treatments-xml/data/0C/BF/52/0CBF527E079157E8A95AA72A8E86DA88.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="0CBF527E079157E8A95AA72A8E86DA88" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia friedmaniorum Kriebel 2016, comb. nov." docType="treatment" docVersion="3" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="178" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="174" updateTime="1668141747722" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kriebel, Ricardo</mods:namePart>
<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-07-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>67</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>326</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226942" LSID="urn:lsid:plazi:treatment:0CBF527E079157E8A95AA72A8E86DA88" httpUri="http://treatment.plazi.org/id/0CBF527E079157E8A95AA72A8E86DA88" lastPageId="177" lastPageNumber="178" pageId="173" pageNumber="174">
<subSubSection pageId="173" pageNumber="174" type="nomenclature">
<paragraph pageId="173" pageNumber="174">
<taxonomicName LSID="0CBF527E-0791-57E8-A95A-A72A8E86DA88" authority="(Almeda &amp; Umana) Kriebel" authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="173" pageNumber="174" phylum="Tracheophyta" rank="species" species="friedmaniorum" status="comb. nov.">
Conostegia friedmaniorum (Almeda &amp; 
<normalizedToken originalValue="Umaña">Umana</normalizedToken>
) Kriebel
</taxonomicName>
<taxonomicNameLabel pageId="173" pageNumber="174">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 158" captionStartId="F158" captionText="Figure 158. Conostegia friedmaniorum. A Habit. Note asymmetric leaf base B Habit showing deflexed inflorescence C Close up of flower D Infructescence E Flower bud F Longitudinal section of a flower bud G-H Pickled flowers I Petal J Stamen. Photos of A, D of specimen vouchered R. Kriebel 5497, and B, D R. Kriebel 5641." figureDoi="10.3897/phytokeys.67.6703.figure158" httpUri="https://binary.pensoft.net/fig/96516" pageId="173" pageNumber="174">Fig. 158</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="174" pageNumber="175" type="reference_group">
<paragraph pageId="174" pageNumber="175">
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="174" pageNumber="175" phylum="Tracheophyta" rank="species" species="friedmaniorum" status="comb. nov.">
<pageBreakToken pageId="174" pageNumber="175" start="start">Conostegia</pageBreakToken>
friedmaniorum
</taxonomicName>
(Almeda &amp; 
<normalizedToken originalValue="Umaña">Umana</normalizedToken>
) Kriebel. Basionym: 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia friedmaniorum" order="Myrtales" pageId="174" pageNumber="175" phylum="Tracheophyta" rank="species" species="friedmaniorum" status="comb. nov.">Miconia friedmaniorum</taxonomicName>
Almeda &amp; 
<normalizedToken originalValue="Umaña">Umana</normalizedToken>
, Novon 3(1): 5, f. 1. 1993. Type: Costa Rica. Alajuela: Upala, Colonia Libertad, subiendo hasta el Llano Aguacatales, 
<geoCoordinate degrees="10" direction="north" minutes="48" orientation="latitude" precision="15" seconds="25" value="10.806945">10 48'25&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="85" direction="west" minutes="17" orientation="longitude" precision="15" seconds="50" value="-85.29722">85 17'50&quot;W</geoCoordinate>
, 1500 m, 28 April 1988, G. Herrera 1900 (holotype: CR!, isotype: BR!, CAS!, F!, MEXU!, MO!, PMA, SAC, USJ).
</paragraph>
</subSubSection>
<subSubSection lastPageId="176" lastPageNumber="177" pageId="174" pageNumber="175" type="description">
<paragraph pageId="174" pageNumber="175">Description.</paragraph>
<paragraph lastPageId="176" lastPageNumber="177" pageId="174" pageNumber="175">
Shrub to small tree 2-5 m tall with somewhat tetragonal and ridged stems in newer branches that are densely covered with inconspicuously stalked reddish-orange asperous-headed hairs usually also with some simple multicellular hairs on the distal internodes; the nodal line nodal line present. Leaves of a pair equal to somewhat unequal in length. Petioles 0.5-4.8 cm. Leaf blades 7.5-18.6 
<normalizedToken originalValue="×">x</normalizedToken>
