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<mods:title id="126D4061846F825271C359DE451FEC71">A review of the Neotropical dung beetle genera Deltorhinum Harold, 1869, and Lobidion gen. nov. (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<paragraph id="847B36E7FFD7F5393ABAFA7E3A8FFA1E" blockId="1.[151,477,1453,1479]" box="[151,477,1453,1479]" pageId="1" pageNumber="36">
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<taxonomicName id="43C44D64FFD7F5393ABAFA7E3A8FFA1E" ID-CoL="42TC" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[151,477,1453,1479]" class="Insecta" family="Scarabaeidae" genus="Deltorhinum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B6B0EAF5FFD7F5393ABAFA7E3A8FFA1E" bold="true" box="[151,477,1453,1479]" pageId="1" pageNumber="36">
<emphasis id="B6B0EAF5FFD7F5393ABAFA7E3A61FA1E" bold="true" box="[151,307,1453,1479]" italics="true" pageId="1" pageNumber="36">Deltorhinum</emphasis>
<bibRefCitation id="E0554B16FFD7F5393B17FA7E3A8FFA1E" author="Harold" box="[314,477,1453,1479]" pageId="1" pageNumber="36" refString="Harold, E. von (1867) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 2, 94 - 100." type="journal article" year="1867">Harold, 1867</bibRefCitation>
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<paragraph id="847B36E7FFD7F5393ABAFA26397CF9D5" blockId="1.[151,824,1525,1868]" box="[151,558,1525,1548]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAFA263A72F9D5" author="Harold" box="[151,288,1525,1548]" pageId="1" pageNumber="36" refString="Harold, E. von (1867) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 2, 94 - 100." type="journal article" year="1867">Harold 1867</bibRefCitation>
: 96 [original description]
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<paragraph id="847B36E7FFD7F5393ABAF9C63A8DF9F5" blockId="1.[151,824,1525,1868]" box="[151,479,1557,1580]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAF9C63A72F9F5" author="Harold" box="[151,288,1557,1580]" pageId="1" pageNumber="36" refString="Harold, E. von (1869) Tom. IV. Scarabaeidae, In: Gemminger, M. &amp; Harold, E. von (eds.) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus autoribus, E. H. Gummi, Munich, pp. 979 - 1346, [1347 - 1353]." type="book chapter" year="1869">Harold 1869</bibRefCitation>
: 1003 [catalogue]
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<paragraph id="847B36E7FFD7F5393ABAF9E63AE7F995" blockId="1.[151,824,1525,1868]" box="[151,437,1589,1612]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAF9E63A40F995" author="Gillet" box="[151,274,1589,1612]" pageId="1" pageNumber="36" refString="Gillet, J. J. E. (1911) Scarabaeidae: Coprinae I. Coleopterorum Catalogus, 38, 1 - 263." type="journal article" year="1911">Gillet 1911</bibRefCitation>
: 54 [catalogue]
</paragraph>
<paragraph id="847B36E7FFD7F5393ABAF9863955F975" blockId="1.[151,824,1525,1868]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAF9863A0FF9B5" author="Luederwaldt" box="[151,349,1621,1644]" pageId="1" pageNumber="36" refString="Luederwaldt, H. (1931) O genero Ontherus (Coleop.) (Lamellic. - Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista, 17, 363 - 422." type="journal article" year="1931">Luederwaldt 1931</bibRefCitation>
: 366 [identification key to genus]
<bibRefCitation id="E0554B16FFD7F5393ABAF9A63A74F955" author="Paulian" box="[151,294,1653,1676]" pageId="1" pageNumber="36" refString="Paulian, R. (1938) Contribution a l'etude des Canthonides americains (Coleopt. Lamellic.). Annales de la Societe Entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
: 234 [identification key to genus]
<bibRefCitation id="E0554B16FFD7F5393ABAF9463A0FF975" author="Blackwelder" box="[151,349,1685,1708]" pageId="1" pageNumber="36" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 205 [checklist]
</paragraph>
