treatments-xml/data/0B/9F/97/0B9F97DA0E9E534588D429E2AF212FBE.xml

<document ID-CLB-Dataset="24048" ID-DOI="http://dx.doi.org/10.3897/zookeys.911.38496" ID-GBIF-Dataset="928675b3-f1d3-48d9-9e7b-bb1a7dc2e4ca" ID-PMC="PMC7031396" ID-Pensoft-Pub="1313-2970-911-1" ID-Pensoft-UUID="4B99B71032185BE68A8FAC14668BA5D9" ID-PubMed="32104136" ID-ZooBank="0E3064E46D1E444CAF818BDA7DC4306B" ModsDocID="1313-2970-911-1" checkinTime="1581612310290" checkinUser="pensoft" docAuthor="Sanoamuang, La-orsri &amp; Watiroyram, Santi" docDate="2020" docId="0B9F97DA0E9E534588D429E2AF212FBE" docLanguage="en" docName="ZooKeys 911: 1-20" docOrigin="ZooKeys 911" docSource="http://dx.doi.org/10.3897/zookeys.911.38496" docTitle="Phyllodiaptomus roietensis Sanoamuang &amp; Watiroyram 2020, sp. nov." docType="treatment" docUuid="59131C6D-A0DE-4BE0-9383-20C0DA8A709D" docUuidSource="ZooBank" docVersion="6" id="4B99B71032185BE68A8FAC14668BA5D9" lastPageNumber="1" masterDocId="4B99B71032185BE68A8FAC14668BA5D9" masterDocTitle="Phyllodiaptomus (Phyllodiaptomus) roietensis, a new diaptomid copepod (Copepoda, Calanoida) from temporary waters in Thailand and Cambodia, with a key to the species" masterLastPageNumber="20" masterPageNumber="1" pageNumber="1" updateTime="1732940226412" updateUser="ExternalLinkService">
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<mods:title id="DFF9F4856EC22A5D2F4821AD05EF3ECD">Phyllodiaptomus (Phyllodiaptomus) roietensis, a new diaptomid copepod (Copepoda, Calanoida) from temporary waters in Thailand and Cambodia, with a key to the species</mods:title>
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<mods:namePart id="CF8F287F9ED8F9F33F1225E8E776BB09">Sanoamuang, La-orsri</mods:namePart>
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<mods:namePart id="B07C4C627B3A930C12E89D81AC3DE0E4">Watiroyram, Santi</mods:namePart>
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<mods:date id="595DAFC407C624512DCCB6954CD00238">2020</mods:date>
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<treatment id="0B9F97DA0E9E534588D429E2AF212FBE" ID-GBIF-Taxon="162136180" LSID="urn:lsid:zoobank.org:act:59131C6D-A0DE-4BE0-9383-20C0DA8A709D" httpUri="http://treatment.plazi.org/id/0B9F97DA0E9E534588D429E2AF212FBE" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Copepoda" scope_family="Diaptomidae" scope_order="Calanoida" scope_subFamily="Diaptominae">
<subSubSection id="AEE4BF61229CD9C3A48C1943FC326979" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="65352F4897578C8059F5B191A413CC4A" pageId="0" pageNumber="1">
<taxonomicName id="69555525C7AF1C5A0C4754CAE38F108D" ID-CoL="7ZGN2" LSID="0B9F97DA-0E9E-5345-88D4-29E2AF212FBE" authority="Sanoamuang &amp; Watiroyram, 2020" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis" status="sp. nov.">Phyllodiaptomus (P.) roietensis</taxonomicName>
<taxonomicNameLabel id="293217ECC18D8E7B77D603BBDD0B3B49" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="4547CB12BA4C6FD38D9274FB04FAE2B6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation id="AE25B4D923661CE5DA622FB85A792C61" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="547D9C50BBDCA273C599A4536A47D796" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation id="84ED918A4CF4657A64CFDFAD4A681637" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllodiaptomus (P.) roietensis sp. nov., female: A P 1 B P 2 C P 3 D P 4 E P 5 F, G right and left P 5 Exp- 2 (black arrows indicate longtitudinal ridges) A-E posterior view F, G anterior view." figureDoi="10.3897/zookeys.911.38496.figure5" httpUri="https://binary.pensoft.net/fig/379465" pageId="0" pageNumber="1">, 5</figureCitation>
<figureCitation id="42E8692D03C10CA23FBCB940EED5274E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">, 6</figureCitation>
<figureCitation id="1E7B0510AF27FC3DE9DDD8F7F617D0CE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">, 7</figureCitation>
<figureCitation id="F312D94EC1EA8E9A85744954FBE77C59" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0252D9909F9CE8B15CCA8AB85B3F953E" pageId="0" pageNumber="1" type="type locality">
<paragraph id="4BA4FE790A668964BFC324D84C94A804" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="D609DF637EF977E03D2083C37807AE29" pageId="0" pageNumber="1">
A pool in the rice field at Ban Nakae, Khilek Subdistrict, Pathum Rat District, Roi Et Province, northeastern Thailand; pH of water 8.6, water conductivity 126 
<normalizedToken id="96C01AE004C61164B697E07AA42D313A" originalValue="µS">µS</normalizedToken>
cm-1.
</paragraph>
</subSubSection>
<subSubSection id="DF5270E2EB3D43D2200F85F466D8E2FA" pageId="0" pageNumber="1" type="type material">
<paragraph id="88DED4C37379E086D3D98152BA6A789D" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="A75D4C70E902C21B70D0AD2BB29286B6" pageId="0" pageNumber="1">
<emphasis id="D062CAF62CBE8A623ED5CF7A5814964F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: one adult male completely dissected (NHMUK 2019.7, one slide), Ban Nakae (
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, 
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), Khilek Subdistrict, Pathum Rat District, Roi Et Province, northeastern Thailand; collected on 12 June 1999 by L. Sanoamuang. 
<emphasis id="4349E0124068C64BADB31F3F2B0A013C" bold="true" italics="true" pageId="0" pageNumber="1">Allotype</emphasis>
: one adult female completely dissected (NHMUK 2019.8, one slide); same data as for holotype. 
<emphasis id="60F80C665694752873BF38A9A75F245D" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: two adult females and three adult males undissected preserved in 70% ethanol (NHMUK 2019.9-13), one adult female completely dissected (KKU-COP-2019-S-01); one adult female with eggs and three adult males undissected preserved in 70% ethanol (KKU-COP-2019-T-01); same data as for holotype.
