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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.665.11792" ID-GBIF-Dataset="f49697cc-fc0c-4a14-bba1-edf1268f59f0" ID-PMC="PMC5523169" ID-Pensoft-Pub="1313-2970-665-93" ID-PubMed="28769629" ID-ZBK="4D5230AC67CB47ADAF2F051CA10D9724" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-665-93" ModsDocOrigin="ZooKeys 665" ModsDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1491378304537" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Wild, Michael, Sumoked, Bob &amp; Balke, Michael" docDate="2017" docId="0BD4201C606D2F01FF391BCEFEF5FF2A" docLanguage="en" docName="ZooKeys 665: 93-120" docOrigin="ZooKeys 665" docSource="http://dx.doi.org/10.3897/zookeys.665.11792" docTitle="Exocelina likui Shaverdo, Wild, Sumoked &amp; Balke, 2017, sp. n." docType="treatment" docUuid="3B1B6701-679A-4646-BCEF-8F09A5ECBF88" docUuidSource="ZooBank" docVersion="5" lastPageNumber="98" masterDocId="DC520217FF93FFF8FFDEFFCFFFA3777E" masterDocTitle="Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="120" masterPageNumber="93" pageNumber="98" updateTime="1668164355145" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wild, Michael</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sumoked, Bob</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>665</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>93</mods:start>
<mods:end>120</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.665.11792</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.665.11792</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-665-93</mods:identifier>
<mods:identifier type="ZBK">4D5230AC67CB47ADAF2F051CA10D9724</mods:identifier>
<mods:identifier type="ZooBank">4D5230AC67CB47ADAF2F051CA10D9724</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="128229140" LSID="urn:lsid:zoobank.org:act:3B1B6701-679A-4646-BCEF-8F09A5ECBF88" httpUri="http://treatment.plazi.org/id/0BD4201C606D2F01FF391BCEFEF5FF2A" lastPageNumber="98" pageId="5" pageNumber="98">
<subSubSection pageId="5" pageNumber="98" type="nomenclature">
<paragraph pageId="5" pageNumber="98">
5.
<taxonomicName LSID="http://zoobank.org/3B1B6701-679A-4646-BCEF-8F09A5ECBF88" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina likui" order="Coleoptera" pageId="5" pageNumber="98" phylum="Arthropoda" rank="species" species="likui">Exocelina likui</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="98">sp. n.</taxonomicNameLabel>
Figs 7, 15
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="type locality">
<paragraph pageId="5" pageNumber="98">Type locality.</paragraph>
<paragraph pageId="5" pageNumber="98">
Papua: Puncak Jaya Regency, south from Iratoi,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-3.3868055">03°23'12.5&quot;S</geoCoordinate>
;
<geoCoordinate direction="east" orientation="longitude" precision="1" value="137.24542">137°14'43.5&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="type material">
<paragraph pageId="5" pageNumber="98">Type material.</paragraph>
<paragraph pageId="5" pageNumber="98">Holotype: male &quot;Indonesia: Papua Province, S Iratoi, forest, 220m, 21.v.2015, -3,38095162063837, 137,311441982164, Sumoked &amp; Balke (Pap036)&quot;, &quot;M.Balke 6980&quot; [green text] (MZB). Paratypes: 1 male, 2 females with the same label as the holotype, one female with an additional label &quot;M.Balke 6981&quot; [green text] (NHMW, ZSM).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="diagnosis">
<paragraph pageId="5" pageNumber="98">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="98">Beetle small, oblong, dark brown to piceous, dorsal punctation dense and coarse, microreticulation distinctly impressed; pronotum without lateral bead; male antennae simple; male protarsomere 4 cylindrical, narrow, with large, thick, strongly curved anterolateral hook-like seta; median lobe slightly curved, with broadly pointed apex in lateral view and abruptly narrowed apically, with apex truncate in ventral view; paramere without dorsal notch, with subdistal setae very dense, strong, long and proximal setae very sparse, thin, small, weakly visible.</paragraph>
<paragraph pageId="5" pageNumber="98">
In shape of median lobe, the species resembles
<taxonomicName lsidName="E. takime" pageId="5" pageNumber="98" rank="species" species="takime">E. takime</taxonomicName>
(Balke, 1998) but distinctly differs from it in absence of the pronotal bead and strong dorsal sculpture. From
<taxonomicName lsidName="E. pui" pageId="5" pageNumber="98" rank="species" species="pui">E. pui</taxonomicName>
sp. n., it differs in dense and coarse dorsal punctation and in having medial lobe apically more pointed in lateral view, without lateral setae.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="description">
<paragraph pageId="5" pageNumber="98">Description.</paragraph>
<paragraph pageId="5" pageNumber="98">Size and shape: Beetle small (TL-H 3.2-3.6 mm, TL 3.8-4.0 mm, MW 1.85-1.95 mm), with oblong habitus, broadest at elytral middle. Coloration: Dorsally dark brown to piceous, with head and pronotum paler; head appendages and legs reddish brown distally (Fig. 7).</paragraph>
<paragraph pageId="5" pageNumber="98">Surface sculpture: Head with dense, coarse punctation (spaces between punctures 1-2 times size of punctures), finer anteriorly and posteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with slightly sparser punctation than head. Elytra with dense, coarse punctation, coarser than on pronotum. Pronotum and elytra with distinctly impressed microreticulation, dorsal surface submatt. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, with sparse but distinct punctation, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and sparse, coarse punctation, especially on four last abdominal ventrites.</paragraph>
<paragraph pageId="5" pageNumber="98">Structures: Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, smooth, broadly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, very slightly convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded apically.</paragraph>
<paragraph pageId="5" pageNumber="98">
Male: Antenna simple (Fig. 7). Pro- and mesotarsomeres 1-3 not dilated. Protarsomere 4 cylindrical, narrow, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 40 and posterior row of 14 relatively long, not pointed setae (Fig. 15D). Median lobe slightly curved, with broadly pointed apex in lateral view, and abruptly narrowed apically, with apex slightly truncate in ventral view. Paramere without dorsal notch, with subdistal setae very dense, strong, long and proximal setae very sparse, thin, small, weakly visible. (Fig. 15
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Abdominal ventrite 6 broadly rounded apically, 5-6 lateral strioles on each side.
</paragraph>
<paragraph pageId="5" pageNumber="98">Holotype: TL-H 3.5 mm, TL 3.9 mm, MW 1.9 mm.</paragraph>
<paragraph pageId="5" pageNumber="98">Female: Pro- and mesotarsi not modified. Abdominal ventrite 6 without lateral strioles.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="distribution">
<paragraph pageId="5" pageNumber="98">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="98">Papua: Puncak Jaya Regency. The species is known only from the type locality (Fig. 19).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="98" type="etymology">
<paragraph pageId="5" pageNumber="98">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="98">
The species is named after Michael
<normalizedToken originalValue="Wilds">Wild's</normalizedToken>
best Wano friend Liku who grew up at Iratoi and hunted many times in the area where the species was collected. The name is a noun in the genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>