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<document ID-DOI="http://dx.doi.org/10.3897/aiep.52.89340" ID-Pensoft-Pub="1734-1515-3-215" ID-Pensoft-UUID="A2AB5425B81B546997A7FB8906135FA2" ID-ZooBank="850A82A7B7C744A695E07583DA61EB11" ModsDocID="1734-1515-52-3-215" checkinTime="1663958826690" checkinUser="pensoft" docAuthor="Borg, Joseph A., Evans, Julian, Knittweis, Leyla &amp; Schembri, Patrick J." docDate="2022" docId="0E98F02F67D65CEFA98B7A53E8F5F3F9" docLanguage="en" docName="ActIchthPisc 52(3): 215-221" docOrigin="Acta Ichthyologica et Piscatoria 52 (3)" docPubDate="2022-09-23" docSource="http://dx.doi.org/10.3897/aiep.52.89340" docTitle="Thorogobius ephippiatus" docType="treatment" docVersion="2" id="A2AB5425B81B546997A7FB8906135FA2" lastPageNumber="215" masterDocId="A2AB5425B81B546997A7FB8906135FA2" masterDocTitle="New and interesting records of marine fishes (Actinopterygii) from the Maltese Islands (central Mediterranean)" masterLastPageNumber="221" masterPageNumber="215" pageNumber="215" updateTime="1663959127312" updateUser="ExternalLinkService">
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<mods:title>New and interesting records of marine fishes (Actinopterygii) from the Maltese Islands (central Mediterranean)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Borg, Joseph A.</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, University of Malta, Msida, Malta</mods:affiliation>
<mods:nameIdentifier type="email">joseph.a.borg@um.edu.mt</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Evans, Julian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7837-5927</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Faculty of Science, University of Malta, Msida, Malta</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Knittweis, Leyla</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, University of Malta, Msida, Malta &amp; Fisheries New Zealand, Ministry for Primary Industries, Wellington 6011, New Zealand</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schembri, Patrick J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6723-7198</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Faculty of Science, University of Malta, Msida, Malta</mods:affiliation>
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<mods:title>Acta Ichthyologica et Piscatoria</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-09-23</mods:number>
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<mods:number>52</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/aiep.52.89340</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0E98F02F67D65CEFA98B7A53E8F5F3F9" httpUri="http://treatment.plazi.org/id/0E98F02F67D65CEFA98B7A53E8F5F3F9" lastPageNumber="215" pageId="0" pageNumber="215">
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<taxonomicName LSID="0E98F02F-67D6-5CEF-A98B-7A53E8F5F3F9" authority="(Lowe, 1839)" baseAuthorityName="Lowe" baseAuthorityYear="1839" class="Actinopterygii" family="Gobiidae" genus="Thorogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thorogobius ephippiatus" order="Perciformes" pageId="0" pageNumber="215" phylum="Chordata" rank="species" species="ephippiatus">Thorogobius ephippiatus (Lowe, 1839)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="215" type="description">
<paragraph pageId="0" pageNumber="215">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photograph of Thorogobius ephippiatus, taken on 27 June 2016 at a depth of 24 m within a cave (36 ° 04.879 N, 014 ° 14.110 E). Photograph: OCEANA / Carlos Minguell © LIFE BaĦAR for N 2 K." figureDoi="10.3897/aiep.52.89340.figure1" httpUri="https://binary.pensoft.net/fig/747670" pageId="0" pageNumber="215">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Image grab from ROV footage collected on 7 July 2016 at a depth of 117 m (36 ° 00.776 N, 014 ° 50.517 E) showing an individual of Thorogobius ephippiatus. Image: OCEANA © LIFE BaĦAR for N 2 K." figureDoi="10.3897/aiep.52.89340.figure2" httpUri="https://binary.pensoft.net/fig/747671" pageId="0" pageNumber="215">, 2</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="215" type="diagnosis">
<paragraph pageId="0" pageNumber="215">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="215">Base coloration greyish with blue-green sheen on back; covered with large, round dark spots. Head, including predorsal area, covered with brown to dark orange, smaller round spots, usually lighter in colour and more reddish than blotches on body. Five to six large, dark brown or brown-purple to black midlateral blotches. Dark blotches above midline smaller than midlateral blotches.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="215" type="remarks">
