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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="0C7D2D2174BE5698B91539251E676276" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Tortulosa tortuosa" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="5E1C59F546AEDC60212D2F8948241503">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="D5D05D64E4C3ECD7AFF02B9B0C443AC0">Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier id="62F67E4719AC4E17B56B469FB98AFB04" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation id="8A51C43E61EC293D802B8E6224DD6BCB">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="C50D120DAE75D526EDD14A25B1E1E9BE">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="D1B89AE98B02D4AC348F2504335289DE">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="CAA06938076921210AC2FF5ABF39A026" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="17CF066F9D779BF511C81DCC562823BD">Panha, Somsak</mods:namePart>
<mods:affiliation id="B1DE46F176B4B2EC47EFF7B3336949EF">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier id="EB531A5A7BADC4ED3F3719B76FC73AE2" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<treatment id="0C7D2D2174BE5698B91539251E676276" ID-GBIF-Taxon="199838938" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276" httpUri="http://treatment.plazi.org/id/0C7D2D2174BE5698B91539251E676276" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupinellinae" scope_superFamily="Cyclophoroidea">
<subSubSection id="1892201E54C112D14AD93156E505EA8D" pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName id="582E1C1D2216F3D4ED80230F706155BE" ID-CoL="57HXN" LSID="0C7D2D21-74BE-5698-B915-39251E676276" authority="(Ferussac, 1821)" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
Tortulosa tortuosa (
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, 1821)
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<figureCitation id="8D1EF5FCA75A14C7CBC5506AFE6432E4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">Figs 8D</figureCitation>
<figureCitation id="7E1ECCA3D2684D4C8BAF1340044CA508" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">, 15</figureCitation>
<figureCitation id="6C5F75A45DE088164D2624D6B6B70192" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Infraspecific variation of shell shape and colour found in the same collecting locality of Tortulosa tortuosa, CUMZ 12166 from Tham Suea Temple, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure16" httpUri="https://binary.pensoft.net/fig/735377" pageId="0" pageNumber="1">, 16</figureCitation>
</paragraph>
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<paragraph id="177B2B54946C08CC6FA8D2AC001D9661" pageId="0" pageNumber="1">
<taxonomicName id="6E300F708739177F375B200CB02D57F7" class="Gastropoda" family="Pupinidae" genus="Turbo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turbo tortuosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosus">Turbo tortuosus</taxonomicName>
-
<bibRefCitation id="B39BCEDD7CCFD0F62E16CECADE27C052" author="Chemnitz, JH" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B21" refString="Chemnitz, JH, 1795. Neues systematisches Conchylien-Cabinet. Bey Gabriel Nikolaus Raspe, Nuernberg, vol.11: 1-310. [pls 174-213]" title="Neues systematisches Conchylien-Cabinet. Bey Gabriel Nikolaus Raspe, Nuernberg, vol. 11: 1 - 310. [pls 174 - 213]" year="1795">Chemnitz 1795</bibRefCitation>
: 158, 159, pl. 195, figs 1882, 1883. Type locality: Nicobarischen Eylanden [Nicobar Islands]. Unavailable name.
</paragraph>
<paragraph id="7DB9789500557AD0026DCE4C229DB2F4" pageId="0" pageNumber="1">
<taxonomicName id="3285B2907EB4172C0B30589A910FE7B3" class="Gastropoda" family="Pupinidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Cochlodina) tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa" subGenus="Cochlodina">Helix (Cochlodina) tortuosa</taxonomicName>
<normalizedToken id="B3784AE5D279B13420F2C1B546379FC4" originalValue="Férussac">Ferussac</normalizedToken>
, 1821: 61.
</paragraph>
<paragraph id="9E937A9C3D8E8F220FC369DF84BD66D6" pageId="0" pageNumber="1">
<taxonomicName id="28C5655E88167C6AAA2FF461D091C000" class="Gastropoda" family="Pupinidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">Helix tortuosa</taxonomicName>
Pupa
<taxonomicName id="49FD07EE9E8BF62562AD170B270C67DD" class="Gastropoda" family="Pupinidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">Helix tortuosa</taxonomicName>
-
<bibRefCitation id="703F35198ECFDAB8E0D8C181D61877CC" author="Gray, JE" journalOrPublisher="Annals of Philosophy, new series" pageId="0" pageNumber="1" pagination="407 - 415" refId="B57" refString="Gray, JE, 1825. A list and description of some species of shells not taken notice of by Lamarck. Annals of Philosophy, new series 9: 407 - 415" title="A list and description of some species of shells not taken notice of by Lamarck." volume="9" year="1825">Gray 1825</bibRefCitation>
: 413.
</paragraph>
<paragraph id="27977E0405F94F51A09B348285102193" pageId="0" pageNumber="1">
<taxonomicName id="613537F437165E5548B1518693B29D8F" authorityName="G.B.Sowerby I" authorityYear="1843" class="Gastropoda" family="Pupinidae" genus="Cyclostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclostoma tortuosum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosum">Cyclostoma tortuosum</taxonomicName>
-Sowerby I 1843: 152, pl. 28, figs 185, 186.
<bibRefCitation id="3B5146796DA6EA651D3ADD2C173B54A9" DOI="https://doi.org/10.5962/bhl.title.119591" author="Pfeiffer, L (1848" journalOrPublisher="Of Sea and Shore" pageId="0" pageNumber="1" refId="B120" refString="Pfeiffer, L (1848, 1849[1843-1850]), Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV, 1-228, pls A, 1-30., [pp. 57-96, pls, 20-25, 1848. Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV1-228pls A1-30.[pp. 57-96pls20-25 (1848) ; pp. 97-144, pls 26-30 (1849 [sic]); pp. 145-176 (1849)] [Published in parts, dates follow Welter-Schultes (1999)] https://doi.org/10.5962/bhl.title.119591" title="Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV 1 - 228 pls A 1 - 30. [pp. 57 - 96 pls 20 - 25 (1848); pp. 97 - 144, pls 26 - 30 (1849 [sic]); pp. 145 - 176 (1849)] [Published in parts, dates follow Welter-Schultes (1999)]" url="https://doi.org/10.5962/bhl.title.119591" year="1848">Pfeiffer 1848</bibRefCitation>
: pl. 24, figs 19, 20.
<bibRefCitation id="17BA3420C526923AEC531EA84418D432" DOI="https://doi.org/10.5962/bhl.title.119591" author="Pfeiffer, L (1848" journalOrPublisher="Of Sea and Shore" pageId="0" pageNumber="1" refId="B120" refString="Pfeiffer, L (1848, 1849[1843-1850]), Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV, 1-228, pls A, 1-30., [pp. 57-96, pls, 20-25, 1848. Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV1-228pls A1-30.[pp. 57-96pls20-25 (1848) ; pp. 97-144, pls 26-30 (1849 [sic]); pp. 145-176 (1849)] [Published in parts, dates follow Welter-Schultes (1999)] https://doi.org/10.5962/bhl.title.119591" title="Die gedeckelten Lungenschnecken. (Helicinacea et Cyclostomacea.). In Abbildungen nach der Natur mit Beschreibungen. Systematisches Conchylien-Cabinet von Martini und Chemnitz 1 (19) [(1)]: I-IV 1 - 228 pls A 1 - 30. [pp. 57 - 96 pls 20 - 25 (1848); pp. 97 - 144, pls 26 - 30 (1849 [sic]); pp. 145 - 176 (1849)] [Published in parts, dates follow Welter-Schultes (1999)]" url="https://doi.org/10.5962/bhl.title.119591" year="1848">Pfeiffer 1849</bibRefCitation>
: 165, 166.
</paragraph>
<paragraph id="F06B45915B0B65CA62FD38CF9999302E" pageId="0" pageNumber="1">
<taxonomicName id="1251B4CCD7C4B896EAADC26AA9029CEF" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">Tortulosa tortuosa</taxonomicName>
<bibRefCitation id="E4A2A9142706928C0A2F90E11490FB22" author="Adams, H" pageId="0" pageNumber="1" refId="B1" refString="Adams, H, Adams, A (1856 [1854-1858]) The genera of Recent Mollusca; arranged according to their organization. van Voorst London, vol., 2, 661, pp. [pp. 285-412, 1856. The genera of Recent Mollusca; arranged according to their organization. van Voorst Londonvol.2661pp. [pp. 285-412 (1856)] [Published in parts, dates follow Trew (1992)]" title="The genera of Recent Mollusca; arranged according to their organization. van Voorst Londonvol. 2661 pp. [pp. 285 - 412 (1856)] [Published in parts, dates follow Trew (1992)]" year="1856">-Adams and Adams 1856</bibRefCitation>
: 285, pl. 86, fig. 2, 2a, b.
<bibRefCitation id="AE32B0BE0626E144C5310202DA0D91A4" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
: 190,?Nicobars; India: Trevandrum.
<bibRefCitation id="C8C5A8995C1DCB761C4118359E38E6FE" author="van Benthem Jutting, WSS" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="10 - 20" refId="B181" refString="van Benthem Jutting, WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
: 11, 12, limestone hill Kaki Bukit, near kampong Wang Tangga, Perlis [Malaysia].
<bibRefCitation id="3982778257D2C3832F170088E17BBF39" author="Berry, AJ" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="1 - 17" refId="B11" refString="Berry, AJ, 1963. An introduction to the non-marine molluscs of Malaya. Malayan Nature Journal 17: 1 - 17" title="An introduction to the non-marine molluscs of Malaya." volume="17" year="1963">Berry 1963</bibRefCitation>
: pl. 6, fig. 31.
<bibRefCitation id="5DF0D73364A0311C322407AF878D52CD" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
: 40, fig. 7, Wat Thum Sua, Nation Valley, near Krabi.
<bibRefCitation id="3DB5822CD1DE25832E73E940085AAB23" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 44.
<bibRefCitation id="6EEF9740110F3A304E981654838ED247" author="Sutcharit, C" journalOrPublisher="BRT, Bangkok" pageId="0" pageNumber="1" refId="B153" refString="Sutcharit, C, Panha, S, 2008. Land snails in Khao Nan National Park. BRT, Bangkok" title="Land snails in Khao Nan National Park." year="2008">Sutcharit and Panha 2008</bibRefCitation>
: 50, 51, with figs, Khao Nan National Park, Nakhon Si Thammarat, Thailand.
<bibRefCitation id="9443626BC5831DFAB9712C74943857F4" author="Egorov, R" journalOrPublisher="Proceedings of the Zoological Society of London, Series A" pageId="0" pageNumber="1" refId="B39" refString="Egorov, R, 2013. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. Treasure of Russian Shells Supplement 3 (Part 3): 1-62." title="A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. Treasure of Russian Shells Supplement 3 (Part 3): 1 - 62." year="2013">Egorov 2013</bibRefCitation>
: 14, 15, fig. 23.
<bibRefCitation id="693C3EEEAF8312F27CDE5DD14528103E" author="Raheem, DC" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B136" refString="Raheem, DC, Taylor, H, Ablett, J, Preece, RC, Aravind, NA, Naggs, F, 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1-294." title="A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1 - 294." year="2014">Raheem et al. 2014</bibRefCitation>
: 53, figs 9e, 30b, c.
