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<document id="C1172214D16E9C5E21BABA53711607BB" ID-DOI="10.3897/jhr.97.127500" ID-ZooBank="F340D011-940A-47BB-9A5B-78D9B3DE1833" ID-publisher-id="127500" URI-arpha="03041528-079C-5F0A-A2DB-9DDB50728C61" URI-zoobank="http://zoobank.org/F340D011-940A-47BB-9A5B-78D9B3DE1833" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1725892545912" checkinUser="pensoft" docAuthor="Benda, Daniel, Pohl, Hans, Beutel, Rolf &amp; Straka, Jakub" docDate="2024" docId="0E4AF8F4762D5F40ACB296709C8040EA" docLanguage="en" docName="Journal of Hymenoptera Research 97: 721-739" docOrigin="Journal of Hymenoptera Research 97" docSource="https://jhr.pensoft.net/article/127500/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Deltoxenos impressus Benda &amp; Pohl &amp; Beutel &amp; Straka 2024, sp. nov." docType="treatment" docUuid="9FE257A5-8010-4607-9285-7364594430E2" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="739" masterDocId="03041528079C5F0AA2DB9DDB50728C61" masterDocTitle="Solitary folded-winged wasps of the genus Zethus Fabricius (Vespidae, Zethinae) parasitised by two new species of Strepsiptera on different continents" masterLastPageNumber="739" masterPageNumber="721" pageNumber="721" updateTime="1725895626845" updateUser="admin">
<mods:mods id="95FE2D93121CF32E2F3319B835D971D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C8C056B418724B771CDF63DA095F9C9">
<mods:title id="B12DAA9E5784F4D2832E3255BEF67E3C">Solitary folded-winged wasps of the genus Zethus Fabricius (Vespidae, Zethinae) parasitised by two new species of Strepsiptera on different continents</mods:title>
</mods:titleInfo>
<mods:name id="F8CEB40F5D51BF3A9D5D509ED40FF741" type="personal">
<mods:role id="AFFD08E110FEDCAC0F472C1B6B3B9C22">
<mods:roleTerm id="DE3F4E70CBE8F7ED8ED9DBBF4431244B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3DF09FA58891FFF5AD36B8A945981B3B">Benda, Daniel</mods:namePart>
<mods:nameIdentifier id="535D60A4783D03CFFFC84FA93BB892AC" type="ORCID">0000-0002-5729-0411</mods:nameIdentifier>
<mods:affiliation id="1FF0A6FC750A8BFE6731BFE88910F94D">Department of Entomology, National Museum of the Czech Republic, Prague, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="2897BBEBC167F4613B5FA9D016E67FCC" type="personal">
<mods:role id="88C0B362F234E0F40A3C5A84C42C4109">
<mods:roleTerm id="D595EFBF94EA26E604CA5195493955A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A752AF841F836ADA2CA097B50739165A">Pohl, Hans</mods:namePart>
<mods:nameIdentifier id="B8DB6F7BFC154459D2109E9C826613B4" type="ORCID">0000-0002-7090-6612</mods:nameIdentifier>
<mods:affiliation id="AFF901090E4EEDBBF97E8B605645B52E">Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität, Jena, Germany</mods:affiliation>
</mods:name>
<mods:name id="CAD2B19BE54F6717CC9BDDF34B9D04CC" type="personal">
<mods:role id="E18AC57FE8847A0552926188D7DEE7A4">
<mods:roleTerm id="6531D588327C3B3548693310B60EE6F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4753545078ED7E27190D741679A0B8C0">Beutel, Rolf</mods:namePart>
<mods:affiliation id="FFD79CFD55542E97A2DBF1A1F3ADEA1D">Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität, Jena, Germany</mods:affiliation>
</mods:name>
<mods:name id="390872D27DE9AA57AE999FF103786884" type="personal">
<mods:role id="03F6C60002CE05B1BA2D4D30F3412AED">
<mods:roleTerm id="D980E76A6FBA9B90FF875671D2BAE68A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16B4126CC6846A006E927B9E58B88E5D">Straka, Jakub</mods:namePart>
<mods:nameIdentifier id="156858BFA34882C8A7162821B5EB5803" type="ORCID">0000-0002-8987-1245</mods:nameIdentifier>
<mods:affiliation id="7C258A127F1C4736DDEDA7706AB42364">Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E2CFA603F2723D53FEE4C5D1A2616E39">text</mods:typeOfResource>
<mods:relatedItem id="73AAC46E0CBD0C1C4B374EECCE6D6F86" type="host">
<mods:titleInfo id="C298255752DF937D2A82B5083C93927B">
<mods:title id="AEB946E9BA802422E284ED112AC34B61">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="C4D175360453925B702D34BD6C442C82">
<mods:date id="430644BE2BE63EADCAAE09610BA9342B">2024</mods:date>
<mods:detail id="90F2CDFD3F8835CC6FA90F6C75598847" type="pubDate">
<mods:number id="B90E4D62D757391035A10B7A9EC13720">2024-09-09</mods:number>
</mods:detail>
<mods:detail id="32F600878E8FD7AD0844577F4E88204F" type="volume">
<mods:number id="180AB68C569EC03B604D6F8A224ADBEA">97</mods:number>
</mods:detail>
<mods:extent id="AE658ED937AA1949FEF54B052EC0516B" unit="page">
<mods:start id="1043CEC91670FF23B34038901F4BB2F8">721</mods:start>
<mods:end id="98348BE363BD3BF1B5A7DBC38A57BCEA">739</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A4CB25AF2DEFEED874DBBD7CC8036CCE">journal article</mods:classification>
<mods:identifier id="38962F57E92B1DD284A69895F8FEDE3F" type="DOI">10.3897/jhr.97.127500</mods:identifier>
<mods:identifier id="F59B7693673C1D52BE332E13139C9F45" type="ZooBank">F340D011-940A-47BB-9A5B-78D9B3DE1833</mods:identifier>
</mods:mods>
<treatment id="0E4AF8F4762D5F40ACB296709C8040EA" ID-arpha="0E4AF8F4-762D-5F40-ACB2-96709C8040EA" ID-zoobank="https://zoobank.org/9FE257A5-8010-4607-9285-7364594430E2" LSID="urn:lsid:zoobank.org:act:9FE257A5-8010-4607-9285-7364594430E2" LSID-ZBK="urn:lsid:zoobank.org:act:9FE257A5-8010-4607-9285-7364594430E2" httpUri="http://treatment.plazi.org/id/0E4AF8F4762D5F40ACB296709C8040EA">
<subSubSection id="5B0F0D844580C587AF786BAA841594D9" type="nomenclature">
<paragraph id="DF6F2986D504F3FC779E5868A973B85C">
<taxonomicName id="5C2F1B6D5FD3F9C0C90637BB79BF4181" authority="Benda &amp; Straka" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Vespidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="73CD723769F506DE3F094B508C19A6F4" italics="true">Deltoxenos impressus</emphasis>
Benda &amp; Straka
</taxonomicName>