<paragraph pageId="0" pageNumber="215">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="215">
In the SCUBA surveys there were six occurrences of this species from depths of between 8 and 32 m within caves, with four occurrences on soft sediment bottoms and two on rock (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Individuals of Thorogobius ephippiatus recorded from the LIFE BaĦAR for N 2 K marine benthic surveys." httpUri="http://table.plazi.org/id/9290C6EC705EBC6F33E4295155EE47FA" pageId="0" pageNumber="215" tableUuid="9290C6EC705EBC6F33E4295155EE47FA">1</tableCitation>
). An additional single record from the ROV surveys was from a depth of 117 m on a rocky bottom having a thin layer of silt (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Individuals of Thorogobius ephippiatus recorded from the LIFE BaĦAR for N 2 K marine benthic surveys." httpUri="http://table.plazi.org/id/9290C6EC705EBC6F33E4295155EE47FA" pageId="0" pageNumber="215" tableUuid="9290C6EC705EBC6F33E4295155EE47FA">1</tableCitation>
and Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Image grab from ROV footage collected on 7 July 2016 at a depth of 117 m (36 ° 00.776 N, 014 ° 50.517 E) showing an individual of Thorogobius ephippiatus. Image: OCEANA © LIFE BaĦAR for N 2 K." figureDoi="10.3897/aiep.52.89340.figure2" httpUri="https://binary.pensoft.net/fig/747671" pageId="0" pageNumber="215">2</figureCitation>
). No cave or rock fissure or any other similar feature was present in the vicinity of this individual. Given that the literature indicates that
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<emphasis italics="true" pageId="0" pageNumber="215">T. ephippiatus</emphasis>
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is typical of cave habitats, crevices, overhangs or deep gullies (
<bibRefCitation DOI="https://doi.org/10.1016/j.ecss.2008.09.023" author="Bussotti, S" journalOrPublisher="Cybium" pageId="0" pageNumber="215" refId="B5" refString="Bussotti, S, Guidetti, P, 2009. Do Mediterranean fish assemblages associated with marine caves and rocky cliffs differ? Estuarine, Coastal and Shelf Science 81(1): 65-73. https://doi.org/10.1016/j.ecss.2008.09.023" title="Do Mediterranean fish assemblages associated with marine caves and rocky cliffs differ? Estuarine, Coastal and Shelf Science 81 (1): 65 - 73." url="https://doi.org/10.1016/j.ecss.2008.09.023" year="2009">Bussotti and Guidetti 2009</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0122632" author="Bussotti, S" journalOrPublisher="Cybium" pageId="0" pageNumber="215" refId="B6" refString="Bussotti, S, Di Franco, A, Francour, P, Guidetti, P, 2015. Fish assemblages of Mediterranean marine caves. PLoS One 10(4): e0122632. https://doi.org/10.1371/journal.pone.0122632" title="Fish assemblages of Mediterranean marine caves. PLoS One 10 (4): e 0122632." url="https://doi.org/10.1371/journal.pone.0122632" year="2015">Bussotti et al. 2015</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3390/jmse9060557" author="Ragkousis, M" journalOrPublisher="Journal of Marine Science and Engineering" pageId="0" pageNumber="215" pagination=": 557" refId="B21" refString="Ragkousis, M, Digenis, M, Kovacic, M, Katsanevakis, S, Gerovasileiou, V, 2021. Rarely reported cryptobenthic fish in marine caves of the eastern Mediterranean Sea. Journal of Marine Science and Engineering 9 (6): 557, DOI: https://doi.org/10.3390/jmse9060557" title="Rarely reported cryptobenthic fish in marine caves of the eastern Mediterranean Sea." url="https://doi.org/10.3390/jmse9060557" volume="9" year="2021">Ragkousis et al. 2021</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5144.1.1" author="Kovacic, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="215" pagination="1 - 103" refId="B18" refString="Kovacic, M, Renoult, JP, Pillon, R, Svensen, R, Bogorodsky, SV, Engin, S, Louisy, P, 2022. Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals. Zootaxa 5144 (1): 1 - 103, DOI: https://doi.org/10.11646/zootaxa.5144.1.1" title="Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals." url="https://doi.org/10.11646/zootaxa.5144.1.1" volume="5144" year="2022">
<normalizedToken originalValue="Kovačić">Kovacic</normalizedToken>
et al. 2022
</bibRefCitation>
), the occurrence of this species outside this type of habitat is unusual.