3.4-7.7 cm, 5-plinerved, with the innermost pair of veins arising up to about 4 cm above the base in opposite or frequently in strongly alternate fashion, elliptic to elliptic-ovate, the base obtuse to oblique, the apex acuminate,the margin entire to inconspicuously denticulate, the adaxial surface mostly glabrous, moderately to sparsely covered with a mixture of inconspicuously asperous headed and scalelike multicellular hairs on the secondary and higher order veins abaxially. Inflorescence a mostly deflexed terminal modified cyme branching at the base, 3.6-9.8 cm long, accessory branches absent, rachis covered with stalked asperous headed hairs, linear oblong to triangular, bracteoles 1.5-4 mm, persistent. Pedicel 1-1.75 mm long. Flowers 5-merous, calyx in bud not calyptrate but closed in bud and crowned by an apiculum, rupturing irregularly at anthesis into 4-5 hyaline irregular lobes 0.3-1.5 mm long, flower buds 3-4 
<normalizedToken originalValue="×">x</normalizedToken>
2-2.5 mm, the hypan 
<pageBreakToken pageId="175" pageNumber="176" start="start">thium</pageBreakToken>
1.5-2.5 
<normalizedToken originalValue="×">x</normalizedToken>
1.75-2.25 mm, campanulate, covered with orangish asperous headed hairs; exterior calyx teeth conspicuous, triangular, 0.5-0.6 mm long, torus glabrous. Petals 3-3.5 
<normalizedToken originalValue="×">x</normalizedToken>
0.9-1.25 mm, translucent pink, linear-oblong, reflexed at anthesis, 
<pageBreakToken pageId="176" pageNumber="177" start="start">glabrous</pageBreakToken>
, the apex acute. Stamens 10, 2.2-3 mm, radially arranged around the style, the filaments 1-1.75 mm, geniculate near the apex, white, anthers 1-1.25 
<normalizedToken originalValue="×">x</normalizedToken>
0.25-0.75 mm, linear-oblong, yellow, laterally compressed, the pore 0.1-0.14 mm wide, terminal and slightly dorsally inclined. Ovary 5 locular, 3/4 inferior, slightly fluted and with glandular hairs on the apex. Style 5.25-5.5 mm long, slightly to strongly bending, vertical distance from anther pore to stigma 1.75-2.25 mm, horizontal distance absent or up to1.3 mm, stigma truncate and somewhat dilated, 0.25-35 mm wide. Berry 3-4 
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm, red turning purple black when mature. Seeds 0.39-0.6 mm, angular pyramidate to somewhat cescent shaped in profile, the testa smooth.
</paragraph>
<caption ID-Zenodo-Dep="973967" doi="10.3897/phytokeys.67.6703.figure158" httpUri="https://binary.pensoft.net/fig/96516" pageId="176" pageNumber="177" start="Figure 158" startId="F158">
<paragraph pageId="176" pageNumber="177">
<emphasis bold="true" pageId="176" pageNumber="177">Figure 158.</emphasis>
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="176" pageNumber="177" phylum="Tracheophyta" rank="species" species="friedmaniorum">
<emphasis italics="true" pageId="176" pageNumber="177">Conostegia friedmaniorum</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="176" pageNumber="177">A</emphasis>
Habit. Note asymmetric leaf base 
<emphasis bold="true" pageId="176" pageNumber="177">B</emphasis>
Habit showing deflexed inflorescence 
<emphasis bold="true" pageId="176" pageNumber="177">C</emphasis>
Close up of flower 
<emphasis bold="true" pageId="176" pageNumber="177">D</emphasis>
Infructescence 
<emphasis bold="true" pageId="176" pageNumber="177">E</emphasis>
Flower bud 
<emphasis bold="true" pageId="176" pageNumber="177">F</emphasis>
Longitudinal section of a flower bud 
<emphasis bold="true" pageId="176" pageNumber="177">G-H</emphasis>
Pickled flowers 
<emphasis bold="true" pageId="176" pageNumber="177">I</emphasis>
Petal 
<emphasis bold="true" pageId="176" pageNumber="177">J</emphasis>
Stamen. Photos of 
<emphasis bold="true" pageId="176" pageNumber="177">A, D</emphasis>
of specimen vouchered 
<emphasis italics="true" pageId="176" pageNumber="177">R. Kriebel 5497</emphasis>
, and 
<emphasis bold="true" pageId="176" pageNumber="177">B, D</emphasis>
<emphasis italics="true" pageId="176" pageNumber="177">R. Kriebel 5641</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177" type="distribution">
<paragraph pageId="176" pageNumber="177">Distribution</paragraph>
<paragraph pageId="176" pageNumber="177">
(Fig. 
<figureCitation captionStart="Figure 159" captionStartId="F159" captionText="Figure 159. Distribution of Conostegia friedmaniorum." figureDoi="10.3897/phytokeys.67.6703.figure159" httpUri="https://binary.pensoft.net/fig/96517" pageId="176" pageNumber="177">159</figureCitation>
). Endemic to the Caribbean slope of the Cordillera de 
<normalizedToken originalValue="Tilarán">Tilaran</normalizedToken>
and 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
Tenorio National Park in Costa Rica, 1250-1700 m in elevation.