<bibRefCitation id="A69712E0F441E57673D0DC2531FA5AA5" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 98.
<bibRefCitation id="C58DF8A26320B34C64326FCBB3E8B24B" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: 159, figs 5-11f, 5-13n.
<bibRefCitation id="CE4406EB20182FA9CAA644DDF4E6C290" author="Thach, NN" journalOrPublisher="48 HrBooks Company, Ohio, USA" pageId="0" pageNumber="1" refId="B164" refString="Thach, NN, 2018. New shells of South Asia seashells-freshwater &amp; land snails, 3 new genera, 132 new species &amp; subspecies. 48HrBooks Company, Ohio, USA" title="New shells of South Asia seashells-freshwater &amp; land snails, 3 new genera, 132 new species &amp; subspecies." year="2018">Thach 2018</bibRefCitation>
: 97 (figure caption), figs 139, 140.
<bibRefCitation id="EAEFCC638A45EF8EEB15A072E697BD5E" author="Meksuwan, P" journalOrPublisher="Burapha Science Journal" pageId="0" pageNumber="1" pagination="246 - 255" refId="B92" refString="Meksuwan, P, Phongsuwan, S, Dumrongrojwattana, P, 2020. Species and distribution of land snails in Phuket and adjacent islands. Burapha Science Journal 25: 246 - 255" title="Species and distribution of land snails in Phuket and adjacent islands." volume="25" year="2020">Meksuwan et al. 2020</bibRefCitation>
: 249, fig. 2, Tonsai Waterfall, Thalang District, Phuket Province.
<bibRefCitation id="6E4CFFF44CE8AFD6C676A046661C3AD4" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="B008E1104E297068300AFBE1DC80AFF9" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
: 41.
</paragraph>
<paragraph id="597E1B4C486874D2876F5F5E48D47C2F" pageId="0" pageNumber="1">
<taxonomicName id="3281113F6811D96AD099DA85A9868615" baseAuthorityName="Gray" baseAuthorityYear="1825" class="Gastropoda" family="Pupinidae" genus="Cataulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataulus (Tortulosa) tortuosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosus" subGenus="Tortulosa">Cataulus (Tortulosa) tortuosus</taxonomicName>
-Sowerby I 1866:
<taxonomicName id="718347804F60A7AD093E43FC1858BD27" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
, pl. 2 (pl. 264),
<taxonomicName id="4363CDE0E4FC1726B202F2CA2E474351" authorityName="L.Pfeiffer" authorityYear="1851" class="Gastropoda" family="Pupinidae" genus="Cataulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataulus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Cataulus</taxonomicName>
, fig. 1.
</paragraph>
<paragraph id="C0DD8C94E6787BBDDFBF9DAC9B84D170" pageId="0" pageNumber="1">
<taxonomicName id="E741A0FB8E633678847F1774936C0E4A" baseAuthorityName="Gray" baseAuthorityYear="1825" class="Gastropoda" family="Pupinidae" genus="Cataulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cataulus tortuosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosus">Cataulus tortuosus</taxonomicName>
-
<bibRefCitation id="3FA2441D9652FDFEDE8E3AF1D1C58FFE" author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve &amp; Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve &amp; Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
:
<taxonomicName id="450E077D85E0E7F106E96592B19B4AC7" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
, pl. 6, sp. 49.
<bibRefCitation id="60DBC62650C878CB7A845C94CF147D6E" author="Nevill, G" journalOrPublisher="Journal of the Asiatic Society of Bengal, Part II" pageId="0" pageNumber="1" pagination="125 - 167" refId="B105" refString="Nevill, G, 1881. New or little-known Mollusca of the Indo-Malayan fauna. Journal of the Asiatic Society of Bengal, Part II 50 (3): 125 - 167" title="New or little-known Mollusca of the Indo-Malayan fauna." volume="50" year="1881">Nevill 1881</bibRefCitation>
: 149.
</paragraph>
<paragraph id="1FE29774E7785932B68308288C1516A8" pageId="0" pageNumber="1">
<taxonomicName id="F63BD5C2D100282C3C1026EF97E9BFAB" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa (Tortulosa) tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa" subGenus="Tortulosa">Tortulosa (Tortulosa) tortuosa</taxonomicName>
-
<bibRefCitation id="1C73BEA7BAE9B99A0F3633E78572435C" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 288, fig. 64.
</paragraph>
<paragraph id="24F0BFAD21B2E2CE4D704A23CF657FD2" pageId="0" pageNumber="1">
<taxonomicName id="16A5B5B42C09D37899AC4C3C89ECCAF1" authorityName="Tomlin" authorityYear="1948" class="Gastropoda" family="Pupinidae" genus="Perlisia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perlisia tweediei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">Perlisia tweediei</taxonomicName>
Tomlin, 1948: 225, 226, pl. 11, fig. 6. Type locality: Kaki Bukit, Perlis [Malaysia].
<bibRefCitation id="BC03A41158CC1BDA32BA86413368E6C6" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="9650D24C23FD2F3DDFD2D73B46804219" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
: 41, fig. 3.
</paragraph>
<paragraph id="F9C75DB3B07A3F13F63F3FE7FCD46CD3" pageId="0" pageNumber="1">
<taxonomicName id="0DDAAEDB918A27EA5EC929AC99A07655" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tweediei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">Tortulosa tweediei</taxonomicName>
-
<bibRefCitation id="36F819C4292EAA8BA4731E5B5886B3C6" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 98.
</paragraph>
<paragraph id="A7F725AF431A53C1143D06E050EA83B7" pageId="0" pageNumber="1">
<taxonomicName id="A34069116CA1A98842948C6043221D75" authorityName="Thach" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa huberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huberi">Tortulosa huberi</taxonomicName>
Thach, 2018: 21, 22, figs 133-138. Type locality: Krabi, South Thailand.
<bibRefCitation id="44498F765E9B61CA0D8AF763FEF57473" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="24DB16DFC8A40CEFD5058BECCC74FF59" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
: 41, fig. 5.
</paragraph>
<paragraph id="C1FE6F93FD4459C39D9A25AC099B6BBF" pageId="0" pageNumber="1">
<taxonomicName id="A4D69752CE80D7889AB8B3D6BB701645" authorityName="Thach &amp; F. Huber" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa schileykoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="schileykoi">Tortulosa schileykoi</taxonomicName>
Thach &amp; Huber in Thach, 2018: 22, figs 142-146. Type locality: Phang Nga, South Thailand.
<bibRefCitation id="C97A4B15F04BB089813E7520DA5291EC" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="940B2E118539D2436B6F462B77B3C0E4" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
: 41, fig. 4.
</paragraph>
</subSubSection>
<subSubSection id="7FF569A5B937A49BB90207FAD5D8D941" pageId="0" pageNumber="1" type="type material">
<paragraph id="C3D19C8527640A64D970DB4F9EDEE1F2" pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph id="D0C52D8B976AF701812ECD5672945890" pageId="0" pageNumber="1">
<emphasis id="52CA283EC3D4398077F3F9D5432E0DA6" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
of
<taxonomicName id="9C4ED269BDDC73A41FA11317523287C9" authorityName="Tomlin" authorityYear="1948" class="Gastropoda" family="Pupinidae" genus="Perlisia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perlisia tweediei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">
<emphasis id="0799C6713876E85889F8302A75D6BB0F" italics="true" pageId="0" pageNumber="1">Perlisia tweediei</emphasis>
</taxonomicName>
NHMUK 1948.10.2.6 (Fig.
<figureCitation id="88D8A919AD81D5FDD5B9E62FABB4EE1E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">15A</figureCitation>
) from Kaki Bukit, Perlis.
<emphasis id="08C1463C0B9A8CE63D5A71EB24F2D5CB" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
of
<taxonomicName id="178F2A30748D7E9C10EA81DADFC5DF03" authorityName="Thach" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa huberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="8C61F4D246D8684953ED94EFB256BA6D" italics="true" pageId="0" pageNumber="1">Tortulosa huberi</emphasis>
</taxonomicName>
MNHN-IM-2000-34054 (Fig.
<figureCitation id="DE237B8EE1FFB0AB88EEE1E10E588A09" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">15B</figureCitation>
) from Krabi Province, Thailand.
<emphasis id="1F63A16C32E8EC45F1C5D09292FD8830" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
of
<taxonomicName id="EFE4824ECD7FBBE04384D3722ADDCBC9" authorityName="Thach &amp; F. Huber" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa schileykoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="schileykoi">
<emphasis id="728F10277F1B9F9F363109CAF2BD86E6" italics="true" pageId="0" pageNumber="1">Tortulosa schileykoi</emphasis>
</taxonomicName>
MNHN-IM-2000-34055 (Fig.
<figureCitation id="59F6B5D1AAACF6B9DD9DD25E224A23BD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">15C</figureCitation>
) from Phang Nga Province, Thailand.
</paragraph>
</subSubSection>
<subSubSection id="F6A67BA3F6FFC76286B607103B52AEB7" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="098A586122D97803A7989D504682A1B5" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="3E1FB22E57AAE6F8A1D82DEA46393E27" pageId="0" pageNumber="1">
<materialsCitation id="C2D307900F1987EC3C50949918B238FD" collectingDate="2009-01-16" country="Nicobar Islands" county="Krabi Province" location="Nai-Chong Silvicultural Research Station" municipality="Mueang Krabi District" specimenCount="1">
NHMUK 20100643/1-2 (2 shells) from the
<collectingCountry id="D0A7F834796580B78276A6E9C50F575B" name="India">Nicobar Islands</collectingCountry>
figured in
<bibRefCitation id="7F0F5C778DBB7544C7AF9D9101DCF935" author="Raheem, DC" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B136" refString="Raheem, DC, Taylor, H, Ablett, J, Preece, RC, Aravind, NA, Naggs, F, 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1-294." title="A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1 - 294." year="2014">Raheem et al. (2014</bibRefCitation>
: figs 30b, c). CUMZ 12154 (1 shell) from
<location id="A37AAFEEFF24E4B774A1A0997FD45867" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:A37AAFEEFF24E4B774A1A0997FD45867" country="Nicobar Islands" county="Krabi Province" municipality="Mueang Krabi District" name="Nai-Chong Silvicultural Research Station">Nai-Chong Silvicultural Research Station</location>
,
<collectingMunicipality id="6484C6A7110287FFA991929488866ACA">Mueang Krabi District</collectingMunicipality>
,
<collectingCounty id="496967C39D3E7BFD6EAB1B40E206DBB6">Krabi Province</collectingCounty>
,
<collectingDate id="DDB0DE3BA0CF510EE3D35C8BFE8E5A0C" value="2009-01-16">16 Jan. 2009</collectingDate>
</materialsCitation>
.