<taxonomicNameLabel id="F15F67D665DFA11CFADFBE8E77165787" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUTAE" type="type material">
<paragraph id="AE753C4B003FEFC98034DDA79CFE7D9A">
<heading id="2E6DA0E24931C996A4C3CE037BF0BC2C" reason="title">Type material.</heading>
</paragraph>
<paragraph id="D0A1D08814E3D8031D518EA20D7E16DD">
<materialsCitation id="CBBBCB33403CF830D1DCA9FF0F3E96FE" collectedFrom="Zethus favillaceus Walker, 1871" collectingDate="2007-04-18" collectionCode="OLML" collectorName="M. Halada" country="Kenya" county="Mwingi" location="Kangonde vadi" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="51CFC09D00854C230CA0B1DBCF0FDB27" bold="true">
<emphasis id="3F62B8DCA234DC625A773D87A108CC1A" bold="true" italics="true">
<typeStatus id="CAA9EF5D9727AC346A2CDCF6127BB3DF" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="BC65BD03E00CE70DB1D836C777F6C73B" name="Kenya">Kenya</collectingCountry>
•
<specimenCount id="329CDC9935A5190903782CC116145D58" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="116C72CE37A72BDC3C414B9AA2355B95">Mwingi</collectingCounty>
,
<location id="4AB50C93565349B8438C8023E5C1D5D8" LSID="urn:lsid:plazi:treatment:0E4AF8F4762D5F40ACB296709C8040EA:4AB50C93565349B8438C8023E5C1D5D8" country="Kenya" county="Mwingi" name="Kangonde vadi">Kangonde vadi</location>
,
<date id="95D1217B2F909B026E02A72E1E584C92" value="2007-04-18">
<collectingDate id="9FF25B0E2CFAD985A2AB2DEE668A4997" value="2007-04-18">18 Apr. 2007</collectingDate>
</date>
;
<collectorName id="EA894F3877C93A5DF0CC0763DC4C22BC">M. Halada</collectorName>
leg.;
<named-content id="C518F3B97FD9F6CC8D1655AC271BD365" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">
<collectionCode id="879B5B91C6DDAB38102CD134722E0F69" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" type="Museum">OLML</collectionCode>
</named-content>
; host:
<taxonomicName id="0240A11ADF3BA64586A9B39017789C55" authority="Walker, 1871" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<collectedFrom id="A436DDFE55FA38FCD3DDBE8EB2E31290">
<emphasis id="A4B773C1A18402F51518538B032EF98A" italics="true">Zethus favillaceus</emphasis>
Walker, 1871
</collectedFrom>
</taxonomicName>
.
</materialsCitation>
</paragraph>
<paragraph id="748FEB2B4B3D896BAAA3E65BDE49CFF1">
<materialsCitation id="2AA231A61D526C37F3BC04E446892BD7" collectedFrom="Zethus favillaceus Walker, 1871" collectingDate="2007-04-18" collectionCode="OLML" collectorName="M. Halada" country="Kenya" county="Mwingi" location="Kenya" specimenCount="5" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
<emphasis id="150BDB8696D5D8248A7F263B0D9DBC1E" bold="true">
<emphasis id="6AAA6E1B86CA3F7945C3C0BE04EB4BD8" bold="true" italics="true">
<typeStatus id="4170503FDB32561023B27DC31DC23F93" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="AD256691B57AEC66F1979BC5F5CAE61C" name="Kenya">Kenya</collectingCountry>
•
<specimenCount id="A23F7AC5E2FE0356B1BD9585E9EE6C8E" count="2" type="female">2 ♀</specimenCount>
;
<specimenCount id="8546C565FABC1389E1900B3CD720D637" count="1" type="female">1 ♀</specimenCount>
+ 2 empty male puparia;
<specimenCount id="283B657AECDB01F3329CCADDDFB30FBA" count="1" type="female">1 ♀</specimenCount>
+ 1 male puparium;
<specimenCount id="213426CDB6195F60D79AFAB44E655439" count="1" type="female">1 ♀</specimenCount>
+ 1 male puparium + 2 empty male puparia; same locality and host as for holotype,
<date id="7865CD23A696378F3FC532EDCC808CC7" value="2007-04-18">
<collectingDate id="C34321A052C45001E536A3D3FFBEE53C" value="2007-04-18">18 Apr. 2007</collectingDate>
</date>
;
<collectorName id="99373F0AF05F63BFEAB4E97C8C5D00D5">M. Halada</collectorName>
leg.;
<named-content id="E6CEC9B980138EF928C421E6B6A1B156" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">
<collectionCode id="9A324824F1D7EED3CB922FC9D4C50553" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" type="Museum">OLML</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZUAE" type="diagnosis">
<paragraph id="EB0DA927C4702D320CBF3DE6A409FF6E">
<heading id="4FA47106344846A8A535FD75B12AD44D" reason="title">Diagnosis of female cephalothorax.</heading>
</paragraph>
<paragraph id="5BAF4C0037CC2CEC33ACA58F13511E26">
This species is easily distinguished from other representatives of the genus
<taxonomicName id="157D6198B4952A8B3AB0AC7B2E60E683" authorityName="Benda, Pohl, Nakase, Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="genus">
<emphasis id="7339A036F97A1E5070034ACD6E430731" italics="true">Deltoxenos</emphasis>
</taxonomicName>
by mandibles distinctly anteromedially directed (angle of 65 °) and by conspicuous impressions on the surface of the lateral extensions at the site of the reduced compound eyes. These impressions are best visible on
<abbrev id="ABBRID0EGVAE" xlink_title="Scanning electron microscopy">SEM</abbrev>
images (im; Fig.
<figureCitation id="A2F396E079C92106B562F2428DB88562" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 A</figureCitation>
) but also recognizable with a light microscope. The surface of the lateral extensions is slightly wrinkled or smooth in other species of
<taxonomicName id="D0DBFE51B6FCE434B19CA0830159DFF7" authorityName="Benda, Pohl, Nakase, Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="genus">
<emphasis id="52CA5D9FD3FF382DD4387A197660B82F" italics="true">Deltoxenos</emphasis>
</taxonomicName>
. The anterior head margin is rounded, and the clypeal lobe is merged with the head capsule, in contrast to
<taxonomicName id="8D565FEE4FF7A882B83B4D5259749BD4" authority="(Kinzelbach, 1971)" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="63242A4ADCC3EDA46038DE152367DA7C" italics="true">Deltoxenos rueppelli</emphasis>
(Kinzelbach, 1971)
</taxonomicName>
, where the clypeal lobe protrudes distinctly from the head capsule but is blunted on top. On the ventral side the clypeal area is very smooth and lacks sensilla (cl; Fig.
