 39.1119 ° N, 38.6043 ° E, 976 m alt. 05.05.2007, leg. B. A. Guemues. - All figures scaled x 5." httpUri="https://binary.pensoft.net/fig/14968" pageId="6" pageNumber="23">5-8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="reference_group">
<paragraph pageId="6" pageNumber="23">
<taxonomicName class="Gastropoda" family="Bithyniidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="frivaldskyi">Bulimus frivaldskyi</taxonomicName>
L. Pfeiffer, Zeitschrift
<normalizedToken originalValue="für">fuer</normalizedToken>
Malakozoologie, 1847, 4 (12): 191-192 [Type locality: &quot;prope Brussa Natoliae&quot;].
</paragraph>
<paragraph pageId="6" pageNumber="23">
<taxonomicName class="Gastropoda" family="Bithyniidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="fuscus">Bulimus fuscus</taxonomicName>
L. Pfeiffer, Zeitschr. Malakozool., 1847, 4 (12): 192. Type locality: not given. Nomen nudum (in synonymy).
</paragraph>
<paragraph pageId="6" pageNumber="23">
<taxonomicName class="Gastropoda" family="Bithyniidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="raynevalianus">Bulimus raynevalianus</taxonomicName>
Bourguignat, Amen. Malac. 1855, 29: 128, pl. 8 fig. 5-9 (shell). Type locality: &quot;les vignes des environs de Gallipoli,
<normalizedToken originalValue="oú">ou</normalizedToken>
il vit sous les mottes de terre&quot;.
</paragraph>
<paragraph pageId="6" pageNumber="23">
<taxonomicName class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="friwaldskyi" subGenus="Brephulus">Buliminus [Brephulus] friwaldskyi</taxonomicName>
f.
<emphasis italics="true" pageId="6" pageNumber="23">minor</emphasis>
Westerlund, 1887, Fauna pal. Reg. Binnenconch., 3: 10. Type locality: not given.
</paragraph>
<paragraph pageId="6" pageNumber="23">
<taxonomicName class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="trojanus">Buliminus trojanus</taxonomicName>
Kobelt, Icon., 1893, (2) 6 (5-6): 81, pl. 171 fig. 1100 (shell).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="type locality">
<paragraph pageId="6" pageNumber="23">Type locality.</paragraph>
<paragraph pageId="6" pageNumber="23">&quot;im westlichen Theile der Troas&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="type specimens">
<paragraph pageId="6" pageNumber="23">Type specimens.</paragraph>
<paragraph pageId="6" pageNumber="23">
<emphasis italics="true" pageId="6" pageNumber="23">frivaldskyi</emphasis>
: Syntype SMF 63750,
<normalizedToken originalValue="“Bursa”">&quot;Bursa&quot;</normalizedToken>
, coll. Kobelt ex L. Pfeiffer;
<emphasis italics="true" pageId="6" pageNumber="23">raynevalianus</emphasis>
: Syntype MHNG 12106.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="specimens examined">
<paragraph pageId="6" pageNumber="23">Specimens examined.</paragraph>
<paragraph pageId="6" pageNumber="23">
Vil.
<normalizedToken originalValue="Çanakkale">Canakkale</normalizedToken>
, Truva, 10.06.2003, leg. B. A.
<normalizedToken originalValue="Gümüş">Guemues</normalizedToken>
, NMBE 28357/3 (preserved).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="description of the genital organs">
<paragraph pageId="6" pageNumber="23">Description of the genital organs</paragraph>
<paragraph pageId="6" pageNumber="23">
(
<figureCitation captionStart="Figure 23" captionStartId="F2" captionText="Figure 2 - 3. 2 Genital organs of Meijeriella canaliculata (Kaz Dagi) 3 Genital organs of Meijeriella frivaldskyi (ruins of Truva)." httpUri="https://binary.pensoft.net/fig/14966" pageId="6" pageNumber="23">Fig. 3</figureCitation>
, shell size of dissected animal: 10.6
<normalizedToken originalValue="×">x</normalizedToken>
3.7 mm)
<emphasis bold="true" pageId="6" pageNumber="23">.</emphasis>
Penis without penial appendix, subdivided in a distal and proximal section; proximal section containing a centrally perforated verge; epiphallus very long, reaching ca. 5
<normalizedToken originalValue="×">x</normalizedToken>
the length of penis, with a small caecum at its central part, a short flagellum present; penial retractor muscle attaching at the border of penis and epiphallus; vagina as long as penis, pedunculus with a long diverticulum largely surmounting the bursa copulatrix in length.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="23" type="remarks">
<paragraph pageId="6" pageNumber="23">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="23">
The principal morphological structure of the genital organs does not differ from the holotype of
<taxonomicName authorityName="Bank" authorityYear="1985" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="canaliculata">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella canaliculata</emphasis>
</taxonomicName>
(cf.