<materialsCitation id="3EC6B2E189DDBC821EB9D999E58C3100" collectingDate="2010-05-10" collectorName="Figs" country="India" location="Mueang Krabi District" municipality="Krabi Province" specimenCount="1">
CUMZ 12166 (&gt; 500 shells;
<collectorName id="872F7ECB5B867F76FE7C3001225C2464">Figs</collectorName>
<figureCitation id="93B5A070232133E7BB353AFE0C3BA633" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">15D</figureCitation>
,
<figureCitation id="A06158D4268162438CC6EC1F7BC064E2" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Infraspecific variation of shell shape and colour found in the same collecting locality of Tortulosa tortuosa, CUMZ 12166 from Tham Suea Temple, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure16" httpUri="https://binary.pensoft.net/fig/735377" pageId="0" pageNumber="1">16</figureCitation>
) from Tham Suea Temple,
<location id="BE2806FA0C132A9673A623452E9AD356" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:BE2806FA0C132A9673A623452E9AD356" country="India" municipality="Krabi Province" name="Mueang Krabi District">Mueang Krabi District</location>
,
<collectingMunicipality id="5E4BBEFD8A70CBD71A32A51A432D5B97">Krabi Province</collectingMunicipality>
,
<collectingDate id="F48B323311A68BB8AD96DA5C1F8C5EA1" value="2010-05-10">10 May 2010</collectingDate>
</materialsCitation>
.
<materialsCitation id="FA9A3D061FAACE2D87B0612307E000C2" collectingDate="2017-07-09" collectorName="Fig., Tham Suea Temple" country="India" location="Mueang Krabi District" municipality="Krabi Province" specimenCount="12">
CUMZ 12155 (
<specimenCount id="728366EC383476EAF6602772C81DA5BF" type="generic">12 specimens</specimenCount>
in ethanol;
<collectorName id="5233B7168337C09D65E5265CA20A25B8">Fig.</collectorName>
<figureCitation id="747064931463F5A6BD61F71EF5CF499F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8D</figureCitation>
) from
<collectorName id="1DCA7C1303547E315027D607BBDB6179">Tham Suea Temple</collectorName>
,
<location id="82CFBDD49E9E9639D9651081649D5133" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:82CFBDD49E9E9639D9651081649D5133" country="India" municipality="Krabi Province" name="Mueang Krabi District">Mueang Krabi District</location>
,
<collectingMunicipality id="0AB3B4401ABB9FD54AFC80F3874C0A70">Krabi Province</collectingMunicipality>
,
<collectingDate id="7F48D4F636C32B17C5FA456947270439" value="2017-07-09">9 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="74EB6CC26FB37AB7D9245364B919C067" collectingDate="2016-08-08" country="India" county="Phang Nga Province" location="Phung Chang Cave" municipality="Mueang Phang Nga District" specimenCount="2">
CUMZ 12156 (
<specimenCount id="2B92BA032D5303977E0538FCDFBBB1C4" type="generic">2 specimens</specimenCount>
in ethanol) from
<location id="8A8B31EE62B514F7A01064FBD798F150" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:8A8B31EE62B514F7A01064FBD798F150" country="India" county="Phang Nga Province" municipality="Mueang Phang Nga District" name="Phung Chang Cave">Phung Chang Cave</location>
,
<collectingMunicipality id="D30C47ED06B3E1AB2CD95B5033B68694">Mueang Phang Nga District</collectingMunicipality>
,
<collectingCounty id="43CE9A55B2AA9954355CE626C3D97A83">Phang Nga Province</collectingCounty>
,
<collectingDate id="80F75934B348D3D2460F2CAEE6F68D4F" value="2016-08-08">8 Aug. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation id="4236783584214D502B3A362A0519601B" collectingDate="2018-07-31" country="India" county="Phang Nga Province" location="Phung Chang Cave" municipality="Mueang Phang Nga District" specimenCount="1">
CUMZ 12157 (1 shell) from
<location id="E64F09A3D6150428A58A526667D258F8" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:E64F09A3D6150428A58A526667D258F8" country="India" county="Phang Nga Province" municipality="Mueang Phang Nga District" name="Phung Chang Cave">Phung Chang Cave</location>
,
<collectingMunicipality id="0A7221002DEDFD6086370507AEED8D45">Mueang Phang Nga District</collectingMunicipality>
,
<collectingCounty id="1B72CDF3018068E91B7B4824690DEED2">Phang Nga Province</collectingCounty>
,
<collectingDate id="2E415EAEE2015F3223B8AB5A4B50FA7E" value="2018-07-31">31 July 2018</collectingDate>
</materialsCitation>
.
<materialsCitation id="5FA994B607AB648B2FE235E95A80FBDD" collectingDate="2010-10-07" country="India" county="Phang Nga Province" location="Nam Phut Cave" municipality="Mueang Phang Nga District" specimenCount="1">
CUMZ 12188 (2 shells) from
<location id="8981422DA9DF5D8FD24D9197DB95818C" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:8981422DA9DF5D8FD24D9197DB95818C" country="India" county="Phang Nga Province" municipality="Mueang Phang Nga District" name="Nam Phut Cave">Nam Phut Cave</location>
,
<collectingMunicipality id="3190845A6AFE7EED4486E531C7A9978F">Mueang Phang Nga District</collectingMunicipality>
,
<collectingCounty id="A1FC2133BCBAE8A2A39964DDBAC50BB0">Phang Nga Province</collectingCounty>
,
<collectingDate id="08F807637FDA1C638DD49F3E55706A63" value="2010-10-07">7 Oct. 2010</collectingDate>
</materialsCitation>
.
<materialsCitation id="3F55566FD88CE89A166C2A223D77DADF" collectingDate="2016-08-07" collectorName="Ban Yai" country="India" location="Phanom District" municipality="Surat Thani Province" specimenCount="1">
CUMZ 12158 (
<specimenCount id="C1ACFE375CBD250FDF11267CC6608053" type="generic">1 specimen</specimenCount>
in ethanol) from
<collectorName id="F814A238E16AFA7ECB93594895D1DEC0">Ban Yai</collectorName>
,
<location id="D7B8EBD30EACCBC41956FDCCD20F1BAF" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:D7B8EBD30EACCBC41956FDCCD20F1BAF" country="India" municipality="Surat Thani Province" name="Phanom District">Phanom District</location>
,
<collectingMunicipality id="2B8568C17C8609AEE00B22D3CEA488F5">Surat Thani Province</collectingMunicipality>
,
<collectingDate id="6DD966059253B29716771DE9BB0FB2AB" value="2016-08-07">7 Aug. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation id="CF4A5CDA97E3E5090E416F0E62987FDC" collectingDate="2017-07-04" collectorName="Khiri Rat Phatthana Temple" country="India" location="Wiang Sa District" municipality="Surat Thani Province" specimenCount="18">
CUMZ 12159 (
<specimenCount id="F0E8A2EB6DD7CE5CCBC6C60C91D24AB8" type="generic">18 specimens</specimenCount>
in ethanol) from
<collectorName id="F150F50D66248BBB033B49DB5C8957D7">Khiri Rat Phatthana Temple</collectorName>
,
<location id="53E1CEF566BBF68ACC2136CED230B478" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:53E1CEF566BBF68ACC2136CED230B478" country="India" municipality="Surat Thani Province" name="Wiang Sa District">Wiang Sa District</location>
,
<collectingMunicipality id="B82D707CA4555C478A0D126D2B24D3D1">Surat Thani Province</collectingMunicipality>
,
<collectingDate id="F6A0D48217CC7F1FB72510322BFC7B5F" value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="B25FB525EBCA9F80110A1F06C2F0BA14" collectingDate="2008-12-08" collectorName="Natural Trail, Ratchaprapha Dam" country="India" location="Ban Ta Khun District" municipality="Surat Thani Province" specimenCount="1">
CUMZ 12189 (3 shells) from
<collectorName id="318D1899E99FE7E2EEF421E81B67DEEC">Natural Trail</collectorName>
,
<collectorName id="059FF72D26ED46A26289C7F48974E513">Ratchaprapha Dam</collectorName>
,
<location id="85DA633AD21DB1BFB777FBED9500D0CC" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:85DA633AD21DB1BFB777FBED9500D0CC" country="India" municipality="Surat Thani Province" name="Ban Ta Khun District">Ban Ta Khun District</location>
,
<collectingMunicipality id="8A073F0943FE3023C799A3DD2BF735B8">Surat Thani Province</collectingMunicipality>
,
<collectingDate id="5FFF85863EA0E8AA9C0C4455A13867E2" value="2008-12-08">8 Dec. 2008</collectingDate>
</materialsCitation>
.