<figureCitation id="74C52CDD60C560141243433F6244BACF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 D</figureCitation>
), whereas sensilla are present ventrally in
<taxonomicName id="C2D13C83B251E3D77A3E3C5FF26B7A02" authority="(Pasteels, 1950)" baseAuthorityName="Pasteels" baseAuthorityYear="1950" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="684F475138A34E101FDD5A87561E1482" italics="true">Deltoxenos bidentatus</emphasis>
(Pasteels, 1950)
</taxonomicName>
and
<taxonomicName id="87ED3DC8709BE981C31A3CB6FA553883" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="780AA453A168CE9483D174500A8BF481" italics="true">D. rueppelli</emphasis>
</taxonomicName>
. In contrast to
<taxonomicName id="B1043E69F632B17FFBC4C31AAD385118" authority="(Luna de Carvalho, 1956)" baseAuthorityName="Luna de Carvalho" baseAuthorityYear="1956" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="78C9415ADF19CA50F86A339A936B1669" italics="true">Deltoxenos bequaerti</emphasis>
(Luna de Carvalho, 1956)
</taxonomicName>
, the cuticle of the thoracic segments on the ventral side is reticulate, with scattered inconspicuous or more distinct and pigmented papillae, mainly visible on the metasternum. In contrast, the thoracic segments on the ventral side are evenly scattered with conspicuous papillae in
<taxonomicName id="40440523C3540FA1CB94034924A2F120" baseAuthorityName="Luna de Carvalho" baseAuthorityYear="1956" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="439F46EFF536D0AE72712FF6EE778606" italics="true">D. bequaerti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F156DCA3B55D075BD6FE9ADDE8F040A7">
Other characters that distinguish
<taxonomicName id="ABA6B54B03D85B7CD188DE3A75A0B9A5" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="37A312610BD767A0DC24B6BD5ED36F58" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5742D03913FF115F2C185CC48B2838ED" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="094DB1A115C401D9CE48E73516E22EFC" baseAuthorityName="Pasteels" baseAuthorityYear="1950" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="B240C1598F421309C94C8C90BA744696" italics="true">D. bidentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="40E4CA842DA57BBECB7C2C9A574690CE" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="9406A4D818EA169A392F31AC1D80D0D0" italics="true">D. rueppelli</emphasis>
</taxonomicName>
: ventral field wider than long, elliptic, nearly circular versus more flattened and not nearly circular; dorsal field arcuate, slightly raised versus dorsal field distinctly arcuate and distinctly raised; mandible anteromedially directed at angle of 65 ° versus a maximum of 45 ° in
<taxonomicName id="D7B4229D6CFCB48C500C48E512D5EBE4" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="37BF02A190677F975C94B86FF59BF3E5" italics="true">D. rueppelli</emphasis>
</taxonomicName>
; mandibular bulge distinctly raised, elongated, slightly curved laterally versus mandibular bulge slightly raised and anteriorly directed; cuticle of mandible completely smooth versus almost completely wrinkled.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5YAE" type="description">
<paragraph id="04B87A8DA7036B64EA382801D3776589">
<heading id="1024C6D28E8E14E23DB0509D5BC25454" reason="title">Description of female cephalothorax.</heading>
</paragraph>
<paragraph id="EFDA6D3464CB77C2EB8C18F89F3656DD">
<emphasis id="8035BE7191A2A80F920399A67160D631" bold="true">
<emphasis id="DEFE1FEE88B5960028F058928DD4C056" bold="true" italics="true">Shape and colouration</emphasis>
.
</emphasis>
Size of
<typeStatus id="D1FB74D5A1BB85A6C4DD9BC1E641CABB" type="holotype">holotype</typeStatus>
cephalothorax: length
<quantity id="7D3F29DFCA62ED1251D1A13FF2A177A9" metricMagnitude="-3" metricUnit="m" metricValue="1.06" unit="mm" value="1.06">1.06 mm</quantity>
, width
<quantity id="860CEA0AE2966E7673675FA74F5A3381" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.9 mm</quantity>
. Cephalothorax variable in size but always distinctly longer than wide, length
<quantity id="5EE3F900B4C85D547A361BB2ACBD4823" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="10.6" metricValueMin="9.0" unit="mm" value="0.98" valueMax="1.06" valueMin="0.9">0.9–1.06 mm</quantity>
, width
<quantity id="4A02DCC64052BAA78B5ABC28673F1C82" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="9.0" metricValueMin="7.4" unit="mm" value="0.82" valueMax="0.9" valueMin="0.74">0.74–0.9 mm</quantity>
. Pro-mesothoracic and meso-metathoracic segmental border only slightly constricted laterally (Fig.
<figureCitation id="060F6AF8355B56DB281F313600C10A5A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, female cephalothorax A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with female cephalothorax C – D holotype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle." figureDoi="10.3897/jhr.97.127500.figure4" httpUri="https://binary.pensoft.net/fig/1130330">4 C, D</figureCitation>
). Abdominal segment I not protruding laterally, corner below spiracles rounded. Anterior head margin rounded, not distinctly protruding from remaining head capsule. Thorax elongated, very slightly widening posteriorly. Cephalothorax with conspicuously contrasting light and dark colour pattern.