<taxonomicName baseAuthorityName="Lowe" baseAuthorityYear="1839" class="Actinopterygii" family="Gobiidae" genus="Thorogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thorogobius ephippiatus" order="Perciformes" pageId="0" pageNumber="215" phylum="Chordata" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">Thorogobius ephippiatus</emphasis>
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is a new record for Maltese coastal waters; it is a NE Atlantic and Mediterranean species. In the Mediterranean its distribution appears to be mostly along the northern shores of both west and east basins including the Aegean Sea, Cyprus, and Israel (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5144.1.1" author="Kovacic, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="215" pagination="1 - 103" refId="B18" refString="Kovacic, M, Renoult, JP, Pillon, R, Svensen, R, Bogorodsky, SV, Engin, S, Louisy, P, 2022. Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals. Zootaxa 5144 (1): 1 - 103, DOI: https://doi.org/10.11646/zootaxa.5144.1.1" title="Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals." url="https://doi.org/10.11646/zootaxa.5144.1.1" volume="5144" year="2022">
<normalizedToken originalValue="Kovačić">Kovacic</normalizedToken>
et al. 2022
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). According to
<bibRefCitation author="Froese, R" journalOrPublisher="International Journal of Scientific and Technological Research" pageId="0" pageNumber="215" publicationUrl="http://www.fishbase.org" refId="B12" refString="Froese, R, Pauly, D, 2022. FishBase. [Version 02/2022] http://www.fishbase.org" title="FishBase. [Version 02 / 2022]" url="http://www.fishbase.org" year="2022">Froese and Pauly (2022)</bibRefCitation>
, the depth range for
<taxonomicName lsidName="T. ephippiatus" pageId="0" pageNumber="215" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">T. ephippiatus</emphasis>
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is 6-40 m, with the fish being commonest in the 6-12 m range. The record from a depth of 117 m is therefore noteworthy, especially as the present authors are aware of only one previous deep-water record of this species, by
<bibRefCitation DOI="https://doi.org/10.1111/jai.13625" author="Stern, N" journalOrPublisher="Journal of Applied Ichthyology" pageId="0" pageNumber="215" pagination="681 - 683" refId="B23" refString="Stern, N, Weissman, A, Makovsky, Y, 2018. East and deep: Range extension and depth record for the leopard-spotted goby Thorogobius ephippiatus (Lowe, 1839) (Osteichthyes: Gobiidae). Journal of Applied Ichthyology 34 (3): 681 - 683, DOI: https://doi.org/10.1111/jai.13625" title="East and deep: Range extension and depth record for the leopard-spotted goby Thorogobius ephippiatus (Lowe, 1839) (Osteichthyes: Gobiidae)." url="https://doi.org/10.1111/jai.13625" volume="34" year="2018">Stern et al. (2018)</bibRefCitation>
, who recorded it from a depth of 156 m off the north Israeli coast.
