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<document ID-CLB-Dataset="229647" ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-GBIF-Dataset="6ba6d2ca-5dad-4270-914f-1042327d503c" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="0C12234D630E5A04AB9A64A81C0B686A" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana Domahovski &amp; Cavichioli 2023, gen. nov." docType="treatment" docUuid="C82FE565-FF2B-4B90-99AE-C115857B6A29" docUuidSource="ZooBank" docVersion="5" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1732669275789" updateUser="ExternalLinkService">
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<mods:title id="656556843C591C7DBDE30BA9515C6FAA">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:namePart id="99DF67624762E2721F1B92C03E0A492D">Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier id="BA2E9AB8AE03A570DB8BF9ED246C56FB" type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation id="1E83E82354897328BE5557461A361DCE">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier id="46CD4BB37441E8FC4FAE1C9E59A14D1A" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="BB1D9EDA02AD65DAD558C99659FA0925">Cavichioli, Rodney Ramiro</mods:namePart>
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<mods:affiliation id="473BC62A56F0E3BA052EED02624BECE9">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:date id="9AB41E88194643EF47494AE2F1381F75">2023</mods:date>
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<treatment id="0C12234D630E5A04AB9A64A81C0B686A" ID-GBIF-Taxon="209013697" LSID="urn:lsid:zoobank.org:act:C82FE565-FF2B-4B90-99AE-C115857B6A29" httpUri="http://treatment.plazi.org/id/0C12234D630E5A04AB9A64A81C0B686A" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
<subSubSection id="6CFC78112A7ACFF4BBF2DCEC61112572" pageId="0" pageNumber="79" type="nomenclature">
<paragraph id="AFFE5013B940EBBECCCF35EFD6DF7BCE" pageId="0" pageNumber="79">
3.3.2.
<taxonomicName id="3CBFC46631BC094E2DA396F79A8D214C" LSID="https://zoobank.org/C82FE565-FF2B-4B90-99AE-C115857B6A29" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus" status="gen. nov.">Coarctana</taxonomicName>
<taxonomicNameLabel id="5C760F19B492AC64F7187990DBBBBF21" pageId="0" pageNumber="79">gen. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C43C4349EF0A8B9F6F0AF585D10DCD18" pageId="0" pageNumber="79" type="description">
<paragraph id="1D4866F76957351E45583B175BF4F344" pageId="0" pageNumber="79">
<figureCitation id="117696CAD230721FCF06CE5DB4114F66" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">Figs 17</figureCitation>
<figureCitation id="09C529E200FE1A7344C188A6D689D965" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">, 18</figureCitation>
<figureCitation id="1CADE8DEA74C62EE823013473D350C98" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">, 19</figureCitation>
<figureCitation id="48180134AB83189FFCA9C687EA2F273F" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">, 20</figureCitation>
<figureCitation id="268744CB8C1AE4B73DA2CC49B2345C6F" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">, 21</figureCitation>
<figureCitation id="DD6FFCC1D825EE51A1B4EE61EF407C28" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">, 22</figureCitation>
<figureCitation id="3079BE490A8B55AC14C8AD567F36F93E" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">, 23</figureCitation>
<figureCitation id="14BFCD802FB669619487CEECE17A8A14" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">, 24</figureCitation>
<figureCitation id="BD2E7B4A25D0592284744951F2C07887" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">, 25</figureCitation>
<figureCitation id="3F9CB49F77D834FF29528CC8BA753477" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">, 26</figureCitation>
<figureCitation id="A059AACDCC293B55F3B9B321F976E3CD" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">, 27</figureCitation>
<figureCitation id="483315A29F43D2CD9BFC0971AE0D7A07" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">, 28</figureCitation>
<figureCitation id="A72E11828BE8111F73B8BDC88D5E5888" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Coarctana vilavelha sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure29" httpUri="https://binary.pensoft.net/fig/798182" pageId="0" pageNumber="79">, 29</figureCitation>
<figureCitation id="0C7E04D0B1D85A75BBC437D086C0983C" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Coarctana vilavelha sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure30" httpUri="https://binary.pensoft.net/fig/798183" pageId="0" pageNumber="79">, 30</figureCitation>
<figureCitation id="8A784942E4CB78A93F48C8A6C1A9E461" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31</figureCitation>
<figureCitation id="13878EF42D7184B2EF3AEB552B233D74" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DB59105ECE8B2009BB8744B694041292" pageId="0" pageNumber="79" type="type species">
<paragraph id="6DDBB7787077C6C971A2EEAD954CDED6" pageId="0" pageNumber="79">Type species.</paragraph>
<paragraph id="DBAC45BD4DB576DAFA4A5FF7EC68DF80" pageId="0" pageNumber="79">
<taxonomicName id="730C6ED52FC92FC5FF7883D5746E0791" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Acuera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuera rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana">
<emphasis id="D22269CC3B408378EAB56F0EC54786E9" italics="true" pageId="0" pageNumber="79">Acuera rana</emphasis>
</taxonomicName>
DeLong &amp; Freytag, 1982: 309.