</paragraph>
<caption id="16CDD4657916916AB2D6319C552D355C" ID-DOI="10.3897/jhr.97.127500.figure4" ID-arpha="A019CF79-4875-5696-A10F-C5B4DFA67D25" httpUri="https://binary.pensoft.net/fig/1130330" startId="F4">
<paragraph id="659E7089AC084BFCD164AD3932A1E4A5">
<label id="C556A237E22460489E285C081B885CBF">Figure 4.</label>
</paragraph>
<paragraph id="A67158F2580F145D29F91B9B19935648">
<taxonomicName id="E497A32E7686915DEF11DEFAF700ED3A" authority="Benda &amp; Straka" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="857A44B59385CB98B21D82202F1D0413" italics="true">Deltoxenos impressus</emphasis>
Benda &amp; Straka
</taxonomicName>
,
<taxonomicNameLabel id="529BB626DBA9C58933F494025A1AD618" rank="species">sp. nov.</taxonomicNameLabel>
, host, female cephalothorax
<emphasis id="C46743D1914E6B66AA6836307BC07C7B" bold="true">A</emphasis>
<taxonomicName id="4EE9DA9494BAE87C5C9F18FB57957033" authority="Walker, 1871" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="D05D9215104C673F1DD76F2B2477B773" italics="true">Zethus favillaceus</emphasis>
Walker, 1871
</taxonomicName>
stylopised by
<taxonomicName id="2DF9542698927719384030DC12C16B1E" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="5842EE21274B615765596F80083E9A94" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F59F5342BEC38EF7FC82BD21173AAB51" rank="species">sp. nov.</taxonomicNameLabel>
, lateral view
<emphasis id="245DCC38CE05DCD1F3B7CF61A34894E4" bold="true">B</emphasis>
detail of host abdomen of
<taxonomicName id="087AE1E15B17B36D6651F5AD721D4071" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="F92D7B8BE0C192D6076112FCBAD9911C" italics="true">Z. favillaceus</emphasis>
</taxonomicName>
, with female cephalothorax
<emphasis id="4FAF7F71D5102464C101D4CD4EEE9AA6" bold="true">C – D</emphasis>
holotype of
<taxonomicName id="FAAC85C6857E208D3BDD60C4DE53034E" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="660B3DBD657DB5A675C7E473E0E8451D" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDC9D89CEE95E51B7F9197AE4E41EEE3" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (
<named-content id="9546245975E71B1458889A0BD0C88C6A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">OLML</named-content>
) from
<taxonomicName id="CB8636083C6E07CB9212D300B3DE88C9" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="1582BC7879FF11478C6DD99CDA9ADA48" italics="true">Z. favillaceus</emphasis>
</taxonomicName>
<emphasis id="9E3B4A5F2A91B27B81AD8D49783B8BDA" bold="true">C</emphasis>
ventral side of cephalothorax
<emphasis id="9CF3AB094B091E6E68127CD369ABC8D0" bold="true">D</emphasis>
dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle.
</paragraph>
</caption>
<paragraph id="BFD64B3CAB240D7C2110A9FC2DF43B02">
<emphasis id="CC013DD9048D8841299E42091DAD2986" bold="true">
<emphasis id="2066F197A1A75CB7AC3C8CFA4FA4F0F9" bold="true" italics="true">Head capsule</emphasis>
.
</emphasis>
Ca 2 / 5 as long as entire cephalothorax including lateral cephalic extension. Colouration forming specific pattern with dark brown anterior part and pale lateral extensions. Surface of lateral extensions at site of reduced compound eyes smooth, with conspicuous impressions visible on
<abbrev id="ABBRID0ER3AE" xlink_title="Scanning electron microscopy">SEM</abbrev>
images (im; Fig.
<figureCitation id="8951AD614944760831C2BF9948DC0C90" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 A</figureCitation>
). Clypeal area well delimited from labral area, arcuate, clypeal lobe merged with head capsule. Surface completely smooth with slightly more than 40 distinctly exposed sensilla on dorsal side (cls; Fig.
<figureCitation id="20F2B91BD235DAFE72E30C6040A9F62C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 F</figureCitation>
). Ventral side of clypeal area smooth and lacking sensilla (cl; Fig.
<figureCitation id="FEF20A1E9F89437C98E5FE340596A2FD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 D</figureCitation>
).
<collectingRegion id="33B45D531248A2FBFE2015A7EFF3E1FE" country="Saudi Arabia" name="Al Hudud ash Shamaliyah">Border</collectingRegion>
between clypeal and frontal region indistinct but still present. Frontal region smooth (fr; Fig.
<figureCitation id="1E985189E0B46B502ED6FEFB621C54DF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 B</figureCitation>
). Segmental border between head and prothorax indicated by distinct mesal furrow on dorsal side (sbhp; Fig.
<figureCitation id="6F4B203A3EF2A9BE67FD8A0DCF5F3FC7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 B</figureCitation>
) and by dorsal transverse stripe of frontal and occipital papillae (p; Fig.
<figureCitation id="304C8837BED3B828F5985BE4FF44B743" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 B</figureCitation>
). Head and prothorax distinctly separated by birth opening ventromedially (bo; Fig.
<figureCitation id="56A60D56CC24D9F0DE565415106F8C5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 A</figureCitation>
) and laterally by suture (sbhp; Fig.
<figureCitation id="4A2D630EA941A7FC8FE43137BE34F38E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 A</figureCitation>
).
</paragraph>
<caption id="E80877722DD59ECA7CE2F068B5BE1992" ID-DOI="10.3897/jhr.97.127500.figure5" ID-arpha="C0B11CD1-9720-580E-A223-C5FF15A436CB" httpUri="https://binary.pensoft.net/fig/1130331" startId="F5">
<paragraph id="154BA2EA1DC4CE60FDC478DF66C85A38">
<label id="9FBC66C65360A050758F72CFE46EE675">Figure 5.</label>
</paragraph>
<paragraph id="6BA380DD895E55DF80B364C790E95E3B">
Holotype of
<taxonomicName id="6F17832776D56E4E2FBDD67CF2DEB8A6" authority="Benda &amp; Straka" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="2754B912E3176EFF2EA47A9F23D527D1" italics="true">Deltoxenos impressus</emphasis>
Benda &amp; Straka
</taxonomicName>
,
<taxonomicNameLabel id="8BA8158ECC13505A41F512994197C0B0" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (
<named-content id="312ED42C6E55F325395F746B92F7BE7B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">OLML</named-content>
),
<abbrev id="ABBRID0EM5AE" xlink_title="Scanning electron microscopy">SEM</abbrev>
micrographs of cephalothorax
<emphasis id="E13313F3B80636ECEF3A89A8DAC3BB01" bold="true">A</emphasis>
ventral side
<emphasis id="D92CA6C67CDC03A7A16A3ACD1F75AB89" bold="true">B</emphasis>
dorsal side
<emphasis id="ABC6C22568466D4725DD05A07DDB8FF1" bold="true">C</emphasis>
right vestigial antenna, dorsal side
<emphasis id="34F3AE506EC9F67809BB2031BF79721A" bold="true">D</emphasis>
left vestigial antenna, dorsal side
<emphasis id="0D292949A9C352F831F0ED5A3C7F8E23" bold="true">E</emphasis>
left lateral border of abdominal segment I below spiracle, dorsal side
<emphasis id="610E0663C7F4B6ED4ED17055018434B3" bold="true">F</emphasis>
detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax.