<bibRefCitation DOI="https://doi.org/10.1111/jai.13625" author="Stern, N" journalOrPublisher="Journal of Applied Ichthyology" pageId="0" pageNumber="215" pagination="681 - 683" refId="B23" refString="Stern, N, Weissman, A, Makovsky, Y, 2018. East and deep: Range extension and depth record for the leopard-spotted goby Thorogobius ephippiatus (Lowe, 1839) (Osteichthyes: Gobiidae). Journal of Applied Ichthyology 34 (3): 681 - 683, DOI: https://doi.org/10.1111/jai.13625" title="East and deep: Range extension and depth record for the leopard-spotted goby Thorogobius ephippiatus (Lowe, 1839) (Osteichthyes: Gobiidae)." url="https://doi.org/10.1111/jai.13625" volume="34" year="2018">Stern et al. (2018)</bibRefCitation>
attribute the deep-water occurrence of
<taxonomicName lsidName="T. ephippiatus" pageId="0" pageNumber="215" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">T. ephippiatus</emphasis>
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to its requirement for dark and cold conditions, which in the eastern Levantine Sea only occur in deep water. However, the species is present in shallow water in Cyprus (
<bibRefCitation DOI="https://doi.org/10.12681/mms.13771" author="Gerovasileiou, V" journalOrPublisher="Mediterranean Marine Science" pageId="0" pageNumber="215" pagination="355 - 384" refId="B13" refString="Gerovasileiou, V, Akel, EHK, Akyol, O, Alongi, G, Azevedo, F, Babali, N, Bakiu, R, Bariche, M, Bennoui, A, Castriota, L, Chintiroglou, CC, Crocetta, F, Deidun, A, Galinou-Mitsoudi, S, Giovos, I, Gokoglu, M, Golemaj, A, Hadjioannou, L, Hartingerova, J, Insacco, G, Katsanevakis, S, Kleitou, P, Korun, J, Lipej, L, Malegue, M, Michailidis, N, 2017. New Mediterranean biodiversity records (July 2017). Mediterranean Marine Science 18 (2): 355 - 384, DOI: https://doi.org/10.12681/mms.13771" title="New Mediterranean biodiversity records (July 2017)." url="https://doi.org/10.12681/mms.13771" volume="18" year="2017">Gerovasileiou et al. 2017</bibRefCitation>
), which raises doubts about this hypothesis. On the other hand, the present deep-water record from Malta confirms that
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has a much wider bathymetric range than previously thought. It may be significant that both the Maltese and Israeli deep-water records were from open rocky bottoms and not from caves, suggesting that the association with caves in shallow water is due to the
<normalizedToken originalValue="fishs">fish's</normalizedToken>
requirement for a dark habitat, while in deep water the species can occur in the open. In fact,
<bibRefCitation author="Kovacic, M" journalOrPublisher="Natura Croatica" pageId="0" pageNumber="215" pagination="423 - 435" refId="B14" refString="Kovacic, M, 1997. Cryptobenthic gobies (Pisces, Perciformes, Gobiidae) and clingfishes (Pisces, Gobiesociformes, Gobiesocidae) in the Kvarner area, Adriatic Sea. Natura Croatica 6 (4): 423 - 435" title="Cryptobenthic gobies (Pisces, Perciformes, Gobiidae) and clingfishes (Pisces, Gobiesociformes, Gobiesocidae) in the Kvarner area, Adriatic Sea." volume="6" year="1997">
<normalizedToken originalValue="Kovačić">Kovacic</normalizedToken>
(1997)
</bibRefCitation>
had already noted that this species occurs deep within caves in very shallow waters, but closer to the mouth at the deeper end of the depth range studied by this author (32 m). It therefore also occurs outside of caves in deeper water.
</paragraph>
<caption ID-Table-UUID="9290C6EC705EBC6F33E4295155EE47FA" httpUri="http://table.plazi.org/id/9290C6EC705EBC6F33E4295155EE47FA" pageId="0" pageNumber="215" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Table 1.</emphasis>
Individuals of
<taxonomicName baseAuthorityName="Lowe" baseAuthorityYear="1839" class="Actinopterygii" family="Gobiidae" genus="Thorogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thorogobius ephippiatus" order="Perciformes" pageId="0" pageNumber="215" phylum="Chordata" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">Thorogobius ephippiatus</emphasis>
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recorded from the LIFE
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for N2K marine benthic surveys.