<materialsCitation id="3E8B9012E9A173428DF26E22AD2F38B5" collectingDate="2017-07-04" collectorName="Fig., Tham Kanlayanamit Temple" country="India" location="Tham Phannara District" municipality="Nakhon Si Thammarat Province" specimenCount="3">
CUMZ 12160 (
<specimenCount id="E1722882C252B35A7B55231682A024F5" type="generic">3 specimens</specimenCount>
in ethanol;
<collectorName id="4419B2308161E6757D3F46723253AFD8">Fig.</collectorName>
<figureCitation id="FED2B863152E20194B16FC66B20A97E7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Tortulosa tortuosa A lectotype of Perlisia tweediei NHMUK 1948.10.2.6 from Kaki Bukit, Perlis B holotype of Tortulosa huberi MNHN-IM- 2000 - 34054 from Krabi C holotype of Tortulosa schileykoi MNHN-IM- 2000 - 34055 from Phang Nga D specimen CUMZ 12166 from Tham Suea Temple, Krabi, and E-H operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing E outer operculum F inner operculum G side view (inner surface up), and H spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (B, C)." figureDoi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1">15E-H</figureCitation>
) from
<collectorName id="FB1FADB3D51B1207B8EB5A3F6CBF76B3">Tham Kanlayanamit Temple</collectorName>
,
<location id="784C42A43F6A996054140FCAB126B95D" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:784C42A43F6A996054140FCAB126B95D" country="India" municipality="Nakhon Si Thammarat Province" name="Tham Phannara District">Tham Phannara District</location>
,
<collectingMunicipality id="964BAFFA06AEF35E3ACCD44E5BA15271">Nakhon Si Thammarat Province</collectingMunicipality>
,
<collectingDate id="5822501FB76E47456CAE76ED24A12338" value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="9A1764F6856AAF7C68CAEAED7386BCF7" collectingDate="2017-07-07" country="India" county="Satun Province" location="Ton Din Cave" municipality="Khuan Don District" specimenCount="3">
CUMZ 12161 (
<specimenCount id="6855AE5E8E1DFF908B597AF570C29AD8" type="generic">3 specimens</specimenCount>
in ethanol) from
<location id="7060309BA45A84423763F333A96E6565" LSID="urn:lsid:plazi:treatment:0C7D2D2174BE5698B91539251E676276:7060309BA45A84423763F333A96E6565" country="India" county="Satun Province" municipality="Khuan Don District" name="Ton Din Cave">Ton Din Cave</location>
,
<collectingMunicipality id="2405FEBCE07E76189463AC9066CC80D9">Khuan Don District</collectingMunicipality>
,
<collectingCounty id="F507B60A2D7380452A7CCF535003F3C8">Satun Province</collectingCounty>
,
<collectingDate id="B7BBA90C3601AEC37988CB5893387778" value="2017-07-07">7 July 2017</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="16A1A362C1079319D6E986E7455751CD" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="02D56F450E2D88E9C4C0303157DAD143" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="D13C60B5558C7579EF2CB63AE44A1A5E" pageId="0" pageNumber="1">Shell rounded, spindle-shaped, translucent whitish to brown. Whorls 7, convex; third to penultimate whorls broader; last whorl narrower, detached, brought forward, with a filiform basal keel broader at the mouth. Aperture almost circular, always with basal indentation; palatal indentation obvious in specimens with thicker shell. Operculum thick cylindrical, corneous, multi-layer, spring-like when extended by force; inner operculum (attached to dorsal side of posterior body) translucent yellow, convex with crater within and conical protrusion in the middle; outer operculum (free surface) dark brown and usually eroded.</paragraph>
</subSubSection>
<subSubSection id="9A4240DF48A09E402A3C88DE425097F3" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="443B055C4AE2BE645998456C08CEFE2F" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="3624DFE5E05D4B174F9C4ED5C8472109" pageId="0" pageNumber="1">
<taxonomicName id="25247170F8A7EA79F39DC847B9FAA0A0" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
<emphasis id="AF20FB90CA21E21204323144B8C7A6BE" italics="true" pageId="0" pageNumber="1">Tortulosa tortuosa</emphasis>
</taxonomicName>
can be distinguished from other species in this genus by a narrower last whorl that is detached from the penultimate whorl and brought forward, a shallower basal indentation, and the presence of a palatal indentation (
<bibRefCitation id="79CFDCE07D56C9CF758CD37669EA2A5E" author="Raheem, DC" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B136" refString="Raheem, DC, Taylor, H, Ablett, J, Preece, RC, Aravind, NA, Naggs, F, 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1-294." title="A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1 - 294." year="2014">Raheem et al. 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D9B98BFF5A21EDBB97249E8FDE0FE8FE" pageId="0" pageNumber="1" type="distribution">
<paragraph id="49C752B860B40D1D02C7241A617621A0" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="92741513351748D526D87308B81B9A5A" pageId="0" pageNumber="1">
Northern Peninsular Malaysia and southern Thailand. The type locality of this species is still controversial while the occurrences in India and Nicobar Islands need further confirmation (
<bibRefCitation id="318B961787AA2B10698B814CB586D58B" author="Sutcharit, C" journalOrPublisher="BRT, Bangkok" pageId="0" pageNumber="1" refId="B153" refString="Sutcharit, C, Panha, S, 2008. Land snails in Khao Nan National Park. BRT, Bangkok" title="Land snails in Khao Nan National Park." year="2008">Sutcharit and Panha 2008</bibRefCitation>
;
<bibRefCitation id="11785C100E7B82B6B082D2973437B18A" author="Raheem, DC" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B136" refString="Raheem, DC, Taylor, H, Ablett, J, Preece, RC, Aravind, NA, Naggs, F, 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1-294." title="A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1 - 294." year="2014">Raheem et al. 2014</bibRefCitation>
,
<bibRefCitation id="3A5D0CFD332794DB61712C834D7A7962" DOI="https://doi.org/10.1080/14772019.2017.1388298" author="Raheem, DC" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="1301 - 1317" refId="B138" refString="Raheem, DC, Schneider, S, Boehme, M, Vasiliyan, D, Prieto, J, 2018. The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia. Journal of Systematic Palaeontology 16 (15): 1301 - 1317, DOI: https://doi.org/10.1080/14772019.2017.1388298" title="The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia." url="https://doi.org/10.1080/14772019.2017.1388298" volume="16" year="2018">2018</bibRefCitation>
;
<bibRefCitation id="FBD3EFB54DCED2E5107219AD68CE7F86" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="F6DBF0AF4BA34F636A50849E49137898" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="339F9A520CE79A7749D3DD9CB54CD59E" pageId="0" pageNumber="1" type="remarks">
<paragraph id="E2C87AF218FC6255068A2383C2AA36CD" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="37DAD3F21A991DD4639D222E3AD4C410" pageId="0" pageNumber="1">
The name
<taxonomicName id="C7DC17410735B22C6E787FA710713660" class="Gastropoda" family="Pupinidae" genus="Turbo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turbo tortuous" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuous">
<emphasis id="923AF91207F88E84261827B23044C49C" italics="true" pageId="0" pageNumber="1">Turbo tortuous</emphasis>
</taxonomicName>
Chemnitz, 1795 was published prior to
<normalizedToken id="37B671921BF3251A255D32195F611274" originalValue="Férussac’s">Ferussac's</normalizedToken>
name, but it is unavailable (
<bibRefCitation id="2326FC3A92FF561BE10838CAB1CFFADA" DOI="https://doi.org/10.1080/14772019.2017.1388298" author="Raheem, DC" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="1301 - 1317" refId="B138" refString="Raheem, DC, Schneider, S, Boehme, M, Vasiliyan, D, Prieto, J, 2018. The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia. Journal of Systematic Palaeontology 16 (15): 1301 - 1317, DOI: https://doi.org/10.1080/14772019.2017.1388298" title="The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia." url="https://doi.org/10.1080/14772019.2017.1388298" volume="16" year="2018">Raheem et al. 2018</bibRefCitation>
). See
<bibRefCitation id="F3102F255F1E4A1D5E109A254A591C6A" author="Raheem, DC" journalOrPublisher="Journal and Proceedings of the Asiatic Society of Bengal, new series" pageId="0" pageNumber="1" refId="B136" refString="Raheem, DC, Taylor, H, Ablett, J, Preece, RC, Aravind, NA, Naggs, F, 2014. A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1-294." title="A systematic revision of the land snails of the Western Ghats of India. Tropical Natural History (Supplement 4): 1 - 294." year="2014">Raheem et al. (2014</bibRefCitation>
,
<bibRefCitation id="51C92F9061962665CAAE68D9CE8D54B1" DOI="https://doi.org/10.1080/14772019.2017.1388298" author="Raheem, DC" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="1301 - 1317" refId="B138" refString="Raheem, DC, Schneider, S, Boehme, M, Vasiliyan, D, Prieto, J, 2018. The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia. Journal of Systematic Palaeontology 16 (15): 1301 - 1317, DOI: https://doi.org/10.1080/14772019.2017.1388298" title="The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia." url="https://doi.org/10.1080/14772019.2017.1388298" volume="16" year="2018">2018</bibRefCitation>
) and
<bibRefCitation id="FC607C7889E1028BB4F446DA32910AE5" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="A6D20D056DC4F0B39D8678DE67A63825" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. (2020)
</bibRefCitation>
for the notes on taxonomy and type specimen of this species. Currently, this is the only extant species in the
<taxonomicName id="D53418E37A41D4F273E2D89BF3F90781" authority="Tortulosa" class="Gastropoda" family="Pupinidae" genus="Turbo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Tortulosa) subsp. subgenus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Tortulosa">subgenus Tortulosa</taxonomicName>
. One extinct species,
<taxonomicName id="175EBB2207D913353385038CE5EB2DF7" lsidName="T. naggsi" pageId="0" pageNumber="1" rank="species" species="naggsi">
<emphasis id="13140E7D2C29FCC7D91912AF0B98C1BF" italics="true" pageId="0" pageNumber="1">T. naggsi</emphasis>
</taxonomicName>
Raheem &amp; Schneider, 2017 in
<bibRefCitation id="2EF525FF1C6BE9C0BEE937933494D9AD" DOI="https://doi.org/10.1080/14772019.2017.1388298" author="Raheem, DC" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="1301 - 1317" refId="B138" refString="Raheem, DC, Schneider, S, Boehme, M, Vasiliyan, D, Prieto, J, 2018. The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia. Journal of Systematic Palaeontology 16 (15): 1301 - 1317, DOI: https://doi.org/10.1080/14772019.2017.1388298" title="The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia." url="https://doi.org/10.1080/14772019.2017.1388298" volume="16" year="2018">Raheem et al. (2018)</bibRefCitation>
was discovered from Son La Province, Northern Vietnam. This species exhibits a terminal part of the body whorl that is fully attached to the penultimate whorl, and thus corresponds more to the
<taxonomicName id="B5325414D46B2AF374E6789CE1D9D2E6" authority="Eucataulus" lsidName="(Eucataulus) subsp. subgenus" pageId="0" pageNumber="1" rank="subGenus" subGenus="Eucataulus">subgenus Eucataulus</taxonomicName>
from South Asia (
<bibRefCitation id="66FBA0E4918E7595EED00E9088BB79D8" DOI="https://doi.org/10.1080/14772019.2017.1388298" author="Raheem, DC" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" pagination="1301 - 1317" refId="B138" refString="Raheem, DC, Schneider, S, Boehme, M, Vasiliyan, D, Prieto, J, 2018. The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia. Journal of Systematic Palaeontology 16 (15): 1301 - 1317, DOI: https://doi.org/10.1080/14772019.2017.1388298" title="The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia." url="https://doi.org/10.1080/14772019.2017.1388298" volume="16" year="2018">Raheem et al. 2018</bibRefCitation>
).
</paragraph>
<paragraph id="9010539903E9A6225FE85C2EAC6D7865" pageId="0" pageNumber="1">
<bibRefCitation id="6AF53917AF763F78D4139116FD0F236B" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen (2001)</bibRefCitation>
treated
<taxonomicName id="A01F0EC0DDE966B31C1A5A5B904C5E55" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tweediei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">
<emphasis id="D1A56E5C7563E24CC02313E959A553C3" italics="true" pageId="0" pageNumber="1">P. tweediei</emphasis>
</taxonomicName>
, and
<bibRefCitation id="F0B4BF8F3F3B724FC2D1B8A1D29FBC7C" DOI="https://doi.org/10.12657/folmal.028.002" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="1" pagination="35 - 76" refId="B115" refString="Pall-Gergely, B, Hunyadi, A, Auffenberg, K, 2020. Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014-2019). Folia Malacologica 28 (1): 35 - 76, DOI: https://doi.org/10.12657/folmal.028.002" title="Taxonomic vandalism in malacology: Comments on molluscan taxa recently described by N. N. Thach and colleagues (2014 - 2019)." url="https://doi.org/10.12657/folmal.028.002" volume="28" year="2020">
<normalizedToken id="84CD344EA7F9B7FABDCE3DA97C16EA47" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. (2020)
</bibRefCitation>
treated both
<taxonomicName id="C9C8E0B7FBD2F137953403593BF2D838" lsidName="T. huberi" pageId="0" pageNumber="1" rank="species" species="huberi">
<emphasis id="0B59F046DECE179A4887CBE81EBDEC01" italics="true" pageId="0" pageNumber="1">T. huberi</emphasis>
</taxonomicName>
and
<taxonomicName id="E1759D085DD6909A1DA5321F6F3524F4" lsidName="T. schileykoi" pageId="0" pageNumber="1" rank="species" species="schileykoi">
<emphasis id="289497B8F916F99FBCE737FA4DE7DD6C" italics="true" pageId="0" pageNumber="1">T. schileykoi</emphasis>
</taxonomicName>
as junior subjective synonyms of
<taxonomicName id="9D45D7C08A8E8D6F24B92267B79B1E4F" lsidName="T. tortuosa" pageId="0" pageNumber="1" rank="species" species="tortuosa">
<emphasis id="CEB02A5A22A8A0AA56B6FDB9562908B3" italics="true" pageId="0" pageNumber="1">T. tortuosa</emphasis>
</taxonomicName>
. We agree on those synonymisations because the specimens we collect from the same locality exhibit a high infraspecific variation in the length of the detached part of the body whorl relative to the shell height, also in shell shape from ovate to elongate, and shell colour from translucent whitish to brown (Fig.