</paragraph>
</caption>
<caption id="A0DAD5569E92508DB5280829F2A20DBA" ID-DOI="10.3897/jhr.97.127500.figure6" ID-arpha="6513ED54-6FFC-52C0-A584-C839FA1F3A9E" httpUri="https://binary.pensoft.net/fig/1130332" startId="F6">
<paragraph id="C01C9843B1C7BCF0FCB390D07DAAD9C1">
<label id="0D92A05E29AA7F1A06FB3DF18E512644">Figure 6.</label>
</paragraph>
<paragraph id="6B8ECC00CDE4EA75774EF8E5124308F4">
Holotype of
<taxonomicName id="C09EED8FBF13DF8E50358D5951D46F9C" authority="Benda &amp; Straka" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="4F05A9F9491C12631E8D31AEEF8CDAD0" italics="true">Deltoxenos impressus</emphasis>
Benda &amp; Straka
</taxonomicName>
,
<taxonomicNameLabel id="CA08EBE047876B195A40BAFB69CD7817" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (
<named-content id="3CF3203887C371673B6FE6BDFBFE6D66" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">OLML</named-content>
),
<abbrev id="ABBRID0EY6AE" xlink_title="Scanning electron microscopy">SEM</abbrev>
micrographs of cephalothorax
<emphasis id="8D374EE5F222E49C5DF462D696684F3B" bold="true">A</emphasis>
anterior part of cephalothorax, ventral side
<emphasis id="010F927C2EEB0BE90BCC66C0C8878649" bold="true">B</emphasis>
anterior part of cephalothorax, dorsal side
<emphasis id="8AB8688916E5217061349F63D58EB776" bold="true">C</emphasis>
mouthparts, ventral side
<emphasis id="884DAAD42FCE824038B97B5B78FB5B92" bold="true">D</emphasis>
detail of anterior border of cephalothorax, ventral side
<emphasis id="423509C78A587FD75364C1CD1A383639" bold="true">E</emphasis>
right mandible and maxilla, ventral side
<emphasis id="13C358CCB4DA63364EE7FCA5D7FAD365" bold="true">F</emphasis>
left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area.
</paragraph>
</caption>
<paragraph id="36A6EA7A52B8832567153803E490E8F1">
<emphasis id="2F6A69FCCB8F1BB4DFBD9AE4E0E4FE82" bold="true">
<emphasis id="104625D6913ECD47275DCBA98CB507DD" bold="true" italics="true">Supra</emphasis>
-
<emphasis id="7347E4B1F9C37E9AB9CCA98BCECFDDEB" bold="true" italics="true">antennal sensillary field</emphasis>
.
</emphasis>
Completely smooth, with dispersed sensilla (Fig.
<figureCitation id="3622C28F8276DF1A885F9B34FDC98A37" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Eupathocera zethi Benda &amp; Straka, sp. nov., ♀ (NMPC), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: asI – abdominal segment I, fr – frontal region, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pra – plate of vestigial antenna, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, ssf – supra-antennal sensillary field." figureDoi="10.3897/jhr.97.127500.figure2" httpUri="https://binary.pensoft.net/fig/1130328">2 C, D</figureCitation>
). Distinctly delimited by furrow on medial side (fssf; Fig.
<figureCitation id="4EFA1D9493E65F828070DA242101828C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 B</figureCitation>
), surface of supra-antennal sensillary field and frontal region with same sculpture.
</paragraph>
<paragraph id="9BF565183ADE553824D6074A57182760">
<emphasis id="E4F0C4064BFC31D87BFB847F0BC112C3" bold="true">
<emphasis id="28E189A0C073613D83708479756E75D7" bold="true" italics="true">Antenna</emphasis>
.
</emphasis>
Preserved as poorly defined area, with several distinct rounded plates and an inconspicuous cavity (Fig.
<figureCitation id="ADF53FBA03BD1C538BD4D9C05ECE458A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 C, D</figureCitation>
). Antennal torulus reduced. Periantennal area expanded, smooth (paa; Fig.
<figureCitation id="5288546802C89D5F89E628FCCBA33391" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 C, D</figureCitation>
).
<collectingRegion id="E4533009D4D45ED07682BA1973FCEDEB" country="Saudi Arabia" name="Al Hudud ash Shamaliyah">Border</collectingRegion>
between antennal area and supra-antennal sensillary field indistinct.
</paragraph>
<paragraph id="2E325F01BF66488FDF97B6CB5C1229DD">
<emphasis id="76944ADAF135049CB69E7B296F454E08" bold="true">
<emphasis id="E92B1BD54A6977AE09CB9B0121FCBA79" bold="true" italics="true">Labrum</emphasis>
.
</emphasis>
Ventral field wider than long, elliptic, nearly circular. Dorsal field arcuate, slightly raised, laterally narrower than medially, 6 × wider than long in midline (vlf, dlf; Fig.
<figureCitation id="64CF3C9349EEAD3D1B18347FA0746AD9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 C, D</figureCitation>
). Dorsal field with 16 sensilla inserted in cavities.
</paragraph>
<paragraph id="B7E565BC7AE4F84740DC63FD42B19025">
<emphasis id="88E7CBB7207A682C5DF3A1D5F5E5D038" bold="true">
<emphasis id="A30BBFB9102CAC1355A7920628C59306" bold="true" italics="true">Mandible</emphasis>
.
</emphasis>
Mandibles anteromedially directed at angle of 65 °, enclosed in mandibular capsule. Mandibular bulge distinctly raised, elongated, slightly curved laterally, with several sensilla (mdb; Fig.
<figureCitation id="E9AD7A7653D464C4EB28C0F650200DEF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 E, F</figureCitation>
). Cuticle of mandible completely smooth. Mandibular tooth slightly curved, pointed apically, almost completely lacking spines (mdt; Fig.
<figureCitation id="64567919D9773C6C4CF1B24501EF2286" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 E, F</figureCitation>
).
</paragraph>
<paragraph id="AA313A9B5238FD60029AC6430E013C07">
<emphasis id="C2EEA716721EC8F9D3C65A5F45E6A560" bold="true">
<emphasis id="CE0F57FB9C1625E5151A79B80B41890C" bold="true" italics="true">Maxilla</emphasis>
.
</emphasis>
Only slightly raised, almost fused with labial area (mx; Fig.
<figureCitation id="3ECE8B4064122E3F2F62D00FEB7C6C7A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 C</figureCitation>
). Cuticle smooth, with longitudinal furrow. Apical maxillary region not projecting beyond mandibular apex. Basal part firmly connected with labium and not overlapping with mandible (mxb; Fig.
<figureCitation id="48971D666447E4D3064FC37AD7772285" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 C</figureCitation>
). Vestige of palp indistinct. Maxillary base distinctly produced anterolaterally as submaxillary groove. Space between prothoracic extension and head extended (sbhp, mxb; Fig.
<figureCitation id="3A5A34B8128F42F97FF87A2F87CE3ADB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 A</figureCitation>
).
</paragraph>
<paragraph id="5A3D9159017C7019091EC95AF08B73DF">
<emphasis id="3A0798E92C8EDA853B6A8D55EA9D91BC" bold="true">
<emphasis id="43512E315305483B2B0BE0B0492F113A" bold="true" italics="true">Labium</emphasis>
.
</emphasis>
Labial area quite indistinct between maxillae, delimited anteriorly by mouth opening and posteriorly by birth opening (lb; Fig.