</paragraph>
</subSubSection>
<subSubSection id="8DA9BC159AC49B836639FB0B44029A08" pageId="0" pageNumber="1" type="other localities">
<paragraph id="41AD29C83273E59BB3DF8DF7F4073E36" pageId="0" pageNumber="1">Other localities.</paragraph>
<paragraph id="490D2E92F6FF2542270437BF61B80A68" pageId="0" pageNumber="1">
(1) a temporary pond, Ban Non Lakki (
<geoCoordinate id="32FC16D944401D79F6532EB772139ADE" degrees="15" direction="north" minutes="10" orientation="latitude" precision="15" seconds="55" value="15.181945">15°10'55&quot;N</geoCoordinate>
, 
<geoCoordinate id="75A5C108060AB72AC437BD5D3D6C6EC3" degrees="102" direction="east" minutes="23" orientation="longitude" precision="15" seconds="46" value="102.39611">102°23'46&quot;E</geoCoordinate>
), Than Lalot Subdistrict, Phimai District, Nakhon Ratchasima Province, northeastern Thailand; collected on 17 October 2017 by N. Plangklang; (2) a roadside canal, Tropeang Chouk village (no geographical co-ordinates), Baray District, Kampong Thom Province, central Cambodia; collected on 14 June 2007 by R. Chaicharoen; (3) a temporary pond, Kropeu village (no geographical co-ordinates), Steung Sen District, Kampong Thom Province, central Cambodia; collected on 14 June 2007 by R. Chaicharoen.
</paragraph>
</subSubSection>
<subSubSection id="B683041D3BE068D971855FC55A3339C5" pageId="0" pageNumber="1" type="description">
<paragraph id="0599FEFBBB91A60798FAD205C257BB21" pageId="0" pageNumber="1">Description of adult female.</paragraph>
<paragraph id="45A35E30B8107D67EE1BD918F4C12C2C" pageId="0" pageNumber="1">
Total body length measured from anterior margin of rostrum to posterior margin of caudal rami, 0.9-1.3 mm. Rostrum (Fig. 
<figureCitation id="4E4E9EDD9FAECCE0E0D5F3ACE2C3CF5B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3G</figureCitation>
) with bifid process in distal margin, pointed backward; each with short spine at tip. Prosome length: urosome plus caudal rami ratio about 2.6:1, ratio of width to length of prosome = 1:2.4, urosomites 1-3 = 1.3:3.0:1.1, caudal ramus = 1:1.5. Prosome (Figs 
<figureCitation id="D55F28E5DE708B6F156D893BA0F1CA00" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2A</figureCitation>
, 
<figureCitation id="E73B893DB111A940EC47D3B16CED3DEF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3A</figureCitation>
) ovoid, cephalosome with transversal groove in anterior part of somite length; Pdg 4 and 5 fused, partly separated laterally, with few tiny hair-like spinules scattered laterally (Fig. 
<figureCitation id="1EB8D66D0FFD0940C0866E19DC25066D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3B, C</figureCitation>
). Pdg 5 (Figs 
<figureCitation id="6D9670F4C71BEA46484EE30E8B038C75" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2C, E</figureCitation>
, 
<figureCitation id="7401157ADE5F13F642D15E1D671CC8C1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3C-F</figureCitation>
) with asymmetrical postero-lateral wings; right one rounded; left one longer and triangular; each wing with dorsal and posterior spines (former spine slightly larger than later one). Urosome (Figs 
<figureCitation id="FAD7430B2E7BA1F3497D33539A623438" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2A</figureCitation>
, 
<figureCitation id="21C04BBF5C3656252ADD4896306F19A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3A</figureCitation>
) with asymmetrical genital double-somite. Genital double-somite (Figs 
<figureCitation id="373902967C2DF101AF128C91CA6875C1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2C-E</figureCitation>
, 
<figureCitation id="FE1CDD6650142D2B06FA8F5619045534" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3C-F</figureCitation>
) longer than urosomite 2, anal somite and caudal ramus combined. Left side with obviously laterally dilated proximal part of genital-double segment; dilatation dorsally with large and blunt spine distally, tip of spine oriented medially. Right side with slightly dilated proximal part of genital double-somite; elongated into triangular outgrowth with blunt spine at tip; spine orientated ventro-laterally. A pair of gonopores located beneath genital operculum, at about one-half length of genital double-segment. Adult female bears one egg sac with 20-25 eggs (Fig. 
<figureCitation id="595B8E1465B1A1E6705900F65FC569C9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2A</figureCitation>
). Urosomite 2 symmetrical, very short. Anal somite (Figs 
<figureCitation id="F3E28A925F2EFE1CCAAC52A3F76F6316" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2C, D</figureCitation>
, 
<figureCitation id="7FE8B74A5376C3F9F16AF741A138708A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B Pdg 4, lateral view C Pdg 5 and genital double-somite, dorsal view D Pdg 5 and genital double-somite spines, lateral view from left side E Pdg 5 and genital double-somite spines, lateral view from right side F Pdg 5 and genital double-somite, lateral view G rostral spines." figureDoi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1">3A</figureCitation>
) as long as wide; anal operculum small, free margin convex. Caudal rami (Fig. 
<figureCitation id="96186BE584C0EB5FB1EDC85B7F53354A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phyllodiaptomus (P.) roietensis sp. nov., female: A habitus, dorsal view B cephalosome with rostrum, lateral view C lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view D urosome, ventral view E Pdg 5 with P 5 and genital double-somite, lateral view from left side." figureDoi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1">2A, D</figureCitation>
) symmetrical, with row of setules along inner and outer margins. Ramus with six setae (seta II-VII), subequal in length, all plumose but dorsal (VII); dorsal seta articulated, longest.
</paragraph>
<caption id="092F57006B255BE0BF49F769F1A60F05" doi="10.3897/zookeys.911.38496.figure2" httpUri="https://binary.pensoft.net/fig/379462" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="125B56AFA27797B81C36E4EAB2C36080" pageId="0" pageNumber="1">
<emphasis id="15D54353CA0F5D341AE93119E451157E" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName id="CEC0B4177BA504F631C9707ABAFA5457" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., female: 
<emphasis id="CA04E234C063F3E923237D337E1B3AB7" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view 
<emphasis id="722F82E54D6C21403B09D4D9F357E592" bold="true" pageId="0" pageNumber="1">B</emphasis>
cephalosome with rostrum, lateral view 
<emphasis id="2687848B71D13E3D056CEF0962ABE5E1" bold="true" pageId="0" pageNumber="1">C</emphasis>
lateral wings on Pdg 5 and urosome (without caudal rami), dorsal view 
<emphasis id="7FA244E30B796A96FDB92A4AD8A52AE5" bold="true" pageId="0" pageNumber="1">D</emphasis>
urosome, ventral view 
<emphasis id="3CAF0DB8363F0049165CD58301F1FE72" bold="true" pageId="0" pageNumber="1">E</emphasis>
Pdg 5 with P5 and genital double-somite, lateral view from left side.