</paragraph>
</subSubSection>
<subSubSection id="EA031996E67DBEC2CD82040AA87F0C75" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="90B40344C2EB02A86346188AFA8A59CE" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="A88BC56BF818AEFF406F89442DEB2ADE" pageId="0" pageNumber="79">
Small to medium size leafhoppers (7.0-10.5 mm). Head, in dorsal view (Figs
<figureCitation id="E519FD767267625D708AC686633362C3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="F50231F089D2C521F4B7E362EF888BED" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
), moderately produced anterad, median length slightly longer than half interocular width; crown surface with transverse striae; ocellus slightly closer to inner margin of eye than to midline. Head, in lateral view (Figs
<figureCitation id="AF4F29C81CC23264106D09DC73EC7072" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
,
<figureCitation id="D01D6A7800B2B22A76A613A287807E4C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
), crown-face transition thin, with 3-6 transverse carinae. Pronotum (Figs
<figureCitation id="3F6FEDC23629180CFC6B87956985935C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="B313D8C4ADD994B56FD1499B53FFBC3C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) with black or brown punctures. Forewing (Figs
<figureCitation id="DAEE9F9694FCABEF34D1BF7674420815" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20D</figureCitation>
,
<figureCitation id="2D2F89B60F3F4C15539B0A9142BED320" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22D</figureCitation>
) M vein with segment after the divergence between R+M and before the cross vein
<emphasis id="D619FCE7FCC6B727B64A3C06EFD3D082" italics="true" pageId="0" pageNumber="79">m-cu</emphasis>
1 2
<normalizedToken id="7F72C291ECFAC804D5887E40D74BCC8A" originalValue="×">x</normalizedToken>
longer than the length of
<emphasis id="45844D5FEB2B08DE6E141559DF99ABA6" italics="true" pageId="0" pageNumber="79">m-cu</emphasis>
1; appendix reduced. Subgenital plate (Figs
<figureCitation id="523A135898766872FA5CFD98BAC35045" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
,
<figureCitation id="F1C62F5D7F725FEEB6494FDC6F26C7B9" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22H</figureCitation>
) usually with long setae. Aedeagus (Figs
<figureCitation id="1169B7F9BCE26655C3C10D7596762161" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L</figureCitation>
,
<figureCitation id="3344048F3D7864E204EC4B1F1E510B17" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18L</figureCitation>
) strongly constricted between dorsal apodeme and atrium; dorsal apodeme with pair of long processes, strongly curved dorsally.
</paragraph>
<caption id="AB0A0B16CBF9910090672AEA8FFA23A4" doi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79" start="Figure 17" startId="F17">
<paragraph id="10F1EA848C8CE32CF7A36615B8A39610" pageId="0" pageNumber="79">
<emphasis id="DB6AC16A55A8182C84C69D6CF75A9CCD" bold="true" pageId="0" pageNumber="79">Figure 17.</emphasis>
<taxonomicName id="C36ABBBAC36BCC619986FE4E3884FB91" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana">
<emphasis id="B622D9B947937B98CF51FE03519CE66C" italics="true" pageId="0" pageNumber="79">Coarctana rana</emphasis>
</taxonomicName>
<emphasis id="D112D4F80888F413E6291D6F249D49BA" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
, male.
<emphasis id="A319FE190166794405D33A2CA6E7BC76" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="CB5D2F0EBC1C5A78CFBA244015E3632A" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="8551C602F672CF4C02E36EAD802B9E10" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="AF69063281AC2B4AB1A7A7D56152539D" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="BC0E6D9E134590DDF6E60C157BA17F8D" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="F12BB91FB626E27A0DEFEB5B8A8259CF" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="A6FA2578EFDD732387DF6005485E3F26" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view;
<emphasis id="CD26F534CDAF32C513B0C26D26CB3480" bold="true" pageId="0" pageNumber="79">g</emphasis>
apical portion of pygofer, dorsal view.