<figureCitation id="B87642E77903D7FDAAB11E3FC399BE66" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Infraspecific variation of shell shape and colour found in the same collecting locality of Tortulosa tortuosa, CUMZ 12166 from Tham Suea Temple, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure16" httpUri="https://binary.pensoft.net/fig/735377" pageId="0" pageNumber="1">16</figureCitation>
). All the specimens in Fig.
<figureCitation id="B5087EE3955C3E48CACE90E5FC4B6A68" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Infraspecific variation of shell shape and colour found in the same collecting locality of Tortulosa tortuosa, CUMZ 12166 from Tham Suea Temple, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure16" httpUri="https://binary.pensoft.net/fig/735377" pageId="0" pageNumber="1">16</figureCitation>
were found together with hundreds of other specimens inside the same decaying log at Tham Suea Temple, Krabi Province, southern Thailand. The distribution of this species in Thailand is provided in Fig.
<figureCitation id="92ECD9395832F31CC8B801BA124D397D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Map of southern Thailand showing the distribution of Coptocheilus sectilabris (filled square), Coptocheilus sumatranus (open square), Rhaphaulus lorraini (filled star), Rhaphaulus ascendens (open star), Rhaphaulus perakensis (asterisk), and Tortulosa tortuosa (circle). Each red symbol indicates the type locality of its respective taxon. Red circles indicate the type localities of Tortulosa huberi (1) and Tortulosa schileykoi (2)." figureDoi="10.3897/zookeys.1119.85400.figure4" httpUri="https://binary.pensoft.net/fig/735365" pageId="0" pageNumber="1">4</figureCitation>
.
</paragraph>
<caption id="1693B4FDE4C93FF57C758530882B4169" doi="10.3897/zookeys.1119.85400.figure4" httpUri="https://binary.pensoft.net/fig/735365" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="E51157EA0F6BAAF3F03C3DF2668805AB" pageId="0" pageNumber="1">
<emphasis id="D6BBAE63BAAE7AAC0D4F15A1CC5B6291" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Map of southern Thailand showing the distribution of
<taxonomicName id="B33D2B36F9E4FFAB6154E594617A82B8" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sectilabris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabris">
<emphasis id="0AD26AA7F9B08ADA4F61264D71BA16C7" italics="true" pageId="0" pageNumber="1">Coptocheilus sectilabris</emphasis>
</taxonomicName>
(filled square),
<taxonomicName id="C9408EAC105AD927A1BA5FDE5CCAE244" authorityName="Dohrn" authorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranus">
<emphasis id="7943A0FDDCAC2214226D064EE0502C59" italics="true" pageId="0" pageNumber="1">Coptocheilus sumatranus</emphasis>
</taxonomicName>
(open square),
<taxonomicName id="33F28540BB5FC8E60001EE4F8D675E09" authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorraini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorraini">
<emphasis id="7C1D28BB656AE628DAB831BFE7769E77" italics="true" pageId="0" pageNumber="1">Rhaphaulus lorraini</emphasis>
</taxonomicName>
(filled star),
<taxonomicName id="52FF56A53CBA523E442302777F881B4F" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus ascendens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ascendens">
<emphasis id="25708F3AFE161092F7F2FB05BA64C42F" italics="true" pageId="0" pageNumber="1">Rhaphaulus ascendens</emphasis>
</taxonomicName>
(open star),
<taxonomicName id="C6A3E37674AA66E07EDE634BA68FAD8A" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="13F64A97E4CD9D34C6C3CBFE9ACCBFBD" italics="true" pageId="0" pageNumber="1">Rhaphaulus perakensis</emphasis>
</taxonomicName>
(asterisk), and
<taxonomicName id="C265FA6B14314C5C8E0A637629F491FC" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
<emphasis id="508BA2A9C6A249935BDD6C12DA87EFEB" italics="true" pageId="0" pageNumber="1">Tortulosa tortuosa</emphasis>
</taxonomicName>
(circle). Each red symbol indicates the type locality of its respective taxon. Red circles indicate the type localities of
<taxonomicName id="44D22E90BC626AA0D5F5A8AA6F36245D" authorityName="Thach" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa huberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="70ABFAF4A7B15D1D241BB744FA3DBA75" italics="true" pageId="0" pageNumber="1">Tortulosa huberi</emphasis>
</taxonomicName>
(1) and
<taxonomicName id="B432AE6E8AA8837CAFBD03F4E0BA8C73" authorityName="Thach &amp; F. Huber" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa schileykoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="schileykoi">
<emphasis id="7DE8FD989821BA6589964645650D2F6A" italics="true" pageId="0" pageNumber="1">Tortulosa schileykoi</emphasis>
</taxonomicName>
(2).
</paragraph>
</caption>
<caption id="60499C459210D3240CA7DB0D71BF24BF" doi="10.3897/zookeys.1119.85400.figure5" httpUri="https://binary.pensoft.net/fig/735366" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="22672744B630B96E2DC78783493D058E" pageId="0" pageNumber="1">
<emphasis id="35460CE955D189D38A73FEA8ECD61584" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<emphasis id="B201902364EF28ECD283BEE97068FBBC" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName id="690A673AC19399B1CE7FA2AB5F40DA17" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sectilabris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabris">
<emphasis id="9A593428FD414AE33897C9410DDC6FBC" italics="true" pageId="0" pageNumber="1">Coptocheilus sectilabris</emphasis>
</taxonomicName>
:
<emphasis id="EA73EDE861E0E4623C81B4A4E5744B49" bold="true" pageId="0" pageNumber="1">A</emphasis>
lectotype MCZ 169361 from Tavoy
<emphasis id="8A56A588B7608DA254EDE9D6ADE6044A" bold="true" pageId="0" pageNumber="1">B</emphasis>
specimen SMF 109813 from Tavoy, and
<emphasis id="45E58BEC4929C50B9CD870414B7D6729" bold="true" pageId="0" pageNumber="1">C</emphasis>
specimen CUMZ OLM-0111 from Kaeng Krachan, Phetchaburi
<emphasis id="A5A557B1AC81841644BA3632D6E2E853" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName id="4CB5EFB70129B761BFD17635E4E937C5" authorityName="Dohrn" authorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranus">
<emphasis id="DB6CDA8EBAD2B5B62FA9B9C7FDAE2765" italics="true" pageId="0" pageNumber="1">Coptocheilus sumatranus</emphasis>
</taxonomicName>
:
<emphasis id="3A734C79F719A721F9F4ECCACBFEA57B" bold="true" pageId="0" pageNumber="1">D</emphasis>
syntype of
<taxonomicName id="FFD444A5604AC9DC8B00551BBEFD448E" authorityName="Fulton" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="1B0E3CC8496835C3669CF16586039030" italics="true" pageId="0" pageNumber="1">Coptocheilus perakensis</emphasis>
</taxonomicName>
NHMUK 1903.11.20.33 from Perak
<emphasis id="BD623F6BF1AD6B8A04ABA74AC254F414" bold="true" pageId="0" pageNumber="1">E</emphasis>
specimen SMF 262529/1 &quot;
<taxonomicName id="BD4DFFF5E5FD46566E1A0212ACD8F738" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis id="0ECB601F3E10A88EC29AE6D0D0C2CAAD" italics="true" pageId="0" pageNumber="1">Schistoloma siamensis</emphasis>
</taxonomicName>
Brandt&quot; from Thailand: an den
<normalizedToken id="40DD37E756C30486F9B985B1562D2504" originalValue="Tanto-Fällen">Tanto-Faellen</normalizedToken>
bei Ban Nong Star; Yala Provinz, and
<emphasis id="339D8238D5BBA0B49B56F4490AA4E53A" bold="true" pageId="0" pageNumber="1">F</emphasis>
specimen NHMUK 1986.4.19.14 &quot;
<taxonomicName id="08E7DD999C9EF48991417D5DBCE0EF7C" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">
<emphasis id="229704F7EB56AE9497EF470A6501B139" italics="true" pageId="0" pageNumber="1">Coptocheilus sectilabrum</emphasis>
</taxonomicName>
var.&quot; from Larut near Perak.
</paragraph>
</caption>
<caption id="663CF946DCB89CCAAFE9F347693755E9" doi="10.3897/zookeys.1119.85400.figure6" httpUri="https://binary.pensoft.net/fig/735367" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph id="7577E178592825E7CE64284027C2B0CA" pageId="0" pageNumber="1">
<emphasis id="FE2543E74878676A46401AE1D00B1155" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<emphasis id="9279E10CCF94517C4A854E25BF0151D1" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName id="2560E34E885476228332B7A3B5131A6B" authorityName="Kongim &amp; Panha" authorityYear="2013" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. monochroma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="monochroma">
<emphasis id="AB731863AD80FA1FAC022CC1B48A0D0D" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti monochroma</emphasis>
</taxonomicName>
:
<emphasis id="90613EF766C98EECDDB7913A669DABE7" bold="true" pageId="0" pageNumber="1">A</emphasis>
juvenile specimen CUMZ 12182 from Tham Suea Lueang Temple, Loei and
<emphasis id="63C660509BEFD2D5D65B02D3CAEE1D3E" bold="true" pageId="0" pageNumber="1">B</emphasis>
paratype CUMZ 1562 from Tam Pha Bing Temple, Loei
<emphasis id="3D91CD0DA0BB2BDE8EBB3E944F29F05D" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
<taxonomicName id="451B877C3DF09DC4B28B525CD070EA2F" authority="mouhoti" authorityName="mouhoti (Pfeiffer" authorityYear="1863" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="mouhoti">
<emphasis id="F95354F8870ACBF787A0B12018430455" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti mouhoti</emphasis>
</taxonomicName>
<emphasis id="9DB627FF703447C94B563DEE06424E60" bold="true" pageId="0" pageNumber="1">C</emphasis>
lectotype of
<taxonomicName id="95363FEA4A264F805AE6DF3F65F25088" authorityName="L. Pfeiffer" authorityYear="1863" class="Gastropoda" family="Pupinidae" genus="Hybocystis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hybocystis mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="6514904C1B79302D2AAAE4150E38CB50" italics="true" pageId="0" pageNumber="1">Hybocystis mouhoti</emphasis>
</taxonomicName>
NHMUK 20130071/1 from Lao Mountains, Camboja
<emphasis id="17A2941E69095D4DD65A1A0532F095E7" bold="true" pageId="0" pageNumber="1">D</emphasis>
holotype of '
<taxonomicName id="2D79E055CF0746D0BC8501C3C57EFFD4" authorityName="Thach" authorityYear="2021" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria nicoarlingi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nicoarlingi">
<emphasis id="375097F707583D05DD7B1C345D2036B6" italics="true" pageId="0" pageNumber="1">Pollicaria nicoarlingi</emphasis>
</taxonomicName>
' MNHN-IM-2000-37277, and
<emphasis id="482BBF573C18744961F529DB7B3737E1" bold="true" pageId="0" pageNumber="1">E</emphasis>
specimen CUMZ 12166 from Wang Daeng Cave, Phitsanulok
<emphasis id="DBE407CCB53042007D307E98C56A3EDD" bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName id="ED67778BF2753AC0F07B89A3FAF61C32" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria myersii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="myersii">
<emphasis id="2A9CEA3D912215345CC8DB9E03602122" italics="true" pageId="0" pageNumber="1">Pollicaria myersii</emphasis>
</taxonomicName>
, holotype of '
<taxonomicName id="E7B6286855184F83C0823997789E6AE2" authorityName="Thach" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria huberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="AEAB563AB6A8D464A701633BCFADCF8E" italics="true" pageId="0" pageNumber="1">Pollicaria huberi</emphasis>
</taxonomicName>
' NHMUK 20180253. Photo: F. Prugnaud, MNHN (
<emphasis id="C85C0C84F92EF30D5BA0B41E3C60D863" bold="true" pageId="0" pageNumber="1">D</emphasis>
).