</paragraph>
</caption>
<caption id="5857AF72D724CDD77C4A3AF30EAAB543" doi="10.3897/zookeys.911.38496.figure3" httpUri="https://binary.pensoft.net/fig/379463" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="D12C016EAA6DCC49493645F948D87B03" pageId="0" pageNumber="1">
<emphasis id="1DBA88325B7F489F63924FE8C672E321" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="D724EC9B5071B3C089FCC63A39B82CD8" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., female: 
<emphasis id="93150781376836121A0D074DC29E2D2A" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view 
<emphasis id="C1DB5AC0FD8817A984DF77B91A63D1AB" bold="true" pageId="0" pageNumber="1">B</emphasis>
Pdg 4, lateral view 
<emphasis id="DE8A94C5B8522ADBCAC03B16D70F8208" bold="true" pageId="0" pageNumber="1">C</emphasis>
Pdg 5 and genital double-somite, dorsal view 
<emphasis id="02C51A595B4897BA5C119D6743FB3E7F" bold="true" pageId="0" pageNumber="1">D</emphasis>
Pdg 5 and genital double-somite spines, lateral view from left side 
<emphasis id="663FEEEB5B1912A36475D22510A69C38" bold="true" pageId="0" pageNumber="1">E</emphasis>
Pdg 5 and genital double-somite spines, lateral view from right side 
<emphasis id="D9F1BA65C65374E48183338CC57AA847" bold="true" pageId="0" pageNumber="1">F</emphasis>
Pdg 5 and genital double-somite, lateral view 
<emphasis id="6A530BC189EA58A96A7AD9F7CBF72C33" bold="true" pageId="0" pageNumber="1">G</emphasis>
rostral spines.
</paragraph>
</caption>
<paragraph id="46DAD2FB8B7D7F4410C03F1146AA491F" pageId="0" pageNumber="1">
Antennule (Fig. 
<figureCitation id="DA22F169F249E6F4224F88F558952100" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4A</figureCitation>
) symmetrical, 25-segmented, reaching beyond the end of caudal setae. Setal formula (Roman numerals in parentheses refer to segment number): 1+ae (I), 3+ae (II), 1+ae (III), 1(IV), 1+ae (V), 1 (VI), 1+ae (VII), 1+sp (VIII), 2+ae (IX), 1 (X), 1 (XI), 1+ae+sp (XII), 1 (XIII), 1+ae (XIV), 1 (XV), 1+ae (XVI), 1 (XVII), 1 (XVIII), 1+ae (XIX), 1 (XX), 1 (XXI), 2 (XXII), 2 (XXIII), 2 (XXIV), 4+ae (XXV).
</paragraph>
<paragraph id="3399EB934F241BD32071C98906FC0F83" pageId="0" pageNumber="1">
Antenna (Fig. 
<figureCitation id="80893C1653047D846E52F6A493BAC675" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4B</figureCitation>
) biramous. Coxa and basis with one and two inner setae on distal corner, respectively. Exp-1-7 with 1, 3, 1, 1, 1, 1, and 1 inner seta, respectively; Exp-7 with three additional apical setae. Enp-1 with two inner median setae. Enp-2 with eight inner and seven apical setae.
</paragraph>
<paragraph id="57D283286ADAE9CAD4589FDD09857BE6" pageId="0" pageNumber="1">
Mandible (Fig. 
<figureCitation id="08BED2CFE6D37A2AF6476F09B6DD3A5A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4C</figureCitation>
) with six strongly chitinized teeth and one dorsal seta on gnathobase. Basis with four inner setae. Enp-1 with four inner distal setae; Enp-2 with nine apical setae plus tiny spinules along posterior surface. Exp-1-4 with 1, 1, 1, 3 setae, respectively.
</paragraph>
<paragraph id="F2B5A3DC4A41C92D239C50BC7F714DA8" pageId="0" pageNumber="1">
Maxillule (Fig. 
<figureCitation id="28BE943900B58200BAADBA633F44B979" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4D</figureCitation>
) with 13 setae on praecoxal arthrite. Coxal endite and coxal epipodite with four and nine setae, respectively. Proximal and distal endites each with four setae; basal exite with a single seta. Enp with seven apical setae. Exp with six setae.
</paragraph>
<paragraph id="4917EB25FC20D371106EE3CBF5EAA075" pageId="0" pageNumber="1">
Maxilla (Fig. 
<figureCitation id="7E4B0462AEF6BD8A6FF1F2191383858E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4E</figureCitation>
) with two praecoxal and two coxal endites; each with three apical setae. Allobasis with three setae. Enp-1 and 2 with three setae each.
</paragraph>
<paragraph id="14962F796CA817534DFF029C0C755B05" pageId="0" pageNumber="1">
Maxilliped (Fig. 
<figureCitation id="EB7D448F79F798763E66BBF44BF16512" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phyllodiaptomus (P.) roietensis sp. nov., female: A antennule B antenna C mandible D maxillule E maxilla F maxilliped." figureDoi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1">4F</figureCitation>
) with four endites on syncoxa, with 1, 2, 3, 4 apical setae respectively. Basis with three setae along median margin. Enp-1-6 with 2, 3, 2, 2, 2, 4 setae, respectively.
</paragraph>
<caption id="7FF2DE10939D3602C424049B430FFDFE" doi="10.3897/zookeys.911.38496.figure4" httpUri="https://binary.pensoft.net/fig/379464" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="50DE2C7CFDF128C991CB8C6CB8C21042" pageId="0" pageNumber="1">
<emphasis id="13F6DEBC73A3D6170D37CD9A5CEBB3D1" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="3B7F29180C790B0675A99258D2ED8C5A" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., female: 
<emphasis id="03E574280663329861CB38AEB5A57AD6" bold="true" pageId="0" pageNumber="1">A</emphasis>
antennule 
<emphasis id="76A69C47AE698D7A3520F94D143735F2" bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna 
<emphasis id="444EC8D2943C142F4EF8CE717BEC875D" bold="true" pageId="0" pageNumber="1">C</emphasis>
mandible 
<emphasis id="03FF75B6CA0FBD65A260B8019DF66666" bold="true" pageId="0" pageNumber="1">D</emphasis>
maxillule 
<emphasis id="DB769A14FD2116DF7627C2FD47306297" bold="true" pageId="0" pageNumber="1">E</emphasis>
maxilla 
<emphasis id="5257F694631843682E9EFD297166AC67" bold="true" pageId="0" pageNumber="1">F</emphasis>
maxilliped.