<emphasis id="1B7AA47AF59875FA104F79ED444A354D" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="F9A77D1D8785043BEA5969559F077C69" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="57DB6E9C135ECA93197339F840409E58" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="682F26CE28AF6C436B3E35EA20579AE2" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="672E696D0D0AEE102C6F370DCFB30360" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="67F68DD0F665E409AA906DF784E7F234" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
<caption id="BA9836E7694A4269F0BDD03C3DBC0CAA" doi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79" start="Figure 18" startId="F18">
<paragraph id="66B9DB68BF1CDD0E7B5868E60CD7D74E" pageId="0" pageNumber="79">
<emphasis id="5E1E3322EFAF852C2320C33BFA51BD2B" bold="true" pageId="0" pageNumber="79">Figure 18.</emphasis>
<taxonomicName id="9B1A334D85B1D976076E2EFBEB672C91" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana asymmetrica" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="6BE72208D25A3E4805309D018683E751" italics="true" pageId="0" pageNumber="79">Coarctana asymmetrica</emphasis>
</taxonomicName>
<emphasis id="B9B8BE3906F37B9F73BE576370DA7991" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="034BCE0C4100C57E99DEBAA9DF6B0395" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="372E8566E7AFBAA33FDD5888D190F1A4" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="2947A62914F9A412E9ACA2E115015476" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="6AD733125F4922211D89C973BE3DD245" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="6B640B93F6042A18363A0222C182ADC6" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="CCA8801146D37268D715F201058CBF26" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="3BAC308A7CC7FE1CF63550DB004D304E" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="67A3F1177A982378DB8D9E2E0200ECFF" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="7D44C0D209A23CE1E5A3E3D0AF607E8D" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="E39D18F9D76C2E8A5F300FBFCFE07A3E" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="D5ABFB7F83C30E8132F063DB8125C0A2" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="DA31314071C8880DD552D76D152A8B63" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="09991997467E06864DE60BD35CC5AE60" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="87D44617FFA14F12E4A3A2B2E4818016" pageId="0" pageNumber="79" type="coloration">
<paragraph id="9688DE8855ECF3EF997E5977B5FA94A2" pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph id="2A0A728AAE07F73436D44B20842BD5EE" pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation id="F9DD6F6C33C9063281A21E78C56C8AC8" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Coarctana vilavelha sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure29" httpUri="https://binary.pensoft.net/fig/798182" pageId="0" pageNumber="79">29</figureCitation>
,
<figureCitation id="9BC8EA18DB88F1E9C4A6B45822058DF1" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32</figureCitation>
) reddish-brown or yellowish-brown. Head (Figs
<figureCitation id="56B7CE2BED9A6C28B5C51E91CA8D4D73" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="00EFCC9970569C547F5D4588BD8929C0" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Coarctana glabra sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79">19A</figureCitation>
) without black punctures, usually with a pair of small circular black spots, near posterior margin, behind ocelli. Face (Figs
<figureCitation id="6B5D956A3D17D1F78969A7B5BFFC736B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
,
<figureCitation id="91F1017BCD32768078920B84D0D1957D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18B</figureCitation>
) usually without maculae. Pronotum (Figs
<figureCitation id="075BD24ED3D162E8BB8AD1502B68F1BE" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="D89C974F8A1BE915E96DD07281999366" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) with black or brown punctures and frequently with irregular black spots near anterior margin. Forewing (Figs
<figureCitation id="733619192B2D343725E24F9FF1FC2417" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17D</figureCitation>
,
<figureCitation id="EF0064AB7DC11E7DBA44B60D63517E1B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23D</figureCitation>
) with black maculae on apex of anal veins, cross veins of discal and apical cells, usually with additional mottled black spots, or with a transverse brown stripe over subapical cells. Legs (Figs
<figureCitation id="7ACD3191718F3DD2151D08195A4BCE3B" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31</figureCitation>
,
<figureCitation id="9186E765C6B8B4846B9B1F71132876A6" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32</figureCitation>