</paragraph>
</caption>
<caption id="5FCFBF0A449803D0EAE497DC13F09174" doi="10.3897/zookeys.1119.85400.figure7" httpUri="https://binary.pensoft.net/fig/735368" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph id="7D93F61D04EA00F3EAE765CDCE4B3AF1" pageId="0" pageNumber="1">
<emphasis id="C80CD1AD4274F5A42220CA6E4AE0B8E9" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Map of northern Thailand showing the distribution of
<taxonomicName id="547502F442D4F117B4FC75DDCF8E59DB" authority="mouhoti" authorityName="mouhoti (Pfeiffer" authorityYear="1863" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="mouhoti">
<emphasis id="E5924684355368CBD3E2B7348DC41497" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti mouhoti</emphasis>
</taxonomicName>
(filled circle),
<taxonomicName id="43A082A63F0B66CD77A20CC701A975EE" authorityName="Kongim &amp; Panha" authorityYear="2013" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. monochroma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="monochroma">
<emphasis id="9218AEA02BDC93FF71E9D51FE92C33D5" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti monochroma</emphasis>
</taxonomicName>
(open circle),
<taxonomicName id="698954741E6AA4B49FC7E4188CDA2195" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias caligosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="caligosus">
<emphasis id="3119D18B87B371601A2D3611A1E3FA00" italics="true" pageId="0" pageNumber="1">Pseudopomatias caligosus</emphasis>
</taxonomicName>
(square),
<taxonomicName id="92AA1FC549D09E52A6AA62DECB0C7911" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias doiangkhangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="doiangkhangensis">
<emphasis id="EFAD39A6AD12B2174802A5D262201FB5" italics="true" pageId="0" pageNumber="1">Pseudopomatias doiangkhangensis</emphasis>
</taxonomicName>
sp. nov. (hexagon),
<taxonomicName id="975576136459E500B057F55BB5F5E037" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias pallgergelyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallgergelyi">
<emphasis id="5B1F6F591C650F51BC0298D7CB9E97FF" italics="true" pageId="0" pageNumber="1">Pseudopomatias pallgergelyi</emphasis>
</taxonomicName>
sp. nov. (triangle),
<taxonomicName id="B290EFC8ECC312B1FD6D2F196562DD0F" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="545ADE2A4F4E8CC6A32F58B1E2B59268" italics="true" pageId="0" pageNumber="1">Pupinella mansuyi</emphasis>
</taxonomicName>
(cross), and
<taxonomicName id="29073A2A4BE26F8B3CADA30520BD2837" authorityName="Pall-Gergely, Hunyadi &amp; Maassen" authorityYear="2014" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">
<emphasis id="78622D3B9C5B1249893D554B3609EDEF" italics="true" pageId="0" pageNumber="1">Rhaphaulus tonkinensis</emphasis>
</taxonomicName>
(star). Each red symbol indicates the type locality of its respective taxon. The red filled circle denotes the type locality of
<taxonomicName id="CA45390E196CB7EDA5DD2E44E6749105" authorityName="Thach" authorityYear="2021" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria nicoarlingi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nicoarlingi">
<emphasis id="DC59E4B51491BB8C9A386F7ACAAC82EA" italics="true" pageId="0" pageNumber="1">Pollicaria nicoarlingi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="96F0E9888DC40B6D899A1404F0619AA9" doi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph id="4E71B43FEDF630E1B914F491655105EB" pageId="0" pageNumber="1">
<emphasis id="462306F2A2F1700F81D52C2D448B82FC" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Live specimens of
<emphasis id="40BFBE80EAC264795D462DC8443ACA30" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName id="0BFE85A1A0C184DAD6386A4EB53A5252" authority="mouhoti" authorityName="mouhoti (Pfeiffer" authorityYear="1863" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. mouhoti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="mouhoti">
<emphasis id="9BEADA6EFB1A88FBEB4ECAF0B06D039E" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti mouhoti</emphasis>
</taxonomicName>
: specimens
<emphasis id="92F569075579FB42119857D7E6F7DE4B" bold="true" pageId="0" pageNumber="1">A</emphasis>
CUMZ 12166 and
<emphasis id="E0A17D4DC45B5EFAE024694BBFEC8B66" bold="true" pageId="0" pageNumber="1">B</emphasis>
CUMZ 12175 from Wang Daeng Cave, Phitsanulok
<emphasis id="0FD820B136A44A8F54CA4728753CE641" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="972336BCB30D6BEF4806A2D4EE3C4621" authorityName="Kongim &amp; Panha" authorityYear="2013" class="Gastropoda" family="Pupinidae" genus="Pollicaria" kingdom="Animalia" lsidName="Pollicaria mouhoti subsp. monochroma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="mouhoti" subSpecies="monochroma">
<emphasis id="C019436331D86DC4250064BB6EC270AB" italics="true" pageId="0" pageNumber="1">Pollicaria mouhoti monochroma</emphasis>
</taxonomicName>
, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei
<emphasis id="BFD3BFCEEA708D6D700D10CC62C4F3C4" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="810B0324A20E132CAF4C28BB42D01998" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
<emphasis id="BF32B2BC945AE015C9E95948231BB224" italics="true" pageId="0" pageNumber="1">Tortulosa tortuosa</emphasis>
</taxonomicName>
, specimen CUMZ 12155 from Tham Suea Temple, Krabi
<emphasis id="81048C5C667B6565EF2D4F84A85F38D0" bold="true" pageId="0" pageNumber="1">E-H</emphasis>
<taxonomicName id="52354CA25CC653DFE93B3902B415F735" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
<emphasis id="E3CA2E7A551EB68A7EC766FAA1D2F1BD" italics="true" pageId="0" pageNumber="1">Pupina artata</emphasis>
</taxonomicName>
: specimens of
<emphasis id="ED502E2990E8E3C45BA45D3EDE12093B" bold="true" pageId="0" pageNumber="1">E</emphasis>
CUMZ 12006 from Pha Daeng Cave, Mae Hong Son
<emphasis id="C2054720C69DF83B31E68401B5C33AB1" bold="true" pageId="0" pageNumber="1">F</emphasis>
CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and
<emphasis id="F4DDA4CA0DEB9E33EA0791C78C3F5A67" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (
<emphasis id="8EAA68BF30489DDC0B74F760D9A16A85" bold="true" pageId="0" pageNumber="1">G</emphasis>
) and grey (
<emphasis id="E6A8E2858308AA65CADE9DAC840F4536" bold="true" pageId="0" pageNumber="1">H</emphasis>
) shell morphs; All not to scale.
</paragraph>
</caption>
<caption id="F6B0A0D1DD302EB705F806EE1F6311F2" doi="10.3897/zookeys.1119.85400.figure9" httpUri="https://binary.pensoft.net/fig/735370" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph id="9F64F3CD28A4A6A38382FC6A029D9CD9" pageId="0" pageNumber="1">
<emphasis id="FFE04C7F98C2BAA64D5B77A1461EB0DA" bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
<emphasis id="4CC4A46D21DE6C23ACE78B4ED0198BAD" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName id="F98E704C67082AD242F8B0227D432266" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias caligosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="caligosus">
<emphasis id="DF1E0CB7212DE020F1628D9869358F03" italics="true" pageId="0" pageNumber="1">Pseudopomatias caligosus</emphasis>
</taxonomicName>
:
<emphasis id="4BD9CADEE7ED18F3EACB6A194E591A92" bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype HNHM 100176 and
<emphasis id="EB8F1D17C8EDA7BCDE05E30990D453CA" bold="true" pageId="0" pageNumber="1">B</emphasis>
specimen CUMZ 12191 from Pa Tham Wua Temple, Mae Hong Son
<emphasis id="74C0B21C8E101645E95651D973E151D2" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
<taxonomicName id="7E0B17A3C788744E788BA23DF54656DC" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias doiangkhangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="doiangkhangensis">
<emphasis id="FDE9F93967A6C6AC559FEFAA87374D28" italics="true" pageId="0" pageNumber="1">Pseudopomatias doiangkhangensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="22DF7B4DBFB1534381B44DC36249B528" bold="true" pageId="0" pageNumber="1">C</emphasis>
holotype CUMZ 12165/1 and
<emphasis id="58F55F3C241B891F4F956BA724C97806" bold="true" pageId="0" pageNumber="1">D</emphasis>
paratype CUMZ 5219 from Doi Ang Khang, Chiang Mai
<emphasis id="1588213DB0507764643686A5A3394241" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName id="CCE6E6B2E23B9896A31417702438B3E2" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopomatias pallgergelyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallgergelyi">
<emphasis id="D1B9B5DB963D78E5FBE35B29EFCF9B9B" italics="true" pageId="0" pageNumber="1">Pseudopomatias pallgergelyi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="8DE355E18B5DA4F698F1D85F56E39319" bold="true" pageId="0" pageNumber="1">E</emphasis>
holotype CUMZ 12167/1
<emphasis id="8657C9E5BCB9AAC765CE9FF4232E671D" bold="true" pageId="0" pageNumber="1">F</emphasis>
paratype CUMZ 12167/2 from Pha Daeng Cave, Mae Hong Son. Photo: B.
<normalizedToken id="45F398E05933A6271AEEF2271A9CC499" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
(
<emphasis id="9CC47393663A58F74C9FC7F134B25201" bold="true" pageId="0" pageNumber="1">A</emphasis>
).