</paragraph>
</caption>
<paragraph id="309E1BBFCE02D9898AC31D4A4B034037" pageId="0" pageNumber="1">
P1-P4 (Fig. 
<figureCitation id="66F2B999A54C2487452012514E120B18" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllodiaptomus (P.) roietensis sp. nov., female: A P 1 B P 2 C P 3 D P 4 E P 5 F, G right and left P 5 Exp- 2 (black arrows indicate longtitudinal ridges) A-E posterior view F, G anterior view." figureDoi="10.3897/zookeys.911.38496.figure5" httpUri="https://binary.pensoft.net/fig/379465" pageId="0" pageNumber="1">5A-D</figureCitation>
) with round and bare intercoxal sclerite. Coxa with inner seta. P1 basis with reduced outer seta. Exp longer than Enp, Exp and Enp three-segmented except P1 Enp bi-segmented. Armature formula of P1-P4 as follows (Arabic and Roman numerals indicate number of setae and spines, respectively; outer-inner or outer-apical-inner indicate seta/spine):
</paragraph>
<paragraph id="5CD3FDE641323759F9F7E40AF2B5735D" pageId="0" pageNumber="1">
<table id="C5E042CE8D7AB7B20DA05013247E4905" pageId="0" pageNumber="1">
<tr id="F6EA9B5FE2C8E7F8BC8677B27402E3CF" pageId="0" pageNumber="1">
<th id="23B8DF0FFEECFD545F2EC4295ED92ABB" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="2">-</th>
<th id="6A9ACDF46032B4E8B07FF18128248034" colspan="1" pageId="0" pageNumber="1" rowspan="2">Coxa</th>
<th id="6E4E07344C4A4836D78DA15282B8AB4A" colspan="1" pageId="0" pageNumber="1" rowspan="2">Basis</th>
<th id="EC546C8E8343C20C5A5FA907BA66B787" colspan="3" pageId="0" pageNumber="1" rowspan="1">Exp</th>
<th id="EF2F08188C4A56ED0E4EA90F8E285D33" colspan="3" pageId="0" pageNumber="1" rowspan="1">Enp</th>
</tr>
<tr id="0E8388EDB4FF06922C637FE89D826380" pageId="0" pageNumber="1">
<th id="15939EBCD1727CC4AFCB2E210DC8D513" colspan="1" pageId="0" pageNumber="1" rowspan="1">1</th>
<th id="3DDFC870C201EE100DB790993720C974" colspan="1" pageId="0" pageNumber="1" rowspan="1">2</th>
<th id="68D1BE5663B3FB14274AEEA57C7D785D" colspan="1" pageId="0" pageNumber="1" rowspan="1">3</th>
<th id="68BC7C83D6CC7B730F61B9CCECF945F8" colspan="1" pageId="0" pageNumber="1" rowspan="1">1</th>
<th id="76A2165EF6B06F9EDF67054DF02B7982" colspan="1" pageId="0" pageNumber="1" rowspan="1">2</th>
<th id="677BE88975523F0D5F722BAB0A7759DD" colspan="1" pageId="0" pageNumber="1" rowspan="1">3</th>
</tr>
<tr id="10F8729B4BC5D68FB758D0D7B9707E09" pageId="0" pageNumber="1">
<td id="93011E5118B130CBB793EFD183C33996" colspan="1" pageId="0" pageNumber="1" rowspan="1">P1</td>
<td id="E85F7E3A94B9512489FFEAC69B4C2F80" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="36501A077C1EBF8AC3E5A508DB474002" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td id="D000B2E52C2321F7E3D386697FBD385D" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="C82506B3BA06A6983F31E7CFAEA5E59A" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="382B7CDC788B8FEB449DD11ACE526DBC" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-3-2</td>
<td id="19B9D432B86B5A4EC171209BB0261CFE" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="AC7FDD170F37A0BF8A0B2CC9C9AE26FD" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-2-3</td>
<td id="E78A87BC97681CDA7DACB0E6119435F0" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
<tr id="629E12676BB3DC4560125DC899050453" pageId="0" pageNumber="1">
<td id="793FEFE516EF0C226C1FB7FBA591A7A7" colspan="1" pageId="0" pageNumber="1" rowspan="1">P2</td>
<td id="267C12A3D1048937D42904DEE286A527" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="3D9CA52728C1B9D49DC9E88B3C5E6391" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td id="313EDCCB299914BC6852FE961BBE0F6E" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="8F907FF523901E92B637280001159171" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="B7ADED41FAEA9586430280DEF4EBC8D8" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-3-3</td>
<td id="114BED474C5AF94CC8BAF09CDC59D5E6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="3D3E5CF5C1CA9B9A58CD0FDE844AE2E8" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-2</td>
<td id="4A7BCAB0AAF6EBB215DBAF5521798452" colspan="1" pageId="0" pageNumber="1" rowspan="1">2-2-3</td>
</tr>
<tr id="F8AE74A14C73BDB7ACF5A31D164F7AEE" pageId="0" pageNumber="1">
<td id="7685B2FA5EDBE4D2663796750B095D29" colspan="1" pageId="0" pageNumber="1" rowspan="1">P3</td>
<td id="D733BD03E73F956705C693E2C7DB6EEB" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="AFEBE442E911AADBFFA09E08CBEAB9A0" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-0</td>
<td id="EB3ED51B711D1AB1B1C42DDF4F5C484B" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="E5AF289B205E94FD719BAAFF39CC45C5" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="53D49772ACAE906D3AB2FB1E99EC9D96" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-3-3</td>
<td id="47DF6A32C4A4C673B62B16215621A9F6" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="E977576063247240236C242E215B6BF3" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-2</td>
<td id="1821DBF7E6727BAE63FA0BBF3A83C4BE" colspan="1" pageId="0" pageNumber="1" rowspan="1">2-2-3</td>
</tr>
<tr id="3DAB3F572DCB15390D8F7D9C2030299D" pageId="0" pageNumber="1">
<td id="0B65DFCD97CE3AA8EEC87D2C2D905616" colspan="1" pageId="0" pageNumber="1" rowspan="1">P4</td>
<td id="BA071931FB4AEC14E4011BA708A19233" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="A99D13399B2AF755333E9CF1E53A3D0B" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td id="EBDA0989A598A01EE73E8541E3796DED" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="58942B9B05977D3ACB061ED33B8BA8C8" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1</td>
<td id="496C070B2DA36FC1B888DF4C7BF8D4BE" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-3-3</td>
<td id="FECE8AE20924AF2FE641B490DAD8DE00" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="FDCA16D871AF83B4C0DEA9237DA2AE4C" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-2</td>
<td id="6B86B5CF5C766E37A74E806BA67271C0" colspan="1" pageId="0" pageNumber="1" rowspan="1">2-2-3</td>
</tr>
</table>
</paragraph>
<paragraph id="882D96DE73602BC890D0E35DE9076FD5" pageId="0" pageNumber="1">
P5 (Fig. 