) without large black maculae or punctures. Metatibia (Figs
<figureCitation id="0B5BF8466DD7460F4AE9D3EAA4C4CBCD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
,
<figureCitation id="15219F18CF784B628185F78D37245327" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
<caption id="92A5600786173742D274A116A935E346" doi="10.3897/asp.81.e81961.figure19" httpUri="https://binary.pensoft.net/fig/798172" pageId="0" pageNumber="79" start="Figure 19" startId="F19">
<paragraph id="961B51A5721F6557EAF11D8AE3A36336" pageId="0" pageNumber="79">
<emphasis id="A8954BE0B6B706C1D3B4465EFB7070C9" bold="true" pageId="0" pageNumber="79">Figure 19.</emphasis>
<taxonomicName id="6B89CCABB66E8AA424450174F4F8DF1D" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana glabra" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="D8435B4F40CED9385A611D1CD7545F45" italics="true" pageId="0" pageNumber="79">Coarctana glabra</emphasis>
</taxonomicName>
<emphasis id="5B52E418407F163A9494B6063F81B4A6" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="BE153B8C595A2A8860C50ED6145FA609" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="27525C74E9E62104F88607EFF4D6AB83" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="4EA5EFD142BF07ECC9F13B2B4C5E72F0" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="B67F3A8EF29FD77DCBE5B18AA019A25A" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="6DF8354F4506CD606B2666D478EF664A" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="C513F33A5202D49FC5A6924E6A92F3D0" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="2B4F615211C0EE5AE5BC7DCC0129CB19" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="E6804139E6AB2A9D4989E18F23899694" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="4E2B037BD92A8401A4D150B3843039EB" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="BDEECEB9505BDC0876033F1A0F7F036D" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="7FB992476B4E6960B9408F19E1CEF99A" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="84D07C273E9B2464BE896262346AEDF2" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="E2E28B6A226721A1EA801ED4DC905543" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B0AC415AED205533F34335476BF3E4FE" pageId="0" pageNumber="79" type="description">
<paragraph id="0BFAA5066F8BED2CFD6A3D6FBE679C71" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="C8F6E608C6F64C75D720676CC4BA2FD0" pageId="0" pageNumber="79">
Head, in dorsal view (Figs
<figureCitation id="8916ECA72297B2C471DE18ACA8D1C27A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="BE200E996972C99EA54FC725FB9E7CE6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
), moderately produced anterad, median length slightly longer than half interocular width; transocular width 8.0-8.5 tenths of pronotum humeral width; crown surface with transverse striae; ocellus slightly closer to anterior margin or equidistant between anterior and posterior margins of crown and slightly closer to inner margin of eye than to midline. Head, in frontal view (Figs
<figureCitation id="6C0AF123C65EA18F384ECB1B797ED214" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
,
<figureCitation id="52C15A487FE225F8271DECF216093789" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18B</figureCitation>
), face wider than high; frons texture shagreen, surface just below the crown-face transition not excavated; frontogenal suture surpassing antennal ledge, extending to anterior margin of crown; supra-antennal lobe oblique, advancing over the frons for short distance; gena with ventrolateral margin rounded and slightly angled medially; maxillary plate produced ventrally as far as clypeus apex. Head, in lateral view (Figs
<figureCitation id="87025444AA20B120202EE9C06C80AA17" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
,
<figureCitation id="67E78F98310035823B350698663F990C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
), crown-face transition thin, with 3-6 transverse carinae; anterior margin of crown slightly projected over anterior margin of eye; frons not inflated. Pronotum, in dorsal view (Figs
<figureCitation id="4CC861497141646400470FD5E4FCB87C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="3D8D14251735553FEF47CEC882AFCAC4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
), with transverse striae on disc and posterior third; lateral margins as long as eye length; posterior margin slightly excavated; in lateral view (Figs
<figureCitation id="85A785394246748ED9EB159B0561A887" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
,
<figureCitation id="F82F2CBBFC8FEBF88D0DFE56F950ADAC" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
), moderately declivous, continuous with head declivity. Mesonotum (Figs