</paragraph>
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<paragraph pageId="0" pageNumber="215">
<table pageId="0" pageNumber="215">
<tr pageId="0" pageNumber="215">
<th colspan="1" pageId="0" pageNumber="215" rowspan="1">Survey method</th>
<th colspan="1" pageId="0" pageNumber="215" rowspan="1">Date</th>
<th colspan="1" pageId="0" pageNumber="215" rowspan="1">Geographical coordinates</th>
<th colspan="1" pageId="0" pageNumber="215" rowspan="1">Water depth [m]</th>
<th colspan="1" pageId="0" pageNumber="215" rowspan="1">Habitat type</th>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">8 June 2015</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="35" direction="north" minutes="52.021" orientation="latitude" precision="1" value="35.867016">35°52.021N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="21.174" orientation="longitude" precision="1" value="14.3529">014°21.174E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">32</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Rock covered with a thin layer of detritus within cave</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">10 July 2015</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="36" direction="north" minutes="00.813" orientation="latitude" precision="1" value="36.01355">36°00.813N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="15.883" orientation="longitude" precision="1" value="14.264717">014°15.883E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Soft sediment covered with a thin layer of detritus within cave</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">12 June 2016</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="36" direction="north" minutes="04.879" orientation="latitude" precision="1" value="36.081318">36°04.879N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="14.110" orientation="longitude" precision="1" value="14.235167">014°14.110E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">In rock fissure within cave</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">12 June 2016</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="36" direction="north" minutes="04.879" orientation="latitude" precision="1" value="36.081318">36°04.879N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="14.110" orientation="longitude" precision="1" value="14.235167">014°14.110E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Soft sediment bottom within cave</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">27 June 2016</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="36" direction="north" minutes="04.879" orientation="latitude" precision="1" value="36.081318">36°04.879N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="14.110" orientation="longitude" precision="1" value="14.235167">014°14.110E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Soft sediment bottom within cave</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">ROV</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">7 July 2016</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="36" direction="north" minutes="00.776" orientation="latitude" precision="1" value="36.01293">36°00.776N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="50.517" orientation="longitude" precision="1" value="14.84195">014°50.517E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">117</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Rock covered with a thin layer of detritus</td>
</tr>
<tr pageId="0" pageNumber="215">
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">SCUBA diving</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">25 July 2016</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">
<geoCoordinate degrees="35" direction="north" minutes="50.969" orientation="latitude" precision="1" value="35.849483">35°50.969N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="23.050" orientation="longitude" precision="1" value="14.384167">014°23.050E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="215" rowspan="1">Soft sediment bottom within cave</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/aiep.52.89340.figure1" httpUri="https://binary.pensoft.net/fig/747670" pageId="0" pageNumber="215" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 1.</emphasis>
Photograph of
<taxonomicName baseAuthorityName="Lowe" baseAuthorityYear="1839" class="Actinopterygii" family="Gobiidae" genus="Thorogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thorogobius ephippiatus" order="Perciformes" pageId="0" pageNumber="215" phylum="Chordata" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">Thorogobius ephippiatus</emphasis>
</taxonomicName>
, taken on 27 June 2016 at a depth of 24 m within a cave (
<geoCoordinate degrees="36" direction="north" minutes="04.879" orientation="latitude" precision="1" value="36.081318">36°04.879N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="14.110" orientation="longitude" precision="1" value="14.235167">014°14.110E</geoCoordinate>
). Photograph: OCEANA/Carlos Minguell© LIFE
<normalizedToken originalValue="BaĦAR">BaĦAR</normalizedToken>
for N2K.
</paragraph>
</caption>
<caption doi="10.3897/aiep.52.89340.figure2" httpUri="https://binary.pensoft.net/fig/747671" pageId="0" pageNumber="215" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="215">
<emphasis bold="true" pageId="0" pageNumber="215">Figure 2.</emphasis>
Image grab from ROV footage collected on 7 July 2016 at a depth of 117 m (
<geoCoordinate degrees="36" direction="north" minutes="00.776" orientation="latitude" precision="1" value="36.01293">36°00.776N</geoCoordinate>
,
<geoCoordinate degrees="014" direction="east" minutes="50.517" orientation="longitude" precision="1" value="14.84195">014°50.517E</geoCoordinate>
) showing an individual of
<taxonomicName baseAuthorityName="Lowe" baseAuthorityYear="1839" class="Actinopterygii" family="Gobiidae" genus="Thorogobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thorogobius ephippiatus" order="Perciformes" pageId="0" pageNumber="215" phylum="Chordata" rank="species" species="ephippiatus">
<emphasis italics="true" pageId="0" pageNumber="215">Thorogobius ephippiatus</emphasis>
</taxonomicName>
. Image: OCEANA© LIFE
<normalizedToken originalValue="BaĦAR">BaĦAR</normalizedToken>
for N2K.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>