<figureCitation id="F0BA14B5F493388B3DE47670E4C21AA8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllodiaptomus (P.) roietensis sp. nov., female: A P 1 B P 2 C P 3 D P 4 E P 5 F, G right and left P 5 Exp- 2 (black arrows indicate longtitudinal ridges) A-E posterior view F, G anterior view." figureDoi="10.3897/zookeys.911.38496.figure5" httpUri="https://binary.pensoft.net/fig/379465" pageId="0" pageNumber="1">5E</figureCitation>
) asymmetrical. Coxa with blunt, stout spine on distal outer margin. Basis with thin, bare seta on distal outer margin, reaching beyond 3/4 of Exp-1. Exp-1 sub-rectangular, more than twice as long as wide, longer than Enp. Exp-2 triangular, right side stout and shorter than left one; with row of strong spinules along margins and proximolateral spine at basal Exp-3; with two longitudinal grooves on anterior view (Fig. 
<figureCitation id="3BEF1F3F828734611625B9DC847958FA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllodiaptomus (P.) roietensis sp. nov., female: A P 1 B P 2 C P 3 D P 4 E P 5 F, G right and left P 5 Exp- 2 (black arrows indicate longtitudinal ridges) A-E posterior view F, G anterior view." figureDoi="10.3897/zookeys.911.38496.figure5" httpUri="https://binary.pensoft.net/fig/379465" pageId="0" pageNumber="1">5F, G</figureCitation>
). Exp-3 represented by small distal prominence produced into short distolateral spine and longer medial spine. Enp subconical, Enp-1 slightly rectangular. Enp-2 tipped with circular row of spinules.
</paragraph>
<caption id="EE17C3BAE195C590C44F0A202DBFB064" doi="10.3897/zookeys.911.38496.figure5" httpUri="https://binary.pensoft.net/fig/379465" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="DC50A7BC05DFD1BE3E81E5886FDD797F" pageId="0" pageNumber="1">
<emphasis id="BD4797AF814C850BF32DA149718B7A30" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName id="5D7ABE13B8C67CD10CBC5ED831ADC157" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., female: 
<emphasis id="3317BF39AF821B825096C4A947B1D381" bold="true" pageId="0" pageNumber="1">A</emphasis>
P1 
<emphasis id="1A98914D75390ED28A15F2432CA16993" bold="true" pageId="0" pageNumber="1">B</emphasis>
P2 
<emphasis id="493727B1DAF239A6C10F43042AD4CCC2" bold="true" pageId="0" pageNumber="1">C</emphasis>
P3 
<emphasis id="5BE635C86B481028BB1118C1BECF7462" bold="true" pageId="0" pageNumber="1">D</emphasis>
P4 
<emphasis id="B6A9ED527393493F254CDC5BB3AB40F9" bold="true" pageId="0" pageNumber="1">E</emphasis>
P5 
<emphasis id="97FCAA2E36D1AAC972EF2F6410B1CFD7" bold="true" pageId="0" pageNumber="1">F, G</emphasis>
right and left P5 Exp-2 (black arrows indicate longtitudinal ridges) 
<emphasis id="755F55DEDEDA9F6CC91B994EEB3912FA" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
posterior view 
<emphasis id="DC48D676BAADDDF86D4F4CFB8E3FB365" bold="true" pageId="0" pageNumber="1">F, G</emphasis>
anterior view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7FFDE0F5A7D7A8A42C77ABC06C5EA642" pageId="0" pageNumber="1" type="description">
<paragraph id="F1FBD3297C53F0A619DBFB327C054780" pageId="0" pageNumber="1">Description of adult male.</paragraph>
<paragraph id="C002CD9FE54EF8E7D6A15B42F1283EC0" pageId="0" pageNumber="1">
Body length (Figs 
<figureCitation id="57E0224C76B2E8D84BAF6A89C2CC6DBC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6</figureCitation>
, 
<figureCitation id="1E8B50D996801EA2610A2ECB0E112986" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7A</figureCitation>
) without caudal setae, 0.8-1.1 mm (mean = 1.0 mm, n = 5), smaller than female. Prosome length: urosome plus caudal rami ratio about 2.1:1, ratio of width to length of prosome = 1:2.1, urosomites 1-5 = 2.3:1.0:1.0:1.3:1.0, caudal ramus = 1:1.9. Prosome similar to that of female except lateral wings on Pdg 5. Lateral wings (Figs 
<figureCitation id="9D4335D3862FF63FC70EC80733B70A49" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6A</figureCitation>
, 
<figureCitation id="84A3C6220BB2293057152CAD8035AF5C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7B-D</figureCitation>
) asymmetrical, round on right and more triangular on left side; posterior spine on right wing larger compared to left side. Urosome (Figs 
<figureCitation id="8AF5AC8922E3E0E5EA6CADB949D3EFE0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6A, B</figureCitation>
, 
<figureCitation id="D82E630EAF5B13FFC7AB49D8B37C1237" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7A</figureCitation>
) 5-segmented, asymmetrical, curved downward to right side. Genital somite (Figs 
<figureCitation id="C758EE8F5E0ACD0AEFCEE6E9AE17CF04" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6</figureCitation>
, 
<figureCitation id="426236E2C510875AA5A6615465B61691" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7B</figureCitation>
) dilated postero-laterally on right side, with spine at distal outer corner; longer than that on Pdg 5 wings. Genital aperture located on mid-ventral region. Urosomites 2-3 (Figs 
<figureCitation id="E289124E57A5E51C837E012B02DDC9B5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6B</figureCitation>
, 
<figureCitation id="4CDB7F2BF31D9FD7540C4D098BAFD9AE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7D</figureCitation>
) without ornamentation. Urosomite 4 (Fig. 
<figureCitation id="F79432FBA14DBADCAD0F6319BBCE8C7E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6A, B</figureCitation>
) with irregularly dilated posterior margin. Anal somite (Fig. 