<figureCitation id="49F7C612EE1748F97816DF3D001F392D" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="FC88C2D0D38AE2763BBEC40FC7F91CC1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
), slightly wider than long; scutellum (Figs
<figureCitation id="F34C95BCA87F60F04B68F0CE49E7B0A7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
,
<figureCitation id="F272AC43A593D82816D8101BE2C0BF19" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) slightly swollen. Forewing (Figs
<figureCitation id="F5449DE42AFB147C840A3453166AA240" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Coarctana nigromedia sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79">20D</figureCitation>
,
<figureCitation id="7FEDB27F70531949DB5DBFF0515FBE9F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22D</figureCitation>
) without extra numerary veins; M vein with segment after the divergence between R+M and before the cross vein
<emphasis id="CF0261995AC15119AA749EE5C8A2FB50" italics="true" pageId="0" pageNumber="79">m-cu</emphasis>
1 2
<normalizedToken id="F8E5B7AA2B0640B2F20B1838EE4CB377" originalValue="×">x</normalizedToken>
longer than the length of
<emphasis id="0D40D5E62ABE9B6E8C65F7E7AC9CF890" italics="true" pageId="0" pageNumber="79">m-cu</emphasis>
1; appendix reduced, bordering first to second apical cells; apex rounded. Profemur moderately elongated, 3.5
<normalizedToken id="A5301CDF2358992273DB8963646DB83C" originalValue="×">x</normalizedToken>
longer than wide; AD, AM, and PD rows reduced and poorly defined, with exception of apical setae AD1, AM1, and PD1, respectively; IC row formed by slightly arched comb of fine setae, beginning at distal half of femur and extending to AM1; AV row with 4-6 setae restricted to basal half; PV with 1-3 setae. Protibia, in cross-section, semi-circular, dorsal surface with longitudinal carina adjacent to PD row; AV row formed by long setae, slightly longer and thicker towards apex; dorsal rows with apical AD1 and PD1 setae developed; AD row without differentiated setae. Mesotibia with dorsal surface rounded. Metafemur with setal formula 2:2:1. Metatibia with AD row with up to 4 intercalary setae between macrosetae; PV row with setae of apical half formed by sequence of a longer and thicker seta, interspersed with 2-4 thinner and shorter setae, ending with a long and thick seta. Metatarsomere I ventral surface with rows of non-cucullate setae; outer row absent or with setae very reduced in size; inner row with 5-8; pecten with 3-4 platellae, flanked by one inner and one outer tapered seta. Metatarsomere II pecten with 2 platellae, flanked by two inner and one outer tapered seta.
</paragraph>
<caption id="234C8A906AE68590B01E375304B57CA7" doi="10.3897/asp.81.e81961.figure20" httpUri="https://binary.pensoft.net/fig/798173" pageId="0" pageNumber="79" start="Figure 20" startId="F20">
<paragraph id="8645D1C9CB143764381CA1ED3F978C70" pageId="0" pageNumber="79">
<emphasis id="5D27318DB4D6BEBEF0FD956428B283E0" bold="true" pageId="0" pageNumber="79">Figure 20.</emphasis>
<taxonomicName id="B37EE7493C64C3DCFF148839C0898E21" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana nigromedia" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigromedia">
<emphasis id="34F9EB6A65910886062EF43A79CB6FFC" italics="true" pageId="0" pageNumber="79">Coarctana nigromedia</emphasis>
</taxonomicName>
<emphasis id="17967640F0F37BB8473F934983B796BF" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="345F27471E9EE128FAAC8F32CD403B6A" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="6563E08B1DBD25AA31C1DA5D347650ED" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="96B28089CBB32E139F9880544DD2CF00" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="B966EA1DC0BB8FFF9C17A53A5903D782" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="6C4C39D57AF53C151CB0E6192BC6F322" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="322A49C57DD829FEB39772201C16E52C" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="3BDE79EF8B17FAB97DCADA680DF66BBE" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="92D39C93EA4BB134D288D35F5D40E0FA" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="BE98B7134CEFF4627920C7F33828F8E6" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="D68F01343C8FC91C243C2C112373C72C" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="4F042DDD29285595F3FDF11ECD7DCED4" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="B72291AE1E190B39CA995C7009971DB2" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="0B8B2D0ADC3887265153C4FFC9625C26" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="379DF86897CF94A9A5F76ABF2E194960" pageId="0" pageNumber="79" type="male terminalia">
<paragraph id="DD99130FE1A30067CB30AF705805534E" pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph id="84369D8A524A5622206C75C46E84D8D8" pageId="0" pageNumber="79">