<bibRefCitation author="Bank, RA" journalOrPublisher="Heldia" pageId="9" pageNumber="26" pagination="41 - 44" refId="B1" refString="Bank, RA, 1985. Eine neue Enide von der griechischen Insel Mytilini (Gastropoda: Pupillacea). Heldia 1 (2): 41 - 44" title="Eine neue Enide von der griechischen Insel Mytilini (Gastropoda: Pupillacea)." volume="1" year="1985">Bank 1985</bibRefCitation>
) from the Island of Lesvos nor from that from the southern slope of the Kaz
<normalizedToken originalValue="Dağı">Dagi</normalizedToken>
(
<figureCitation captionStart="Figure 23" captionStartId="F2" captionText="Figure 2 - 3. 2 Genital organs of Meijeriella canaliculata (Kaz Dagi) 3 Genital organs of Meijeriella frivaldskyi (ruins of Truva)." httpUri="https://binary.pensoft.net/fig/14966" pageId="6" pageNumber="23">Fig. 2</figureCitation>
). For this reason, the species
<taxonomicName class="Gastropoda" family="Bithyniidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="frivaldskyi">
<emphasis italics="true" pageId="6" pageNumber="23">Bulimus frivaldskyi</emphasis>
</taxonomicName>
is here confined to the genus
<taxonomicName authorityName="Bank" authorityYear="1985" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella</emphasis>
</taxonomicName>
and removed from the genus
<taxonomicName class="Gastropoda" family="Enidae" genus="Ena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="6" pageNumber="23" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="23">Ena</emphasis>
</taxonomicName>
, as already suspected by other authors (
<bibRefCitation author="Bank, RA" journalOrPublisher="Heldia" pageId="9" pageNumber="26" pagination="41 - 44" refId="B1" refString="Bank, RA, 1985. Eine neue Enide von der griechischen Insel Mytilini (Gastropoda: Pupillacea). Heldia 1 (2): 41 - 44" title="Eine neue Enide von der griechischen Insel Mytilini (Gastropoda: Pupillacea)." volume="1" year="1985">Bank 1985</bibRefCitation>
: 42,
<bibRefCitation author="Hausdorf, B" journalOrPublisher="Journal of Natural History" pageId="10" pageNumber="27" pagination="1627 - 1638" publicationUrl="10.1080/002229301317092360" refId="B12" refString="Hausdorf, B, 2001. The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35 (11): 1627 - 1638, 10.1080/002229301317092360" title="The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae)." url="10.1080/002229301317092360" volume="35" year="2001">Hausdorf 2001</bibRefCitation>
). The shell of
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1847" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="frivaldskyi">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella frivaldskyi</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Bank" authorityYear="1985" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="canaliculata">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella canaliculata</emphasis>
</taxonomicName>
by its rounded and not canaliculated last whorl; additionally, the majority of specimens of
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1847" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="frivaldskyi">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella frivaldskyi</emphasis>
</taxonomicName>
differ by presence of a palatal swelling of the labium, which is said to miss in the Lesvian specimens of
<taxonomicName authorityName="Bank" authorityYear="1985" class="Gastropoda" family="Enidae" genus="Meijeriella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="6" pageNumber="23" phylum="Mollusca" rank="species" species="canaliculata">
<emphasis italics="true" pageId="6" pageNumber="23">Meijeriella canaliculata</emphasis>
</taxonomicName>
(and which is also true for the Kaz
<normalizedToken originalValue="Dağı">Dagi</normalizedToken>
population).
</paragraph>
</subSubSection>
</treatment>
</document>