<paragraph id="847B36E7FFD7F5393ABAF9663940F935" blockId="1.[151,824,1525,1868]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAF9663ACFF915" author="Vulcano" box="[151,413,1717,1740]" pageId="1" pageNumber="36" refString="Vulcano, M. A. &amp; Pereira, F. S. (1967) Sinopse dos Passalidae e Scarabaeidae s. str. da Regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica, Vol. 5 (Zoologia), 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967</bibRefCitation>
: 533603 [identification key to genus] Vaz-de-Mello 2000: 192 [checklist]
</paragraph>
<paragraph id="847B36E7FFD7F5393ABAF926395EF8D5" blockId="1.[151,824,1525,1868]" box="[151,524,1781,1804]" pageId="1" pageNumber="36">
<bibRefCitation id="E0554B16FFD7F5393ABAF9263A31F8D5" author="Medina" box="[151,355,1781,1804]" pageId="1" pageNumber="36" refString="Medina, C. A., Lopera-Toro, A., Vitolo, A. &amp; Gill B. (2001) Escarabajos Coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana, 2 (2), 131 - 144." type="journal article" year="2001">Medina et al. 2001</bibRefCitation>
: 134 [checklist]
</paragraph>
<paragraph id="847B36E7FFD7F5393ABAF8C6397EF895" blockId="1.[151,824,1525,1868]" pageId="1" pageNumber="36">
Vaz-de-Mello 2008: 12 [systematic position]
<bibRefCitation id="E0554B16FFD7F5393ABAF8E63AD8F895" author="Almeida" box="[151,394,1845,1868]" pageId="1" pageNumber="36" refString="Almeida, L. M. &amp; Mise, K. M. (2009) Diagnosis and key of the main families and species of South American Coleoptera of forensic importance. Revista Brasileira de Entomologia, 53 (2), 227 - 244." type="journal article" year="2009">Almeida &amp; Mise 2009</bibRefCitation>
: 237 [mention]
</paragraph>
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<paragraph id="847B36E7FFD7F53A3ABAF8A9381DFDFE" blockId="1.[151,1437,1914,2015]" lastBlockId="2.[151,1437,152,551]" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
<emphasis id="B6B0EAF5FFD7F5393ABAF8A93A1CF84D" bold="true" box="[151,334,1914,1940]" pageId="1" pageNumber="36">Redescription.</emphasis>
Small to moderate in size (6.514.0 mm). Body oval to elongate-oval. Color light brown in teneral specimens to black, with distinct metallic sheen only in
<taxonomicName id="43C44D64FFD7F5393986F8733F11F860" box="[939,1091,1952,1977]" class="Insecta" family="Scarabaeidae" genus="Deltorhinum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B6B0EAF5FFD7F5393986F8733F11F860" box="[939,1091,1952,1977]" italics="true" pageId="1" pageNumber="36">D. robustum</emphasis>
</taxonomicName>
. Clypeus regularly tapering toward apex, which is moderately to strongly upturned; apex at most broadly emarginate, never with two distinct teeth. Clypeofrontal suture always distinctly carinate. Pronotal anterior portion modified, produced into a single broad or two narrow lobate projections and additional modifications; simple only in
<taxonomicName id="43C44D64FFD4F53A3F29FF6E3EC6FF0F" box="[1284,1428,189,214]" class="Insecta" family="Scarabaeidae" genus="Deltorhinum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B6B0EAF5FFD4F53A3F29FF6E3EC6FF0F" box="[1284,1428,189,214]" italics="true" pageId="2" pageNumber="37">D. robustum</emphasis>
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, which has a somewhat triangular impression on each side of midline. Pronotal disc with a deep and well defined longitudinal sulcus along most of midline. Elytra strongly convex, with 7 striae on disc, stria 8 distinct from stria 9 only at apex. Elytral pseudepiplera concealed in dorsal view. Pygidium narrowly sulcate basally, sulcus broadly arcuate or slightly bisinuous in distal view. Protibia obliquely truncate anteriorly; protarsi atrophied and usually broken off in non teneral individuals. Mesotibiae and metatibiae very wide at apex in ventral view; tarsi reduced in length, first tarsomere at most 1.5 X longer than wide at apex, usually as wide at apex as long. Sternite 8 unmodified medially. Aedeagus unmodified, parameres simply tapering toward apex, lacking denticles, hooks or rough surface. Internal sac with variously shaped sclerites, usually with a large apical sclerite seemingly forming a guide for the flagellum.
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