</paragraph>
</caption>
<caption id="E3E500F413E842003855049C33CEAFB2" doi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph id="4840FDA7115D5AB3613C2BE4586C4F51" pageId="0" pageNumber="1">
<emphasis id="3ACA241A3820722FF883DA5E87A4044A" bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
Umbilical, columellar and parietal views of
<emphasis id="ECEBDAE48436ED3F86A3B2E21914E72C" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="E4261DE3DB52E6596E2C4A39C4401B8F" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="E1F5EFBB9F86E0E917FE9AC8DDDB47A3" italics="true" pageId="0" pageNumber="1">Pupinella mansuyi</emphasis>
</taxonomicName>
, specimen CUMZ 12148 from Pha Chu, Nan
<emphasis id="D3B38A86BD8DBBF18235DD347DF48061" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="04C57C0D2FA06F1D8A6595119F00C54C" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
<emphasis id="B21DD9B17CD308DB624DB156979AEE06" italics="true" pageId="0" pageNumber="1">Pupina artata</emphasis>
</taxonomicName>
from the
<taxonomicName id="82564482BDA8D5F0FCE03D62A4A0ECF8" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
<emphasis id="8CCB5F8AC1766720A66264E8639973F9" italics="true" pageId="0" pageNumber="1">Pupina artata</emphasis>
</taxonomicName>
species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai
<emphasis id="9C35F1B2AB1DB4CA210228D176005139" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="4C3B7D1ECF3A683527F60FE84B2C41C8" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni">
<emphasis id="EB6F22C7E80C169A59AAA34BF209648B" italics="true" pageId="0" pageNumber="1">Pupina godwinausteni</emphasis>
</taxonomicName>
sp. nov. from the
<taxonomicName id="0354855CBE4B3A375ED1486A021EE944" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
<emphasis id="1DDBFE8B5E83C01B808C2EAE67FA1537" italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
</taxonomicName>
species group, holotype CUMZ 12090/1
<emphasis id="BA044C5D52FE7BB76EFE1A286FFA1AA6" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="EE92A08D8030149A473E9A9192BACBDC" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="BE35C81B8391235AD61664ED270D4A98" italics="true" pageId="0" pageNumber="1">Pupina aureola</emphasis>
</taxonomicName>
from the
<taxonomicName id="7B723DFBF9FD7681AE484DECCD172DFF" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="F1C0E05B958AC16987350CBF8C98E8AF" italics="true" pageId="0" pageNumber="1">Pupina aureola</emphasis>
</taxonomicName>
species group, specimen CUMZ 12130 from Sra Morakot, Krabi.
</paragraph>
</caption>
<caption id="60ED20FAA5115EC1A25E9F570824558D" doi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph id="066278D52619EC3ACE61C1CB9BBF78FB" pageId="0" pageNumber="1">
<emphasis id="EC10BE6E4D3B38BF687757C5E0AE5C43" bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
Shells of
<taxonomicName id="1109AA6CC515E674792A742655111E57" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="4B95641F5747BB4315224AEF44DD5B1A" italics="true" pageId="0" pageNumber="1">Pupinella</emphasis>
</taxonomicName>
species from mainland Southeast Asia.
<emphasis id="872BDAAAE73CCE790B5B5B08B93BC40D" bold="true" pageId="0" pageNumber="1">A-G</emphasis>
<taxonomicName id="91217BA50FCB17512A1646A62ED1BCED" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="5C9E3D44B9B907C5D89952AE3E64BE4D" italics="true" pageId="0" pageNumber="1">Pupinella mansuyi</emphasis>
</taxonomicName>
:
<emphasis id="3C285258814E9B4B789701B50D979875" bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype of
<taxonomicName id="5FF0E5B4C57A6BE2AB4AB7ADBE579518" authorityName="Dautzenberg &amp; H. Fischer" authorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Eupupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupupina mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="B38C0112ED92CD5D913A04705023E714" italics="true" pageId="0" pageNumber="1">Eupupina mansuyi</emphasis>
</taxonomicName>
MNHN-IM-2000-30756 from Deux-Ponts
<emphasis id="9A9E9ECBD94C8A3BF9D5C78BA935E332" bold="true" pageId="0" pageNumber="1">B</emphasis>
syntype of
<taxonomicName id="2CE3E264C6FDE610A336F5A7288AF621" authorityName="Dautzenberg &amp; H. Fischer" authorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Eupupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupupina mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="A57B5547C1500DE8550F560C99A1DE14" italics="true" pageId="0" pageNumber="1">Eupupina mansuyi</emphasis>
</taxonomicName>
MNHN-IM-2000-36067 from Deux-Ponts
<emphasis id="F3771979F3265F9997B8E9B494E0C637" bold="true" pageId="0" pageNumber="1">C</emphasis>
syntype of
<taxonomicName id="EEE0FA291DAA220E3284C8A1D12DF244" authorityName="Dautzenberg &amp; H. Fischer" authorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Eupupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupupina mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="21BBC3E6E4E370A620FAD10B13F0E29D" italics="true" pageId="0" pageNumber="1">Eupupina mansuyi</emphasis>
</taxonomicName>
MNHN-IM-2000-36068 from Quang-Huyen
<emphasis id="71CEA685CEF0CC84E6ED54B82F57E689" bold="true" pageId="0" pageNumber="1">D</emphasis>
syntype of
<taxonomicName id="1816958AD6C92B4E2E09DA5453BF47D8" authorityName="Dautzenberg &amp; H. Fischer" authorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Eupupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupupina mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="43EE05FC83FA30D6B88C42120DA56EC8" italics="true" pageId="0" pageNumber="1">Eupupina mansuyi</emphasis>
</taxonomicName>
RBINS MT970/1 from Quang-Huyen
<emphasis id="1DB44FF7946AC918E9F24D30EF6515AD" bold="true" pageId="0" pageNumber="1">E</emphasis>
holotype of
<taxonomicName id="76676BDE4CA39CB04506A5C8C3334B3A" authorityName="Thach &amp; F. Huber" authorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella frednaggsi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="frednaggsi">
<emphasis id="8500DE0A3441650BA0555EAC4826A300" italics="true" pageId="0" pageNumber="1">Pupinella frednaggsi</emphasis>
</taxonomicName>
NHMUK 20170285
<emphasis id="F8F9D47A475DFD963DDD35B9A88A60FB" bold="true" pageId="0" pageNumber="1">F</emphasis>
specimen CUMZ 12148 from Pha Chu Mount, Nan, and
<emphasis id="5AE47BF7174FF05CFAA03C7CD60E4D5E" bold="true" pageId="0" pageNumber="1">G</emphasis>
specimen CUMZ 12149 from Pha Tub Cave, Nan
<emphasis id="E69F0FCE0F52CD9064738EB27E7EAEBE" bold="true" pageId="0" pageNumber="1">H</emphasis>
<taxonomicName id="567E5056E0ADD7FF2A7DB4CB6EC7F418" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis">
<emphasis id="ED6DBC37AD1CA6BB339F2C02B6FCB830" italics="true" pageId="0" pageNumber="1">Pupinella sonlaensis</emphasis>
</taxonomicName>
, paratype ZRC.MOL.9377
<emphasis id="AB73FCAB949D0ED705696A3FB344035C" bold="true" pageId="0" pageNumber="1">I-L</emphasis>
<taxonomicName id="CBA899E188EC47E4258DE5B24361E07F" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
<emphasis id="F94EDEBED31518BA50756A1332EC67E8" italics="true" pageId="0" pageNumber="1">Pupinella illustris</emphasis>
</taxonomicName>
<emphasis id="266C0BDFA558B96095FB4B372B612F1C" bold="true" pageId="0" pageNumber="1">I, J</emphasis>
syntypes of
<taxonomicName id="222F30199428C309E45A2F9B677BC723" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
<emphasis id="F696F4135471E7B19F31BCE0FBA1A77D" italics="true" pageId="0" pageNumber="1">Pupina illustris</emphasis>
</taxonomicName>
MNHN-IM-2000-35842 from Tonkin
<emphasis id="A64326079616354E351ED39D4D16A06A" bold="true" pageId="0" pageNumber="1">K</emphasis>
lectotype of
<taxonomicName id="3738125B32D0475A46609AD8EE28595C" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="6E5D8F97E05693B18BF6B904525677C3" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
</taxonomicName>
MNHN-IM-2000-35838 from Lang-Son et
<normalizedToken id="D73D2605DBF563DC4253518121198E56" originalValue="That-Khé">That-Khe</normalizedToken>
, and
<emphasis id="FB4E577AB81E302B8B54A63B4910B7DC" bold="true" pageId="0" pageNumber="1">L</emphasis>
paralectotype of
<taxonomicName id="03F759ED254C33009766CBD8E5609E56" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="8E820A02C354FD76DEC7D7021AFBE6A4" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
</taxonomicName>
SMF 109932/10 from Tonkin:
<normalizedToken id="189E328BA72EE1CA21DFD22F16C7FC6C" originalValue="That-khé">That-khe</normalizedToken>
. Photo: A. Lardeur, P. Maestrati, MNHN (
<emphasis id="87FE2C4C16FEEDF3A355D2BD4CA0BC20" bold="true" pageId="0" pageNumber="1">A-C, I-K</emphasis>
), F. Trus, RBINS (
<emphasis id="AC9BABDBE5E8E5D5F4F31FB68BE8DA5F" bold="true" pageId="0" pageNumber="1">D</emphasis>
), S.K. Tan, ZRC (
<emphasis id="5C4430001B91C5E7051364AC5263B9FD" bold="true" pageId="0" pageNumber="1">H</emphasis>
).
</paragraph>
</caption>
<caption id="E1E36BA712DB896AA6566B533F69D44A" doi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph id="D81BF5657AA5D29A84293F01EE8E1D02" pageId="0" pageNumber="1">
<emphasis id="90915C1A861FFF5546DE3F4E31CBAE04" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<emphasis id="C4E305B46DEFF544FE1A5271C1DCFAE5" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName id="F58DA9493DF992DEF0CFE92B8CBEEEB6" baseAuthorityName="Dautzenberg &amp; Fischer" baseAuthorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="754BCDBDCBAF166B5AC7FE900E1E1C26" italics="true" pageId="0" pageNumber="1">Pupinella mansuyi</emphasis>
</taxonomicName>
:
<emphasis id="A9E151FB13FC55474AAA1314177946B2" bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype of
<taxonomicName id="9FA8889DF93C7F68B5D89C516704C457" authorityName="Dautzenberg &amp; H. Fischer" authorityYear="1908" class="Gastropoda" family="Pupinidae" genus="Eupupina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupupina mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mansuyi">
<emphasis id="0D3951662EAF243D39D6ED2563041AB6" italics="true" pageId="0" pageNumber="1">Eupupina mansuyi</emphasis>
</taxonomicName>
RBINS MT970/1 from Quang-Huyen
<emphasis id="1EBDF85F64EA1BE252EE6EE57A883AFD" bold="true" pageId="0" pageNumber="1">B</emphasis>
specimen CUMZ 12148 from Pha Chu Mount, Nan, and
<emphasis id="FA1BB7D2223A72E7857E853AD0F05BC8" bold="true" pageId="0" pageNumber="1">C</emphasis>
specimen CUMZ 12149 from Pha Tub Cave, Nan.