<figureCitation id="A23A9A5DABDAAD6CFB21F8C4DD30F5BB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6A, B</figureCitation>
) asymmetrical and twisted to right side. Caudal rami (Figs 
<figureCitation id="4FC9A1924343709C47B9E1E507400DEA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6A, B</figureCitation>
, 
<figureCitation id="7F5F217ECBAA1B22834A888BACFF34D3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7E, F</figureCitation>
) asymmetrical, right ramus with two triangular prominences: one proximolateral and one distoventral; setation similar to female.
</paragraph>
<paragraph id="B51A996358A9FBAC87090EA60E0C555F" pageId="0" pageNumber="1">
Antennule (Figs 
<figureCitation id="2237F1C300B9F06D52C8EB26F1C353F2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6C</figureCitation>
, 
<figureCitation id="6091611477F910E444E2B4670ED1EA5A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B Pdg 5 and genital somite, dorsal view C Pdg 5 lateral wing, left view D Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view E, F right caudal ramus in dorsal (E) and ventral (F) views G, H right antennule (white arrows indicate spines on segments 8, 10 - 18, 20." figureDoi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1">7G, H</figureCitation>
) asymmetrical, with geniculated right side. Right antennule 22-segmented, with setal formula as 1+ae (I), 3+ae (II), 1+ae (III), 1 (IV), 1+ae (V), 1 (VI), 1+ae (VII), 1+sp (VIII), 2+ae (IX), 1+sp (X), 1+sp (XI), 1+ae+sp (XII), 1+ae+sp (XIII), 2+ae+sp (XIV), 2+ae+sp (XV), 2+ae+sp (XVI), 1+sp (XVII), 1+sp (XVIII), 2+ae+sp (XIX), 3+sp (XX), 2 (XXI), 4+ae (XXII); geniculated between segments 18 and 19; segment 20 (antepenultimate) with serrated process distally (3-4 teeth), and with longitudinal hyaline membrane along outer margin.
</paragraph>
<caption id="B1D801345CA78C5515D6400119B2CD0F" doi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph id="C3172F44D1629D302B4171F5A4EF96BA" pageId="0" pageNumber="1">
<emphasis id="E5B489A75655015AC428470EDC8F2D2E" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
<taxonomicName id="1CDCF94B86FD651C60CD0AB67186196F" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., male: 
<emphasis id="2946FE59E889E64EE9735B774628AC85" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view 
<emphasis id="33C2AEDFC8EF14B347B0B6B22D1850CD" bold="true" pageId="0" pageNumber="1">B</emphasis>
urosome, ventral view 
<emphasis id="AB679A11F207549F98496C32E1231676" bold="true" pageId="0" pageNumber="1">C</emphasis>
right antennule, with grey objects indicating antennular spines 
<emphasis id="E37909622A2001F92DF74FD520E0B2BC" bold="true" pageId="0" pageNumber="1">D</emphasis>
P5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view.
</paragraph>
</caption>
<caption id="7BCBFB89B9800F2753D3F60391757BB8" doi="10.3897/zookeys.911.38496.figure7" httpUri="https://binary.pensoft.net/fig/379467" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph id="AE811AFFB0E3F8B13A10A9F91DF7AF4C" pageId="0" pageNumber="1">
<emphasis id="AABCF19A827CE93A56EC01ED6B9B7CA4" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<taxonomicName id="F7037B9CE7B506AB71B90461FA6F2588" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., male: 
<emphasis id="A406137EB76B644F4C8B2474521E8D92" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view 
<emphasis id="2A8CDFD2ACA3603853FC9453317BD992" bold="true" pageId="0" pageNumber="1">B</emphasis>
Pdg 5 and genital somite, dorsal view 
<emphasis id="60680A21A2643417904FCE9BDE7A2F8C" bold="true" pageId="0" pageNumber="1">C</emphasis>
Pdg 5 lateral wing, left view 
<emphasis id="9C75223982B38B1960B2917BC68C55D9" bold="true" pageId="0" pageNumber="1">D</emphasis>
Pdg 5, genital somite and urosomites 2 and 3, ventrolateral view 
<emphasis id="8AC58003FDBDA1F2A64B811266527D2A" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
right caudal ramus in dorsal (
<emphasis id="F0811F13A838AEE409F00CD123089881" bold="true" pageId="0" pageNumber="1">E</emphasis>
) and ventral (
<emphasis id="EF12CAB1C820106B3C5BB8DF6D072C5B" bold="true" pageId="0" pageNumber="1">F</emphasis>
) views 
<emphasis id="1B63C994D4139225D7507BDCEA411AA1" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
right antennule (white arrows indicate spines on segments 8, 10-18, 20.
</paragraph>
</caption>
<paragraph id="DB2C2CB628E2C5A93744D9989AEE4349" pageId="0" pageNumber="1">Left antennule, antenna, mouthparts, and P1-P4 as in female.</paragraph>
<paragraph id="FF8920F8A00DA3583E5A238245CA2BC3" pageId="0" pageNumber="1">
P5 (Figs 
<figureCitation id="814B2788814D656772DCCFF3FC8C419E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6D</figureCitation>
, 
<figureCitation id="98C3E540C05AD59B547B2995348EB6FB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8A, G</figureCitation>
) intercoxal sclerite with rounded lobe on free margin. Right P5: coxa with acute, stout spine on posterior lobe. Basis (Fig. 
<figureCitation id="280D4D65979514911501B08FC2BF69B5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8B, G</figureCitation>
) with large proximomedial triangular lamella at one-fourth length of inner margin; with large three-lobed chitinous medial prominence on posterior surface; distal outer margin with long, thin seta, slightly extending beyond Exp-1. Enp (Fig. 
<figureCitation id="93F54882574637731A4067280554C065" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8B, H, G</figureCitation>
) with bi-lobed distal margin, tipped with spinules and hyaline lamella on inner and outer lobes, respectively; reaching downward to approximately one-third of Exp-2. Exp-1 (Fig. 
<figureCitation id="B5199C49B087F77FD064EC13A8E2EAA0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8A, B, H</figureCitation>
) with semi-circular knob on distomedial margin; distolateral margin with small acute process. Exp-2 (Fig. 
<figureCitation id="7FE1C5B3DA2014BF5CF5B4A2C1BD7C07" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8C, H</figureCitation>
) elliptic, with three accessory lateral spines, one proximal, middle, and distal on lateral margin. Principal lateral spine articulated, located at two-third length of Exp-2, flat, thick, digitiform, with sharp tip; long, with approximately half of segment bearing it; slightly twisted in posterolateral direction. End claw (Figs 
<figureCitation id="5F53AAD3ACE4FBD7B42EF17C65E2EC23" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6D</figureCitation>
, 
<figureCitation id="040785FB3C50B4CB6B86A89E8EB11F13" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8A</figureCitation>
) medially sickle-shaped, slender towards tip, more than 1.5 times as long as Exp-2; medial margin serrated with row of tiny spinules.