Sternite VIII not covering subgenital plates. Pygofer, in lateral view (Figs
<figureCitation id="D6661A806773363F581A513377522C2E" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22G</figureCitation>
,
<figureCitation id="B84129B0A2D7AB719D797DD8F81C9A24" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Coarctana oricula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79">23G</figureCitation>
), with basodorsal processes usually developed and extending ventrally. Anal tube membranous, without processes. Subgenital plate (Figs
<figureCitation id="9267524E6304CC44DD4C4D2D096C92F8" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
,
<figureCitation id="5B535AF11E3CFF5F6F1AD18B76809007" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coarctana occultata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79">22H</figureCitation>
) usually with long setae. Aedeagus (Figs
<figureCitation id="945F5ED7A0096CB7AEDE17B4C1AF7930" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L</figureCitation>
,
<figureCitation id="A6856936AA498113713E2ADD694B1C02" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18L</figureCitation>
) strongly constricted between dorsal apodeme and atrium, dorsal apodeme bent ventrally, with pair of long processes, strongly curved dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C914E0CBE2DAEC879EC574F0953BBDED" pageId="0" pageNumber="79" type="female terminalia">
<paragraph id="E2DD76F1471E2FE5AC9CCA0869BBF20C" pageId="0" pageNumber="79">Female terminalia.</paragraph>
<paragraph id="A4A99EE67702A0341BC6935297F5A24C" pageId="0" pageNumber="79">
Pygofer (Fig.
<figureCitation id="71D9CB160944C2A7A1F11AE1695FAAFE" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21B</figureCitation>
,
<figureCitation id="8A9ECF7AC31806B9213EEF1C6F1D7F18" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Coarctana oricula sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79">24B</figureCitation>
) with macrosetae distributed on posteroventral quadrant and dorsoapical third. Ovipositor slightly curved dorsally. First valvula (Figs
<figureCitation id="511ED179FD7E2F5C1EA4B1CC32C60AB6" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21C</figureCitation>
,
<figureCitation id="71701ED2BAD22F5CF8D86C00C25970A0" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27C</figureCitation>
) not broadened medially; dorsal sculptured area strigate, beginning after half length of valvula; ventral interlocking device distinct on basoventral 2/5 of blade; apex tapered and acute. Second valvula (Figs
<figureCitation id="2A501E77EF6BD86CBBC9AD41B5314EE4" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21E</figureCitation>
,
<figureCitation id="F2E34F6848667F7A0F2751D2544FEDC7" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27E</figureCitation>
) wider near half length; dorsal protuberance rounded, located after half length of blade; dorsal margin with few very small, rounded teeth, irregular shaped, present subapically; ventral margin without denticles; apex tapered and subacute. Second valvifer (Figs
<figureCitation id="4005F2F34BDDA4446889C8EEE44B1B9B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21G</figureCitation>
,
<figureCitation id="39E490EA062ABD435C1B194EABBB7F49" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27G</figureCitation>
) 2.8
<normalizedToken id="E28E73609FCCC7C620A02EEBF08CC9C5" originalValue="×">x</normalizedToken>
longer than wide. Gonoplac (Figs
<figureCitation id="C8E5FDAB9800846FA0F1AF6034E7DA8A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coarctana nigromedia sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79">21G</figureCitation>
,
<figureCitation id="7F1FEB1AE175846C206535F176120C2E" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27G</figureCitation>
) posterodorsal margin straight, about 4 tenths of blade length; external surface with dentiform cuticular projections; ventral margin broadly rounded; apex slightly tapered and rounded.
</paragraph>
<caption id="052F597D7FCD89D4F8B849E70C9B7076" doi="10.3897/asp.81.e81961.figure21" httpUri="https://binary.pensoft.net/fig/798174" pageId="0" pageNumber="79" start="Figure 21" startId="F21">
<paragraph id="89E756DD3640312232225749FFBCA1E9" pageId="0" pageNumber="79">
<emphasis id="C0BAF34271F21E38ACE6795C532BF821" bold="true" pageId="0" pageNumber="79">Figure 21.</emphasis>
<taxonomicName id="AF470B600AC3EEA8E7A96BB2FD060926" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana nigromedia" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigromedia">
<emphasis id="C57174E27F50531D87EF710BC51D52F6" italics="true" pageId="0" pageNumber="79">Coarctana nigromedia</emphasis>
</taxonomicName>
<emphasis id="155A2A3E7EAD3CEFDA71D462DE11C9C4" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, paratype female.