<figureCitation id="1F0B583E958C819EFF70ED43DE3BF643" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 C</figureCitation>
). Flat, approximately as long as wide. Cuticular surface smooth.
</paragraph>
<paragraph id="133E26FE64A9F6DF939E8018909DB1C7">
<emphasis id="435495BDDCF9753EF79D5B34C6AD6D89" bold="true">
<emphasis id="507C155A907D372D6AEC2936B0CA7A62" bold="true" italics="true">Mouth opening</emphasis>
.
</emphasis>
Widely arcuate, almost semicircular, sclerotised along margin (os; Fig.
<figureCitation id="1C1BCA9BEE1251E8ACEFF4DE69CE0B13" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A anterior part of cephalothorax, ventral side B anterior part of cephalothorax, dorsal side C mouthparts, ventral side D detail of anterior border of cephalothorax, ventral side E right mandible and maxilla, ventral side F left mandible and maxilla, ventral side. Abbreviations: bo – birth opening, cl – clypeal area, dlf – dorsal labral field of labral area, im – impression, lb – labial area, md – mandible, mdb – mandibular bulge, mdt – mandibular tooth, mx – vestige of maxilla (maxilla), mxb – maxillary base (at mandible base), os – mouth opening, p – papilla, sbhp – segmental border between head and prothorax, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure6" httpUri="https://binary.pensoft.net/fig/1130332">6 C</figureCitation>
).
</paragraph>
<paragraph id="C648B75BDEDEC402BE795E9BE3A1D5AA">
<emphasis id="190EC6343B184C139953B4BE94EF01AC" bold="true">
<emphasis id="2504CCBAF7140CA3AFA414224C6F6107" bold="true" italics="true">Thorax</emphasis>
.
</emphasis>
Pro-mesothoracic and meso-metathoracic borders visible ventrally as slightly imprinted mesal furrows (sbpm, sbmm; Figs
<figureCitation id="0CB5671A461400D50ACA96AD3168B0A3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, female cephalothorax A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with female cephalothorax C – D holotype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle." figureDoi="10.3897/jhr.97.127500.figure4" httpUri="https://binary.pensoft.net/fig/1130330">4 C</figureCitation>
,
<figureCitation id="0223D5F489A538576E4B3AE17112607A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 A</figureCitation>
). On dorsal side separated by conspicuous dark mesal furrows, distinctly contrasted with pale thoracic segments (sbpm, sbmm; Figs
<figureCitation id="419D38B9A8BBB3D34FDA08935CA04F21" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, female cephalothorax A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with female cephalothorax C – D holotype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle." figureDoi="10.3897/jhr.97.127500.figure4" httpUri="https://binary.pensoft.net/fig/1130330">4 D</figureCitation>
,
<figureCitation id="D52755B3FA9B0802B7392B3810D0ED2C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 B</figureCitation>
).
<collectingRegion id="3DBF0085096743BC8EB1D4A0D3D4951A" country="Saudi Arabia" name="Al Hudud ash Shamaliyah">Border</collectingRegion>
between metathorax and abdomen indicated by ventral ridge on ventral side or indicated by change in colour and cuticular sculpture. Cuticle of thoracic segments on ventral side reticulate with scattered inconspicuous or more distinct pigmented papillae, mainly visible on metasternum. Prosternum conspicuously dark, mesosternum brown, and metasternum light brown. Prosternal extension undifferentiated. Dorsal side of thorax usually completely smooth, rarely with papillae on prothorax. All thoracic segments dorsally pale, but dark laterally. Meso- and metathorax transverse, rarely slightly elongated.
</paragraph>
<paragraph id="57B2D2EEB250DFB076EBA22CB6B37611">
<emphasis id="1115820897D8ADBD9C9B8A1B80308EE7" bold="true">
<emphasis id="966EC1FE0318E3C85339B7DA782A8D17" bold="true" italics="true">Abdominal segment I and spiracles</emphasis>
.
</emphasis>
Setae and cuticular spines present on lateral region of abdominal segment I posterior to spiracle (Fig.
<figureCitation id="D20BF882EE0B653192AC851816958B1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Deltoxenos impressus Benda &amp; Straka, sp. nov., ♀ (OLML), SEM micrographs of cephalothorax A ventral side B dorsal side C right vestigial antenna, dorsal side D left vestigial antenna, dorsal side E left lateral border of abdominal segment I below spiracle, dorsal side F detail of anterior border of cephalothorax, dorsal side. Abbreviations: a – vestigial antenna, asI – abdominal segment I, cls – clypeal sensillum, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, paa – periantennal area, pst – prosternum (prosternal extension), sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax." figureDoi="10.3897/jhr.97.127500.figure5" httpUri="https://binary.pensoft.net/fig/1130331">5 E</figureCitation>
). Spiracles on posterior ~ 2 / 5 of cephalothorax slightly elevated, with anterolateral orientation. Cephalothoracic part of abdominal segment I below spiracles dark brown and medially light brown on both sides (asI, Fig.
<figureCitation id="EA231FF07E5460E7979AF98B92367CB4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, female cephalothorax A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with female cephalothorax C – D holotype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle." figureDoi="10.3897/jhr.97.127500.figure4" httpUri="https://binary.pensoft.net/fig/1130330">4 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXEAG" type="diagnosis">
<paragraph id="ED971919092C2D48FD0DFD51B1BF17EF">
<heading id="729E292DC2D468C0522A1484F134F785" reason="title">Diagnosis of male cephalotheca.</heading>
</paragraph>
<paragraph id="C700CC6237293C92F45519E6227815FE">
This species is distinguished from other representatives of the genus
<taxonomicName id="1FD6CAB4481834C1693B0A4D9D58C16B" authorityName="Benda, Pohl, Nakase, Beutel &amp; Straka" authorityYear="2022" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="genus">
<emphasis id="09167EFE2EAB08BAE2AB295DBDC188FB" italics="true">Deltoxenos</emphasis>
</taxonomicName>
by a combination of the following characters: clypeal lobe distinctly arcuate in frontal view versus conspicuously bulging as in
<taxonomicName id="EA0470135F0AE75BB230A69C5C4D1398" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="B961FD38ACA91F22A1C991D52F3A1F77" italics="true">D. rueppelli</emphasis>
</taxonomicName>
; shape of cephalotheca rounded in frontal view, very slightly flattened (Fig.