</paragraph>
<paragraph id="95A030437F7C1F15657157C3CD48BD87" pageId="0" pageNumber="1">
Left P5 (Figs 
<figureCitation id="CA96FD8755AA05881BC902834744CC95" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phyllodiaptomus (P.) roietensis sp. nov., male: A habitus, dorsal view B urosome, ventral view C right antennule, with grey objects indicating antennular spines D P 5, with grey and dotted objects indicating hyaline lamella and chitinous prominences respectively, posterior view." figureDoi="10.3897/zookeys.911.38496.figure6" httpUri="https://binary.pensoft.net/fig/379466" pageId="0" pageNumber="1">6D</figureCitation>
, 
<figureCitation id="6744B06990F3A2B4DF71E7B48533A858" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8D</figureCitation>
): coxa with moderate strong seta inserted on posterior lobe at distal inner corner, slightly shorter than distal margin of basis. Basis with flap of longitudinal hyaline lamella at medial margin; with long, thin seta at posterolateral margin, extending to approximately half of Exp-1. Exp-1 (Fig. 
<figureCitation id="0A71A8E8B8EBC4EADD0739EBD70A3E7A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8F</figureCitation>
) tapering towards posterior margin, medial margin concave, with field of setules and tiny spinules. Exp-2 smaller than Exp-1, conical; with large seta at mid-length of medial margin, as long as Exp-2 and apical process combined; with few setules proximally and widespread with spinules distally along inner margin, thickness of spinules increased from proximal to distal; apical process stout, bare, and blunt-tip. Enp (Fig. 
<figureCitation id="FAD454A5E63BDEFC17C41F1D764F2800" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Phyllodiaptomus (P.) roietensis sp. nov., SEM photographs, male: A P 5 in posterior view B right P 5 basis, Exp- 1 and Enp, posterior view C right P 5 Exp- 2 in posterior view (white arrows indicate accessory spines) D left lobe of P 5, posterior view E left P 5 Exp- 2 and Enp, posterior view F left P 5 Enp, posterior view G P 5, anterior view H right P 5 Exp- 1 - 2 and Enp, anterior view I left P 5 basis, Exp and Enp, anterior view J left P 5 Enp (white arrow indicates Enp segmented point), anterior view." figureDoi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1">8F, J</figureCitation>
) bi-segmented, longer than Exp-1, Enp-2 tipped with row of spinules distally.
</paragraph>
<caption id="E0B63AE81ECAA5732340062D5AB5874D" doi="10.3897/zookeys.911.38496.figure8" httpUri="https://binary.pensoft.net/fig/379468" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph id="DE4A13E96F76F965D4E8EBC918E90C07" pageId="0" pageNumber="1">
<emphasis id="D5CD92FA4F38CA9543AE1E543F51AEAA" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
<taxonomicName id="4E3BC3B29F7963F6B8C1702610C624D8" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov., SEM photographs, male: 
<emphasis id="34A2E0EE09E06D3A50C18C1FD59E6AA3" bold="true" pageId="0" pageNumber="1">A</emphasis>
P5 in posterior view 
<emphasis id="E0D8559DDB2E550144D776ACB154B048" bold="true" pageId="0" pageNumber="1">B</emphasis>
right P5 basis, Exp-1 and Enp, posterior view 
<emphasis id="E0A8A796F023FA1CA998B3C59B8F8F20" bold="true" pageId="0" pageNumber="1">C</emphasis>
right P5 Exp-2 in posterior view (white arrows indicate accessory spines) 
<emphasis id="7924F6E79472F64A780E1B2B2F5D33E0" bold="true" pageId="0" pageNumber="1">D</emphasis>
left lobe of P5, posterior view 
<emphasis id="9CD61A5476531DF41F75C9D772DB935D" bold="true" pageId="0" pageNumber="1">E</emphasis>
left P5 Exp-2 and Enp, posterior view 
<emphasis id="BEFF6A31200DB1C8AD649AF87BCAA2A8" bold="true" pageId="0" pageNumber="1">F</emphasis>
left P5 Enp, posterior view 
<emphasis id="27B44C473A419EFFFED563583267521C" bold="true" pageId="0" pageNumber="1">G</emphasis>
P5, anterior view 
<emphasis id="4322A78AFCCEA14C9DDF8EF8EAEE0C68" bold="true" pageId="0" pageNumber="1">H</emphasis>
right P5 Exp-1-2 and Enp, anterior view 
<emphasis id="B365CA876340B7C7D13A28FC7179BD0A" bold="true" pageId="0" pageNumber="1">I</emphasis>
left P5 basis, Exp and Enp, anterior view 
<emphasis id="A1B2FBBA912C843B2BEC862E2F13ABD3" bold="true" pageId="0" pageNumber="1">J</emphasis>
left P5 Enp (white arrow indicates Enp segmented point), anterior view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E8CEDF4951F6DAB79CBDC2B34242CD5B" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="380E19C8C6CBFD4E963803C2C05B8281" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="3B4F1C0224E1BE4F6973F670D857235B" pageId="0" pageNumber="1">
<taxonomicName id="29B2CFFADD3D097E56DA54D63B131C44" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">Phyllodiaptomus (P.) roietensis</taxonomicName>
sp. nov. with the male P5 Exp-2 displays an affinity to the subgenus 
<taxonomicName id="7A2B04F1B366B3BB061888F96DFD6B53" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus">Phyllodiaptomus</taxonomicName>
sensu 
<bibRefCitation id="C18DEAF9F16D7110DB4A7D1202E5450A" DOI="https://doi.org/10.1007/BF00017591" author="Dumont, HJ" journalOrPublisher="Hydrobiologia," pageId="0" pageNumber="1" pagination="139 - 148" refId="B9" refString="Dumont, HJ, Ranga Reddy, Y, Sanoamuang, L, 1996. Description of Phyllodiaptomus christineae n. sp. from Thailand, and distinction of two subgenera within Phyllodiaptomus Kiefer, 1936 (Copepoda, Calanoida). Hydrobiologia, 323: 139 - 148, DOI: https://doi.org/10.1007/BF00017591" title="Description of Phyllodiaptomus christineae n. sp. from Thailand, and distinction of two subgenera within Phyllodiaptomus Kiefer, 1936 (Copepoda, Calanoida)." url="https://doi.org/10.1007/BF00017591" volume="323" year="1996">Dumont et al. (1996)</bibRefCitation>
: the lateral side of the right Exp-2, medially concave in posterior view, principal lateral spine inserted on distal to mid-outer margin and three accessary spines arranged from proximal, middle, and distal, respectively; the left Exp-2 with patch of strong spinules along medial margin.