<emphasis id="459025F2632B42E312A1C03193418B03" bold="true" pageId="0" pageNumber="79">A</emphasis>
distal portion of abdomen, ventral view.
<emphasis id="7A10C3808B47EC4A121BD2EAC3877F93" bold="true" pageId="0" pageNumber="79">B</emphasis>
distal portion of abdomen, lateral view.
<emphasis id="52106EC1FF73158CC914EA80B4B60B5D" bold="true" pageId="0" pageNumber="79">C</emphasis>
first valvifer and first valvula, lateral view.
<emphasis id="81418C2FA241E806EDF0F0DF16E72CC5" bold="true" pageId="0" pageNumber="79">D</emphasis>
apical portion of first valvula.
<emphasis id="EEDC8CE757779560CBB243414268BC21" bold="true" pageId="0" pageNumber="79">E</emphasis>
second valvula, lateral view.
<emphasis id="E3147047535B03EF0F1205685F25187E" bold="true" pageId="0" pageNumber="79">F</emphasis>
apical portion of second valvula.
<emphasis id="4DF42EF929E6BC7D0B59802A789AEEB6" bold="true" pageId="0" pageNumber="79">G</emphasis>
second valvifer and gonoplac, lateral view.
<emphasis id="D0F36C919E104D2E3269600F2D669F34" bold="true" pageId="0" pageNumber="79">H</emphasis>
apical portion of gonoplac. Scale bars in mm.
</paragraph>
</caption>
<caption id="D2D144EEECDBF9921709A14A7A448EF6" doi="10.3897/asp.81.e81961.figure22" httpUri="https://binary.pensoft.net/fig/798175" pageId="0" pageNumber="79" start="Figure 22" startId="F22">
<paragraph id="5AB7961261736057C591C6059E0145DE" pageId="0" pageNumber="79">
<emphasis id="8969576D5A9EA6530E46B72891AA52FF" bold="true" pageId="0" pageNumber="79">Figure 22.</emphasis>
<taxonomicName id="CFDCFA5971471EDA70DD4F81D8DB4D1B" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana occultata" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="occultata">
<emphasis id="DD28B07DB3C2F2C70377121D16916AEB" italics="true" pageId="0" pageNumber="79">Coarctana occultata</emphasis>
</taxonomicName>
<emphasis id="D4044798DFF1E4B172EF20C86F1E4CE5" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="BEFB9E55B31097F8B90494DA11B4060A" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="3AB2A48391B4446E5C36E741024ACAFC" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="ECEE8652AD7802885020B8303E7CA979" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="E697D875187EE9300C8AA7630CE4CED0" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="DE7BAFE7856906EBDA24168BE0F5F4F7" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="D899FAEE0AA358AE3E024538957B7385" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="E8FB6B2FC4B902A8E102F9BE84B9D5B3" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="DF5FA1ED1BE272DB12801FC9314E2D90" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="4F3998D52105A043CFEE629FA75E5C0F" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="95BE72E4EECE4CCB0D3ECE4A79A82CA4" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="3614FA4E3BA00B7DD439F01D427D7283" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="21CF1179517FCF87B32DC45CDCCD4CCD" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="4A4C4664713CF40ADD86FBEE1DB95631" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
<caption id="A868BF1D3260E3B45E6979CEF1CBA637" doi="10.3897/asp.81.e81961.figure23" httpUri="https://binary.pensoft.net/fig/798176" pageId="0" pageNumber="79" start="Figure 23" startId="F23">
<paragraph id="25AC89203E7CFFE05345CA66B35FCBC6" pageId="0" pageNumber="79">
<emphasis id="92CB8E893AF4142CB7B376FBC34B377C" bold="true" pageId="0" pageNumber="79">Figure 23.</emphasis>
<taxonomicName id="510235E0EB37BCFFC0E9165A99404ACA" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="oricula">
<emphasis id="D7CA45F09D9EACF8AE676654A58F1EAE" italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis id="0E823686C4872C919042D7EEDAD5061C" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="AB7537C792A272A20293024A29F1CC0B" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="2E67A2F25C2D5130E36405FA168ED900" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="ED4567FA51FB3F7D7D78E035B3F33DBD" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="5CC70F8166AD845B45E60F51E6A4941C" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="72D4D3AF0A4AE70E4D68F527430A6B69" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="31249D9285CD35055CC1CA6EB11F3109" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="125F8147EFD7D511988904D30D4B6E95" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="5A2CBC2203B84BFDF31ACEC412E45623" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="170589883C7369C3256A552E390B5C57" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="FAC75C4D7592D1B361FF3C9BCED40F71" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="0D412BA8FBC8B343775A847A8FE5717B" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="A2488C2867CED473A1867C801F05DB46" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="EB0F3E31D5F381E4DC983560504FD4DE" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
<caption id="C4AFD9B3F1657C7DAF683E29C6B1E4FB" doi="10.3897/asp.81.e81961.figure24" httpUri="https://binary.pensoft.net/fig/798177" pageId="0" pageNumber="79" start="Figure 24" startId="F24">
<paragraph id="F0C1B4EB0D49941924580353F0F05F99" pageId="0" pageNumber="79">
<emphasis id="3879F96E2A27382FDE9267E0D5AD5D06" bold="true" pageId="0" pageNumber="79">Figure 24.</emphasis>
<taxonomicName id="AD8A71FEB37A95891918570EAF28B491" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="oricula">
<emphasis id="7F125A5044FC19A04B1347FB96FA45AE" italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis id="23545E3088F1A69C403E95F59FEF5F23" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, paratype female.