<figureCitation id="50490B90EDE9DB8B7C7C3E35B92BA2CA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
) versus more flattened in
<taxonomicName id="1BB10A802082D6DD6A478C75754CDABC" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="64DD208814768D4B3437A8883DE7D42C" italics="true">D. rueppelli</emphasis>
</taxonomicName>
; frontal region with distinct impression versus impression not visible in
<taxonomicName id="7DB698DD8F61BEFBDE710229BD110C3A" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="733D3F615750C01BD0D2167C4FE73CD6" italics="true">D. rueppelli</emphasis>
</taxonomicName>
and
<taxonomicName id="1DD69703C08D33B36B1C9FC06402A7ED" baseAuthorityName="Luna de Carvalho" baseAuthorityYear="1956" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="305D68AA97B6F03ED197BA4317349BE1" italics="true">D. bequaerti</emphasis>
</taxonomicName>
; occipital bulge not protruding from elliptic shape of cephalotheca versus occipital bulge protruding in
<taxonomicName id="E2F167AACDF77E6ED7B602813EDFBF71" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="0236D664FD559611236BAC540BACD00B" italics="true">D. rueppelli</emphasis>
</taxonomicName>
and
<taxonomicName id="EF2F238F35D59C8E7E2A32CF1CF73DBD" baseAuthorityName="Luna de Carvalho" baseAuthorityYear="1956" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="D24D4325E732BC3ACF7A9B1EE9BF88BA" italics="true">D. bequaerti</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="598677097C5320A09E96C7941CF7C8F9" ID-DOI="10.3897/jhr.97.127500.figure7" ID-arpha="C55C938C-CC34-5C2D-8749-0A5573E6B721" httpUri="https://binary.pensoft.net/fig/1130333" startId="F7">
<paragraph id="BC463A8E9C1C3FD139AEEE0B33CA53D8">
<label id="62651CBE56C447A67E321E135F21BF00">Figure 7.</label>
</paragraph>
<paragraph id="29B614DE6D062FCDF93683F32C02694A">
<taxonomicName id="78E0A3FD8079EA98C87A8572F5D39D24" authority="Benda &amp; Straka" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="C7902082B6583E9C60BD0FEC0832EC6A" italics="true">Deltoxenos impressus</emphasis>
Benda &amp; Straka
</taxonomicName>
,
<taxonomicNameLabel id="99D43B96E271630450E0345EAD2A6463" rank="species">sp. nov.</taxonomicNameLabel>
, host, male cephalotheca
<emphasis id="57DFD98D0A41CC593BCEC83A472BD895" bold="true">A</emphasis>
<taxonomicName id="AB81B3987110BBA386A3BAC288FD4B68" authority="Walker, 1871" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="7108076596BA83DA5962F4E6F4041200" italics="true">Zethus favillaceus</emphasis>
Walker, 1871
</taxonomicName>
stylopised by
<taxonomicName id="65D50DC6684C7ECFDBBC0682CEB3C171" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="28C50A998FDE49156D8A1C66827AFF60" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3C5D27D21E1E00FA2531D52F310DA47B" rank="species">sp. nov.</taxonomicNameLabel>
, lateral view
<emphasis id="BE50CF82C59CCCA1EBFD362AF95A4723" bold="true">B</emphasis>
detail of host abdomen of
<taxonomicName id="5E31A6B18BB766D4E83DC000695D1807" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="9A186794815C63D4AD626B204C69A076" italics="true">Z. favillaceus</emphasis>
</taxonomicName>
, with male cephalotheca
<emphasis id="A7508DD389B19EB59C83860A8318E56E" bold="true">C – D</emphasis>
paratype of
<taxonomicName id="EE5EF687246367D87DD780A4C90AFF9D" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="C33415953B5C939B4F62D4CA11BFA9D6" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="912F81CB43E263F088108DA3B098E56E" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (
<named-content id="6D54E565CE1AF01EFCD52848C40D2DAA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ober%C3%B6sterreichisches-landesmuseum" xlink_title="Oberösterreichisches Landesmuseum">OLML</named-content>
) from
<taxonomicName id="37CAC65773DCB241C503E59EAF55302C" authorityName="Walker" authorityYear="1871" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="favillaceus">
<emphasis id="87355E9D52712E147F0EDA5FC9106F8C" italics="true">Z. favillaceus</emphasis>
</taxonomicName>
<emphasis id="65999217EEA06CDEC421B10146C2AEBF" bold="true">C</emphasis>
ventral side of cephalothorax
<emphasis id="53332FC51833609B6992632A0943FF94" bold="true">D</emphasis>
dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area.
</paragraph>
</caption>
<paragraph id="0302DEB6E340C4DD3F87F6BFDA714F9A">
Other characters that distinguish
<taxonomicName id="B4EDCFB3C75F931A20C6C0B5C45DE151" authorityName="Benda &amp; Pohl &amp; Beutel &amp; Straka" authorityYear="2024" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="impressus" status="sp. nov.">
<emphasis id="03E28D70CACEC40FC8D3BABF7F1EF860" italics="true">D. impressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="459DFD93D260D5038A9EBE2747D715D8" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="EB5413DAE5EEBAA79ADDD76473751811" baseAuthorityName="Kinzelbach" baseAuthorityYear="1971" class="Insecta" family="Xenidae" genus="Deltoxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Strepsiptera" phylum="Arthropoda" rank="species" species="rueppelli">
<emphasis id="EFECB998C7A70D6DB1BC393718B8550F" italics="true">D. rueppelli</emphasis>
</taxonomicName>
: clypeal lobe as wide as mandible versus clypeal lobe distinctly wider than mandible; gena completely dark (gn; Fig.
<figureCitation id="29F2E357ABBD8DACE3EE968231C9C043" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
) versus light brown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDLAG" type="description">
<paragraph id="F497A840931523EA2EB83A72BC3245C3">
<heading id="D7B34675532F78CE9538C6922C89D629" reason="title">Description of male cephalotheca.</heading>
</paragraph>
<paragraph id="FACDA6F103D3E83FC7BB7FF0D2FD6220">
<emphasis id="B916DB415BD3738F292AEB91A5A9B2C2" bold="true">
<emphasis id="1EBEF5E32A4912367529F1AC7915D505" bold="true" italics="true">Shape and colouration</emphasis>
.
</emphasis>
In frontal view rounded, very slightly flattened, elliptic, length
<quantity id="B0056FD491307EE43D6430620DFC47D6" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
, width
<quantity id="CE553810992A98A565EC8F14BE43652E" metricMagnitude="-4" metricUnit="m" metricValue="8.6" unit="mm" value="0.86">0.86 mm</quantity>
, in lateral view protruding anteriorly, pointed apically (Fig.
<figureCitation id="AD343D5AE5E8ACBB0685B74CFE0C4483" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C, D</figureCitation>
). Colouration predominantly dark with paler areas.
</paragraph>
<paragraph id="9222A692111FA38689082B0BB9A6CC61">
<emphasis id="6F8FC6D7C012EA77C51E07D322F0244C" bold="true">
<emphasis id="0A0D0E0BFB74D61F5EAEBF08421CB504" bold="true" italics="true">Cephalothecal capsule</emphasis>
.