</paragraph>
<paragraph id="72E8B63291001DDF7F17E2ACEC4491D4" pageId="0" pageNumber="1">The male of the new species has serrated outgrowth on the antepenultimate segment of the right antennule. The right caudal ramus with small chitinous spine near distal margin on ventral side and triangular prominence along proximal one-third length of outer margin. The P5 intercoxal sclerite produced, with convex distal margin. The right P5 with (1) short, strong spine on posterior lobe of coxa, (2) triangular hyaline lamella on proximal inner margin and large chitinous outgrowth on posterior surface of basis, (3) acute distal outer corner of Exp-1 (4) Exp-2 oval and concave, with strong, flat, curved principal spine and three accessary spines, and (5) bi-lobed Enp. The left P5 with long and narrow hyaline lamella along inner margin, Exp-2 with patch of strong spinules along medial margin, and bi-segmented Enp.</paragraph>
<paragraph id="B8A42A5204CE0DE098AA08B3847E551A" pageId="0" pageNumber="1">Female with asymmetrical Pdg 5 wings, left wing more elongated in posterio-lateral direction; posterior and dorsal spines short and strong. Genital double-somite with posterolateral directed process on right side. One pair of genital spines on lateral side slightly symmetrical and strong. P5 Exp-2 with conveyor canal on anterior surface. P5 with bi-segmented Enp.</paragraph>
</subSubSection>
<subSubSection id="7F0A99C2BD9E8EBAB275762074420543" pageId="0" pageNumber="1" type="etymology">
<paragraph id="4872C5EE3E00B7DF32B387B1DAA9B566" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="3D746DE2F1A1C590D99BD2112CC0AA69" pageId="0" pageNumber="1">
The specific name 
<taxonomicName id="6D7F01F0FFF63335551BC150AE595AA9" authorityName="Sanoamuang &amp; Watiroyram" authorityYear="2020" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="roietensis" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="roietensis">
<emphasis id="1204FBF5D322A8650F4ACDC216EA9155" italics="true" pageId="0" pageNumber="1">roietensis</emphasis>
</taxonomicName>
is taken after the type locality, Roi Et Province. The name with the Latin suffix &quot;- 
<emphasis id="238A7236DED49D509C7990A5ED2822C2" italics="true" pageId="0" pageNumber="1">ensis</emphasis>
&quot; is the adjective for a location.
</paragraph>
</subSubSection>
<subSubSection id="C4843D470C248F0B7BFCC61E6361A342" pageId="0" pageNumber="1" type="distribution">
<paragraph id="7AE98630D79495595103A0096BD3B558" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="F604663B540582BFFEEEC88E678C2162" pageId="0" pageNumber="1">
Known only from four temporary water bodies from Roi Et and Nakhon Ratchasima provinces, Thailand and Kampong Thom Province, Cambodia (Fig. 
<figureCitation id="DA626FDA9D3DE238F880E1108C88D667" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Phyllodiaptomus (P.) roietensis sp. nov. and P. (P.) surinensis. Key: black square = city, black circle = P. (P.) roietensis sp. nov., black triangular = P. (P.) surinensis, blue arrows indicate water flow direction." figureDoi="10.3897/zookeys.911.38496.figure1" httpUri="https://binary.pensoft.net/fig/379461" pageId="0" pageNumber="1">1</figureCitation>
). Presence of specimens was recorded in early monsoon period. The new species is rare, as it was found in 0.4% of all the localities sampled in Cambodia. The new species was found together with six diaptomids including 
<taxonomicName id="984134FA5EC6CD82FCB21BA964FAC091" baseAuthorityName="Grochmalicki" baseAuthorityYear="1915" class="Copepoda" family="Diaptomidae" genus="Dentodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dentodiaptomus javanus" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="AEAC35BFF232C79E46B1658574FB4288" italics="true" pageId="0" pageNumber="1">Dentodiaptomus javanus</emphasis>
</taxonomicName>
(Grochmalicki, 1915), 
<taxonomicName id="539031C5347104323CE2DD3609CEA93D" authorityName="Ranga Reddy &amp; Dumont" authorityYear="1998" class="Copepoda" family="Diaptomidae" genus="Eodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eodiaptomus sanoamuangae" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sanoamuangae">
<emphasis id="E34E4E4C91D259BAC6D9680F27F2DD14" italics="true" pageId="0" pageNumber="1">Eodiaptomus sanoamuangae</emphasis>
</taxonomicName>
Ranga Reddy &amp; Dumont, 1998, 
<taxonomicName id="56288FB785D219136C52C12D1E6FF30F" baseAuthorityName="Shen &amp; Tai" baseAuthorityYear="1965" class="Copepoda" family="Diaptomidae" genus="Mongolodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mongolodiaptomus calcarus" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="calcarus">
<emphasis id="C27F4FCBC43B5F88B20145C48ACEB1FF" italics="true" pageId="0" pageNumber="1">Mongolodiaptomus calcarus</emphasis>
</taxonomicName>
(Shen &amp; Tai, 1965), 
<taxonomicName id="87DE6D5BC54EF2FC38ABBC56D48386A9" lsidName="M. malaindosinensis" pageId="0" pageNumber="1" rank="species" species="malaindosinensis">
<emphasis id="178EC2773EDC9F84DF6CD7271C8B056C" italics="true" pageId="0" pageNumber="1">M. malaindosinensis</emphasis>
</taxonomicName>
(Lai &amp; Fernando, 1978), 
<taxonomicName id="5744D1508E36F8EDF4519197E1A478B9" authorityName="Kiefer" authorityYear="1974" class="Copepoda" family="Diaptomidae" genus="Neodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neodiaptomus laii" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="laii">
<emphasis id="BC05EE5B2FEE2F41E0ADAACD67C8ED6D" italics="true" pageId="0" pageNumber="1">Neodiaptomus laii</emphasis>
</taxonomicName>
Kiefer, 1974, and 
<taxonomicName id="3F9BFCC4E7F49F23EF408BED7AEB37E5" authorityName="Dumont, Ranga Reddy &amp; Sanoamuang" authorityYear="1996" class="Copepoda" family="Diaptomidae" genus="Phyllodiaptomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodiaptomus christineae" order="Calanoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="christineae">Phyllodiaptomus (Phyllodiaptomus) christineae</taxonomicName>
Dumont, Ranga Reddy &amp; Sanoamuang, 1996.
</paragraph>
</subSubSection>
</treatment>
</document>