<emphasis id="21E5C7F98B7EA76CAD006EA01ED02561" bold="true" pageId="0" pageNumber="79">A</emphasis>
distal portion of abdomen, ventral view.
<emphasis id="A337BCED34227ECE9CA4496107CB9D2E" bold="true" pageId="0" pageNumber="79">B</emphasis>
distal portion of abdomen, lateral view.
<emphasis id="78F45A436C5556AB2F30E525EE62401F" bold="true" pageId="0" pageNumber="79">C</emphasis>
first valvifer and first valvula, lateral view.
<emphasis id="14396A97533ACF5F2CE17B769EC06352" bold="true" pageId="0" pageNumber="79">D</emphasis>
apical portion of first valvula.
<emphasis id="7F1D5F4A1E8B7BF050A2D0F7BB121652" bold="true" pageId="0" pageNumber="79">E</emphasis>
second valvula, lateral view.
<emphasis id="F65E791FEE83238B0A29E998667AC3CD" bold="true" pageId="0" pageNumber="79">F</emphasis>
apical portion of second valvula.
<emphasis id="30A9AC743319DD2FDB441892EF0FCD9F" bold="true" pageId="0" pageNumber="79">G</emphasis>
second valvifer and gonoplac, lateral view.
<emphasis id="CF4C3EA1F9110DBA864A6951649E4511" bold="true" pageId="0" pageNumber="79">H</emphasis>
apical portion of gonoplac. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6A4D7371F5C76BB05DCB0BD24E1D6F3D" pageId="0" pageNumber="79" type="distribution">
<paragraph id="282CE2B570B96F7327E084283BFEFCF1" pageId="0" pageNumber="79">Distribution.</paragraph>
<paragraph id="D7C9375B3A2801D731BEDA2BC9E5BC9D" pageId="0" pageNumber="79">Colombia, Bolivia, Brazil, Ecuador, Peru and Venezuela.</paragraph>
</subSubSection>
<subSubSection id="113C2A9CB659CE336271EB81BE511404" pageId="0" pageNumber="79" type="etymology">
<paragraph id="F89D8722A55E585C55DF4F7E5B57BECC" pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph id="DBF6E138B758DA3FCB6FF19B9F732638" pageId="0" pageNumber="79">
The generic name
<taxonomicName id="FCE377E0C16F8C6DCD18749EB32435A2" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="0BED7A5C2DA4F7F1FE03A4782A07257D" italics="true" pageId="0" pageNumber="79">Coarctana</emphasis>
</taxonomicName>
(feminine noun) is derived from the Latin word
<normalizedToken id="53D1845F4195742096C75DDBA9707C37" originalValue="“coarctatio”">&quot;coarctatio&quot;</normalizedToken>
meaning constriction. It refers to aedeagus bearing a strong constriction between the atrium and dorsal apodeme. The suffix - ana is common in names of
<taxonomicName id="4618BD0519B77FDA183CE8B95E502478" authorityName="Stal" authorityYear="1870" lsidName="" pageId="0" pageNumber="79" rank="tribe" tribe="Gyponini">Gyponini</taxonomicName>
genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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