<emphasis id="D8FCDC82425110B4F4C1498717A78E7F" bold="true" pageId="0" pageNumber="1">D, E</emphasis>
<taxonomicName id="6B4578F411747318C95D4C24C769023D" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
<emphasis id="B98D4ACFD0613A5086DD4C199007F8D7" italics="true" pageId="0" pageNumber="1">Pupinella illustris</emphasis>
</taxonomicName>
<emphasis id="C397D5FE671255FF093DDF67FDB09180" bold="true" pageId="0" pageNumber="1">D</emphasis>
syntype of
<taxonomicName id="D7EF54EDB5E964BEB01FB71325604DD4" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
<emphasis id="840913774A40FD9B3C8BE90B687FE8CF" italics="true" pageId="0" pageNumber="1">Pupina illustris</emphasis>
</taxonomicName>
MNHN-IM-2000-35842 from Tonkin and
<emphasis id="340C0DFF2BB5121BE9586179534D9B4E" bold="true" pageId="0" pageNumber="1">E</emphasis>
paralectotype of
<taxonomicName id="C2C0DBD5CF490F96D4C0B6A0D88272D9" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="D6CFB8948BA67318DAD944B25FF3BD44" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
</taxonomicName>
SMF 109932/10 from Tonkin:
<normalizedToken id="A9687999111150D1DCE9858A99F3CCB8" originalValue="That-khé">That-khe</normalizedToken>
<emphasis id="8E4B67DD218B1AF825EF835737E55679" bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName id="A7D1C846AEC6795A50DF13573FE7F2D9" authorityName="Jirapatrasilp &amp; Sutcharit &amp; Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis">
<emphasis id="81A3F939EE70ABDB30E442D3844EFD6B" italics="true" pageId="0" pageNumber="1">Pupinella sonlaensis</emphasis>
</taxonomicName>
, paratype ZRC.MOL.9377. Photo: F. Trus, RBINS (
<emphasis id="2BC8C5E00FF51F87100965A9C1C23D81" bold="true" pageId="0" pageNumber="1">A</emphasis>
), P. Maestrati, MNHN (
<emphasis id="1542FD5C92280797ACC5F51EE2196C93" bold="true" pageId="0" pageNumber="1">D</emphasis>
), S.K. Tan, ZRC (
<emphasis id="589D9120E7C2E94C1EC4B287BEE197DE" bold="true" pageId="0" pageNumber="1">F</emphasis>
).
</paragraph>
</caption>
<caption id="F0377305D1A5F98D0BCF32EC3F26DBD6" doi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph id="75AE4031CCCD97FB2F3A8D8D71332752" pageId="0" pageNumber="1">
<emphasis id="0791D398C87F85BF98ECEB70BE66E3EC" bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<emphasis id="E3C15AB86DDEC56305A280CF0576E5CB" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName id="ED65CCECCA6E0D7DA6427E73D1E248A7" authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorraini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorraini">
<emphasis id="3EABCBDF523CFB20198E42B1CC81FE9A" italics="true" pageId="0" pageNumber="1">Rhaphaulus lorraini</emphasis>
</taxonomicName>
:
<emphasis id="6D62FA596E0DD24F32AFC8257C64E1B9" bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype NHMUK 20130454 from Pulo Penang and
<emphasis id="366E908CF3248D8528BA9AF361ED33CD" bold="true" pageId="0" pageNumber="1">B</emphasis>
specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga
<emphasis id="E6062501973589CA71A8A0B565636863" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="1D8E962A694C9E98627CC069595A4992" authorityName="Smith" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="EE5577F127A1A6F555CCF8E689F146D8" italics="true" pageId="0" pageNumber="1">Rhaphaulus perakensis</emphasis>
</taxonomicName>
, syntype NHMUK 1897.3.15.41 from Larut, Perak
<emphasis id="EB8FD843E30DAD0FE03FEBB6DD0BB3BC" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="6222C598D0FCAEB7DB94BBFD6602E27C" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus ascendens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ascendens">
<emphasis id="8ABE9DEE6B969CD6189F36B57C32430E" italics="true" pageId="0" pageNumber="1">Rhaphaulus ascendens</emphasis>
</taxonomicName>
, syntype UMZC I.100025 from Patalung, Malay Peninsula
<emphasis id="DF4F00CD23A8846FD12D7089A53117FB" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
<taxonomicName id="4466336961C2D9D83205967FCD4DE417" authorityName="Pall-Gergely, Hunyadi &amp; Maassen" authorityYear="2014" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">
<emphasis id="6756F5C98673B567BEB803AF23719B10" italics="true" pageId="0" pageNumber="1">Rhaphaulus tonkinensis</emphasis>
</taxonomicName>
<emphasis id="8AE77FA2F915C63E60155750B2CE3B57" bold="true" pageId="0" pageNumber="1">E</emphasis>
holotype HNHM 98757 and
<emphasis id="0121FF799B03E8970BC897AE82E203F3" bold="true" pageId="0" pageNumber="1">F</emphasis>
specimen CUMZ 12163/1 from Luang Cave, Chiang Rai. Photo: B.
<normalizedToken id="6B0F148F340467D91E246B7E64441D2B" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
(
<emphasis id="5C39A1017D10DE922509F08355268D85" bold="true" pageId="0" pageNumber="1">E</emphasis>
)
</paragraph>
</caption>
<caption id="849CBC1AD61E8DA07B34C53D3B04864E" doi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1" start="Figure 14" startId="F14">
<paragraph id="0EF118CCE1DE098D667C289E460ED030" pageId="0" pageNumber="1">
<emphasis id="E5D61BCC77D64DE2EA1E5517C7ABF295" bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
<emphasis id="9F79DCA16D289D65855710B2C7DAE3E8" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="A9404C0DD26F8F9BE4FFF8A1BB07FC5E" authorityName="Pall-Gergely, Hunyadi &amp; Maassen" authorityYear="2014" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">
<emphasis id="7590CCE68BE3AD6CC8D6616A3A4AC585" italics="true" pageId="0" pageNumber="1">Rhaphaulus tonkinensis</emphasis>
</taxonomicName>
, specimen CUMZ 12163/2 from Luang Cave, Chiang Rai.
<emphasis id="1BA4B9F25F8703DB15034CFA86BD3B2C" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
<taxonomicName id="DC8CA7F838DC064D8135CD77AF8151CA" baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">
<emphasis id="72DB34EBCE57E0EA1DA02F41603FE280" italics="true" pageId="0" pageNumber="1">Rhaphaulus chrysalis</emphasis>
</taxonomicName>
<emphasis id="ADEC3E69A66088F6D75B805559EE825C" bold="true" pageId="0" pageNumber="1">B</emphasis>
possible syntype NHMUK 2013.04.16 from Siam
<emphasis id="B7E7A56B83769D9C3CE17B44DE02452C" bold="true" pageId="0" pageNumber="1">C</emphasis>
specimen NHMUK 1871.9.23.52 from Burma, and
<emphasis id="185E90F62C0CF33E48CE94C3C6446B31" bold="true" pageId="0" pageNumber="1">D</emphasis>
specimen NHMUK 1903.7.1.3073 from Molmein.
</paragraph>
</caption>
<caption id="CBA2113931170CD8F265F453A99F236E" doi="10.3897/zookeys.1119.85400.figure15" httpUri="https://binary.pensoft.net/fig/735376" pageId="0" pageNumber="1" start="Figure 15" startId="F15">
<paragraph id="68078D0A748D8A7F845F72FBFC0D90CD" pageId="0" pageNumber="1">
<emphasis id="C7E43337CB831D59A82ADA177CF2AAB7" bold="true" pageId="0" pageNumber="1">Figure 15.</emphasis>
<taxonomicName id="DD6CA6224E48D1BB1E79170DC19A1B58" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
<emphasis id="A94797C14388143535F7BB99E3E04784" italics="true" pageId="0" pageNumber="1">Tortulosa tortuosa</emphasis>
</taxonomicName>
<emphasis id="68D83352A6B49DCE7640B156391826F0" bold="true" pageId="0" pageNumber="1">A</emphasis>
lectotype of
<taxonomicName id="9B93198FC88F9203CD1163183A0F6207" authorityName="Tomlin" authorityYear="1948" class="Gastropoda" family="Pupinidae" genus="Perlisia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perlisia tweediei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tweediei">
<emphasis id="996546261668FE1D146E43465A11C415" italics="true" pageId="0" pageNumber="1">Perlisia tweediei</emphasis>
</taxonomicName>
NHMUK 1948.10.2.6 from Kaki Bukit, Perlis
<emphasis id="C68636113CF0EB0E6A0DB55EFC53C5A7" bold="true" pageId="0" pageNumber="1">B</emphasis>
holotype of
<taxonomicName id="E2F5DBD8F9CA439785ABF5F9279308D5" authorityName="Thach" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa huberi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="huberi">
<emphasis id="51AFA793C696330373866974C2FE63D6" italics="true" pageId="0" pageNumber="1">Tortulosa huberi</emphasis>
</taxonomicName>
MNHN-IM-2000-34054 from Krabi
<emphasis id="99A734449069244A739F9087BE2330A9" bold="true" pageId="0" pageNumber="1">C</emphasis>
holotype of
<taxonomicName id="F83DD0EFD81B78C9A5A2EC8B9D61FD55" authorityName="Thach &amp; F. Huber" authorityYear="2018" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa schileykoi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="schileykoi">
<emphasis id="439A3A703110E45973C19976DF4FF2F8" italics="true" pageId="0" pageNumber="1">Tortulosa schileykoi</emphasis>
</taxonomicName>
MNHN-IM-2000-34055 from Phang Nga
<emphasis id="DAEFFBA24A21066FE5DB43D0E9AF1681" bold="true" pageId="0" pageNumber="1">D</emphasis>
specimen CUMZ 12166 from Tham Suea Temple, Krabi, and
<emphasis id="5266A10ED93521CBE09C35404D293786" bold="true" pageId="0" pageNumber="1">E-H</emphasis>
operculum of specimen CUMZ 12160 from Tham Kanlayanamit Temple, Nakhon Si Thammarat, showing
<emphasis id="2922A88431D18FD282E3F334FC4FC0C6" bold="true" pageId="0" pageNumber="1">E</emphasis>
outer operculum
<emphasis id="9E6DABBB1F03FB1149B3C226295BAF59" bold="true" pageId="0" pageNumber="1">F</emphasis>
inner operculum
<emphasis id="3A21DE20F5B0A5947E76575F8A5FCE49" bold="true" pageId="0" pageNumber="1">G</emphasis>
side view (inner surface up), and
<emphasis id="4275808D4D82A8245DF641F15F785E0E" bold="true" pageId="0" pageNumber="1">H</emphasis>
spring-like inner operculum when extended by force. Photo: M. Caballer, MNHM (
<emphasis id="13D59742D7320808D5AFBC2EDF3626AE" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
).
</paragraph>
</caption>
<caption id="5CBAA706AF31102B31F121D8239910D8" doi="10.3897/zookeys.1119.85400.figure16" httpUri="https://binary.pensoft.net/fig/735377" pageId="0" pageNumber="1" start="Figure 16" startId="F16">
<paragraph id="93B7C09372BB541B6622F9208A63AF60" pageId="0" pageNumber="1">
<emphasis id="0F756DBF8D5F53E8A2F41EB16A4E99A1" bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
Infraspecific variation of shell shape and colour found in the same collecting locality of
<taxonomicName id="5B48B4A34EB561853BD7184DB362E006" baseAuthorityName="Ferussac" baseAuthorityYear="1821" class="Gastropoda" family="Pupinidae" genus="Tortulosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortulosa tortuosa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tortuosa">
<emphasis id="68BBA27FB55CA8E9578AF8CA443E461A" italics="true" pageId="0" pageNumber="1">Tortulosa tortuosa</emphasis>
</taxonomicName>
, CUMZ 12166 from Tham Suea Temple, Krabi.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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