</emphasis>
Compound eyes visible, pale to dark, with dark individual cornea lenses. Clypeal lobe distinctly arcuate in frontal view, prominent in lateral view (cl; Fig.
<figureCitation id="C4FA3A6D7C3781EC1BD9868404C1AA18" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C, D</figureCitation>
). Clypeal area completely dark, clypeal sensilla indistinct. Clypeal lobe as wide as mandible.
</paragraph>
<paragraph id="7DB9A256C3F74A7348F9F31A30916B78">
Frontal region with paired furrow of supra-antennal sensillary field, distinct impression and slightly raised occipital bulge (ssf, fi, ob; Fig.
<figureCitation id="63341E2B5C274E8AB8447E71EEDB761B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Occipital bulge not protruding from elliptic outline of cephalotheca. Gena completely dark (gn; Fig.
<figureCitation id="64FA6E897F08EE2DB9AD8E1A4BAE70C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Diameter of genae between maxillary base and compound eye approximately 3 × larger than diameter of vestigial antenna.
</paragraph>
<paragraph id="DC7CBC3D95B0D4769C04D9E098859CA2">
<emphasis id="75F5120E681F3AAB05AF014E3E56E554" bold="true">
<emphasis id="220C9DDBC1D665C7FCDD4FDA88A86F18" bold="true" italics="true">Supra</emphasis>
-
<emphasis id="B3B0AC39D0D9D9552832F6412525A5FA" bold="true" italics="true">antennal sensillary field</emphasis>
.
</emphasis>
Dark, kidney-shaped and bulging, not delimited medially by distinct furrow (ssf; Fig.
<figureCitation id="11961AE35F77244BF4633E46B7798A2A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7</figureCitation>
). Dark sensilla well visible.
</paragraph>
<paragraph id="305F54FC0721FCCCBF41532C6E8CCE79">
<emphasis id="DF9429EF9B54E1271B850128BA51F033" bold="true">
<emphasis id="C2FD3E52B125BA1D4A37EECC21197C73" bold="true" italics="true">Antenna</emphasis>
.
</emphasis>
Of standard shape, dark, small, with small plates or sensilla and complete torulus (a; Fig.
<figureCitation id="29DA224EBFC0A2C52E1C101B6B75C9EA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Periantennal area not clearly delimited from supra-antennal sensillary field. Small plates and sensilla present.
</paragraph>
<paragraph id="06113B4C2B00187B201107F8338EB709">
<emphasis id="78651B96290024251D3A041A8779571F" bold="true">
<emphasis id="8DBDD30B3C40A1CCB026F5B96079078F" bold="true" italics="true">Labrum</emphasis>
.
</emphasis>
Labral area distinct, slightly less dark medially. Dorsal field arcuate, with dispersed setae visible (dlf; Fig.
<figureCitation id="D5DB78212C657152703B7B165C1F1EEE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Ventral field elliptic.
</paragraph>
<paragraph id="C9FFACEC6938B4145602172187E1F0F5">
<emphasis id="9B5893AEEB6237D41D043A02F369C685" bold="true">
<emphasis id="2FE1560CD7F66BF5517A9CE84AFEFD49" bold="true" italics="true">Mandible</emphasis>
.
</emphasis>
Nearly medially directed (md; Fig.
<figureCitation id="891CD01AB07B48817914DC1849D9225C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, female cephalothorax A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with female cephalothorax C – D holotype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: asI – abdominal segment I, cll – clypeal lobe, csI – constriction of abdominal segment I, lehc – lateral extension of head capsule, mst – mesosternum, mtst – metasternum, pst – prosternum (prosternal extension), sbhp – segmental border between head and prothorax, sbma – segmental border between metathorax and abdomen, sbmm – segmental border between mesothorax and metathorax, sbpm – segmental border between prothorax and mesothorax, sp – spiracle." figureDoi="10.3897/jhr.97.127500.figure4" httpUri="https://binary.pensoft.net/fig/1130330">4 E</figureCitation>
). Colouration predominantly dark but slightly lighter medially. Distance between mandibles very distinctly exceeding mandibular length.
<emphasis id="19EDD97E288671B840829F6D301C29A7" bold="true">
<emphasis id="D086A6431171134CA29CB70C75DDD9C4" bold="true" italics="true">Maxilla</emphasis>
.
</emphasis>
Distinct, prominent, completely dark with inconspicuous paler spot anteriorly. Wide at base, approximately 2 × as wide as mandible (mx; Fig.
<figureCitation id="812BC1483F46657916BF01DB9CB1768B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Vestige of palp present, conspicuous (mxp; Fig.
<figureCitation id="F26DB72204B5DF5A0A6A03841A125FD7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
).
</paragraph>
<paragraph id="3CEEC6908F5D9508908C863A70CBD55D">
<emphasis id="CB63F6768CEFBDC52C0E4BCF60413EB7" bold="true">
<emphasis id="6E3DA5A7A49FAE48805C660CA1E9EC23" bold="true" italics="true">Labium and hypopharynx</emphasis>
.
</emphasis>
Labium distinct between and below maxillae, completely dark. Praementum and postmentum separated by furrow. Hypopharyngeal protuberance present but very indistinct (hyp; Fig.
<figureCitation id="C2ED44E065ED86775219862B56EBFDBA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Deltoxenos impressus Benda &amp; Straka, sp. nov., host, male cephalotheca A Zethus favillaceus Walker, 1871 stylopised by D. impressus sp. nov., lateral view B detail of host abdomen of Z. favillaceus, with male cephalotheca C – D paratype of D. impressus sp. nov., ♀ (OLML) from Z. favillaceus C ventral side of cephalothorax D dorsal side of cephalothorax. Abbreviations: a – vestigial antenna, cl – clypeal area, coe – compound eye, dlf – dorsal labral field of labral area, fr – frontal region, gn – gena, md – mandible, mx – vestige of maxilla, mxp – vestige of maxillary palp, ob – occipital bulge, pom – postmentum, prm – praementum, ssf – supra-antennal sensillary field, vlf – ventral labral field of labral area." figureDoi="10.3897/jhr.97.127500.figure7" httpUri="https://binary.pensoft.net/fig/1130333">7 C</figureCitation>
). Mouth opening well visible, not covered by ventral labral field, distinctly arcuate.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMOAG" type="etymology">
<paragraph id="4DE97A4A35B89B8D6976CCFE94032B9E">
<heading id="6D04A2D336D6FEC27EBF711D26A62304" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="4517328973E3080638C8A510BB054B07">
From the Latin substantive
<emphasis id="A19C185939CB80374792E07D1D0A09AF" italics="true">impressio</emphasis>
, meaning an impression or hole. The specific epithet refers to conspicuous impressions on the lateral extensions of the female cephalothorax. Adjective.
</paragraph>
</subSubSection>
</treatment>
</document>
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