</paragraph>
<caption ID-Zenodo-Dep="973969" doi="10.3897/phytokeys.67.6703.figure159" httpUri="https://binary.pensoft.net/fig/96517" pageId="176" pageNumber="177" start="Figure 159" startId="F159">
<paragraph pageId="176" pageNumber="177">
<emphasis bold="true" pageId="176" pageNumber="177">Figure 159.</emphasis>
Distribution of 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="176" pageNumber="177" phylum="Tracheophyta" rank="species" species="friedmaniorum">
<emphasis italics="true" pageId="176" pageNumber="177">Conostegia friedmaniorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177">
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="176" pageNumber="177" phylum="Tracheophyta" rank="species" species="friedmaniorum">
<emphasis italics="true" pageId="176" pageNumber="177">Conostegia friedmaniorum</emphasis>
</taxonomicName>
can be recognized by its rusty pubescence, leaves which are strongly plinerved with the veins frequently arising in asymmetric fashion, 
<pageBreakToken pageId="177" pageNumber="178" start="start">deflexed</pageBreakToken>
inflorescences and linear-oblong petals. Vegetatively it resembles another Costa Rican endemic, 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia pendula</emphasis>
</taxonomicName>
, as well as the Panamanian endemics 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia galdamesiae" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="galdamesiae">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia galdamesiae</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia papillopetala</emphasis>
</taxonomicName>
. From 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Umana &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia pendula" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia pendula</emphasis>
</taxonomicName>
it can be distinguished because the latter has lanate indument on the stems which is absent in 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Almeda &amp; Umana" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia friedmaniorum" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="friedmaniorum">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia friedmaniorum</emphasis>
</taxonomicName>
. From 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia galdamesiae" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="galdamesiae">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia galdamesiae</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel &amp; Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia papillopetala" order="Myrtales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="species" species="papillopetala">
<emphasis italics="true" pageId="177" pageNumber="178">Conostegia papillopetala</emphasis>
</taxonomicName>
it can be distinguished by its deflexed inflorescence and linear-oblong petals.
</paragraph>
</subSubSection>
<subSubSection pageId="177" pageNumber="178" type="materials_examined">
<paragraph pageId="177" pageNumber="178">Specimens examined.</paragraph>
<paragraph pageId="177" pageNumber="178">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Rio Penas Blancas, R. B, P. N. Volcan Tenorio, Estacion El Pilon, de Tilaran, El Valle" country="COSTA RICA" county="Monteverde Reserve" location="Cordillera de Tilaran" municipality="Valle del Rio Penas Blancas" specimenCount="1" stateProvince="Alajuela">
<emphasis bold="true" pageId="177" pageNumber="178">
<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
. 
<collectingRegion country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
</emphasis>
: 
<collectingCounty>Monteverde Reserve</collectingCounty>
, Cerro Negro, continental divide with Atlantic Slope exposure, Bello 3242 (CAS); San 
<normalizedToken originalValue="Ramón">Ramon</normalizedToken>
, 
<collectingMunicipality>
Valle del 
<collectorName>
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken originalValue="Peñas">Penas</normalizedToken>
Blancas, R. B
</collectorName>
</collectingMunicipality>
. Monteverde, Bello 5155 (CAS); 
<collectorName>
P. N. 
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
Tenorio
</collectorName>
, 
<collectorName>
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
El 
<normalizedToken originalValue="Pilón">Pilon</normalizedToken>
</collectorName>
, sendero hacia Cerro Montezuma, Kriebel and Nicholas 5497 (INB, NY); loc. cit., Kriebel et al. 5641 (INB); Border of 
<collectingRegion country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
, 
<collectingRegion country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
, and 
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
Provinces, 
<location LSID="urn:lsid:plazi:treatment:0CBF527E079157E8A95AA72A8E86DA88:7ABEDF685A67EA32A46492700C7CB073" country="COSTA RICA" county="Monteverde Reserve" municipality="Valle del Rio Penas Blancas" name="Cordillera de Tilaran" stateProvince="Alajuela">
Cordillera 
<collectorName>
de 
<normalizedToken originalValue="Tilarán">Tilaran</normalizedToken>
</collectorName>
</location>
, sendero 
<collectorName>El Valle</collectorName>
, Almeda and Anderson 5416 (CAS); Border of 
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
<collectingRegion country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
border, 
<location LSID="urn:lsid:plazi:treatment:0CBF527E079157E8A95AA72A8E86DA88:A65CA1FEFD470CD129BABD740D743BCE" country="COSTA RICA" county="Monteverde Reserve" municipality="Valle del Rio Penas Blancas" name="Cordillera de Tilaran" stateProvince="Alajuela">
Cordillera de 
<normalizedToken originalValue="Tilarán">Tilaran</normalizedToken>
</location>
, 
<location LSID="urn:lsid:plazi:treatment:0CBF527E079157E8A95AA72A8E86DA88:8E69FF4C926F8F0A1AA079CCDEC7E60A" country="COSTA RICA" county="Monteverde Reserve" municipality="Valle del Rio Penas Blancas" name="Monteverde Cloud Forest Reserve" stateProvince="Alajuela">Monteverde Cloud Forest Reserve</location>
, sendero Brillante along the Continental Divide, Almeda and Daniel 7074 (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>