209 lines
20 KiB
XML
209 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4391157" ID-GBIF-Dataset="39c9ce8d-d95d-467e-9590-ab184d952201" ID-GBIF-Taxon="159879583" ID-Zenodo-Dep="4391157" checkinTime="1562074625476" checkinUser="jeremy" docAuthor="DeMarmels, Jurg" docDate="1992" docId="0B24F87CFFBAFFB5FF02BB93C8B126CE" docLanguage="en" docName="De Marmels - 1992 - Caballitos del diablo (Odonata) de las Sierras de Tapirapeco y Unturán, en el extremo Sur de Venezuela.pdf.imf" docOrigin="Acta Biologica Venezuelica 14 (1)" docStyle="DocumentStyle{}" docTitle="Dimeragrion unturanense DeMarmels 1992, sp. n." docType="treatment" docVersion="12" lastPageId="3" lastPageNumber="59" masterDocId="F71D8004FFB8FFB6FFC7B11BC92F2226" masterDocTitle="Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela" masterLastPageNumber="78" masterPageNumber="57" pageId="2" pageNumber="59" updateTime="1640618713225" updateUser="jeremy">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Dragonflies (Odonata) From The Sierras Of Tapirapeco And Unturan, In The Extreme South Of Venezuela</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DeMarmels, Jurg</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Biologica Venezuelica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1992</mods:date>
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<mods:detail type="volume">
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<mods:number>14</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3266063</mods:identifier>
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<mods:identifier type="GBIF-Dataset">39c9ce8d-d95d-467e-9590-ab184d952201</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3266063</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4391157" ID-GBIF-Taxon="159879583" ID-Zenodo-Dep="4391157" LSID="urn:lsid:plazi:treatment:0B24F87CFFBAFFB5FF02BB93C8B126CE" httpUri="http://treatment.plazi.org/id/0B24F87CFFBAFFB5FF02BB93C8B126CE" lastPageId="3" lastPageNumber="59" pageId="2" pageNumber="59">
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<subSubSection box="[197,808,2696,2735]" pageId="2" pageNumber="59" type="nomenclature">
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<paragraph blockId="2.[194,1153,2693,3273]" box="[197,808,2696,2735]" pageId="2" pageNumber="59">
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<heading box="[197,808,2696,2735]" fontSize="13" level="2" pageId="2" pageNumber="59" reason="1">
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<taxonomicName authority="DeMarmels, 1992" authorityName="DeMarmels" authorityYear="1992" box="[197,681,2696,2734]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="unturanense" status="sp. n.">
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<emphasis box="[197,681,2696,2734]" italics="true" pageId="2" pageNumber="59">Dimeragrion unturanense</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[703,808,2697,2735]" pageId="2" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[240,477,2750,2788]" pageId="2" pageNumber="59" type="description">
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<paragraph blockId="2.[194,1153,2693,3273]" box="[240,477,2750,2788]" pageId="2" pageNumber="59">
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<figureCitation box="[240,477,2750,2788]" captionStart="Plate II" captionStartId="14.[193,288,3057,3091]" captionTargetBox="[212,2070,564,2898]" captionTargetPageId="14" captionText="Plate II: Figure 7. Chalcopteryx scintillans, female ultimate instar exuviae, labium, ventral view. Figures 8-10. Dimeragrion unturanense sp.n. (8) holotype male, abdominal segment 10 with anal appendages, left lateral view; (9) same, dorsal view; (10) female, paratype, tip of abdomen, left lateral view. Figure3 11-12. Megapodagrion megalopus, male. (11) penis, ri.ght lateral view; (12) same, ventral view. Figures 13-14. Allopodagrion oscillans (Venezuela, Tachira, Betania). male. (13) penis, right lateral view; (14) same, ventral view." httpUri="https://zenodo.org/record/3266069/files/figure.png" pageId="2" pageNumber="59">Figures 8-10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="59" type="materials_examined">
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<paragraph blockId="2.[194,1153,2693,3273]" pageId="2" pageNumber="59">
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<materialsCitation ID-GBIF-Occurrence="2433522129" collectingDate="1989-02-15" collectingDateMax="1989-02-18" collectingDateMin="1989-02-15" collectionCode="MIZA" collectorName="J. DeMarmels" country="Venezuela" elevation="950" latitude="1.4166666" location="Unturan (Western Sierra de Unturan)" longitude="-65.3" pageId="2" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
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<specimenCount box="[238,335,2802,2840]" pageId="2" pageNumber="59" type="male">Male</specimenCount>
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(
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<typeStatus box="[379,547,2802,2840]" pageId="2" pageNumber="59">holotype</typeStatus>
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).
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<collectingCountry box="[597,791,2802,2840]" name="Venezuela" pageId="2" pageNumber="59" value="Venezuela">Venezuela</collectingCountry>
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,
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<collectingRegion box="[828,1141,2802,2840]" pageId="2" pageNumber="59">Amazonas State</collectingRegion>
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, Sierra de
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<collectingMunicipality box="[391,549,2855,2893]" pageId="2" pageNumber="59">Unturan</collectingMunicipality>
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,
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<elevation box="[576,700,2855,2893]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="59" unit="m" value="950.0">
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<quantity box="[576,700,2855,2893]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="59" unit="m" value="950.0">950 m</quantity>
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</elevation>
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,
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<geoCoordinate box="[731,879,2855,2893]" direction="north" orientation="latitude" pageId="2" pageNumber="59" precision="925" value="1.4166666">01°25'N</geoCoordinate>
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,
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<geoCoordinate box="[907,1064,2855,2893]" direction="west" orientation="longitude" pageId="2" pageNumber="59" precision="925" value="-65.3">65°18'W</geoCoordinate>
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,
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<collectingDate pageId="2" pageNumber="59" value="1989-02-15" valueMax="1989-02-18" valueMin="1989-02-15">
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<date pageId="2" pageNumber="59" value="1989-02-15" valueMax="1989-02-18" valueMin="1989-02-15">15- 18 Feb 1989</date>
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</collectingDate>
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,
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<collectorName box="[461,788,2900,2948]" pageId="2" pageNumber="59">J DEMARMELS</collectorName>
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leg
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</materialsCitation>
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. Material (5 ♂, 1 ♀, including
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<typeStatus box="[493,655,2960,2998]" pageId="2" pageNumber="59">holotype</typeStatus>
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and
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<typeStatus box="[771,960,2960,2998]" pageId="2" pageNumber="59">paratypes</typeStatus>
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).
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<materialsCitation ID-GBIF-Occurrence="2433522144" collectingDate="1989-02-15" collectingDateMax="1989-02-18" collectingDateMin="1989-02-15" collectionCode="MIZA" collectorName="J. DeMarmels" country="Venezuela" elevation="950" latitude="1.4166666" location="Unturan (Western Sierra de Unturan)" longLatPrecision="1308" longitude="-65.3" pageId="2" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
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<specimenCount box="[1005,1060,2957,2998]" pageId="2" pageNumber="59" type="male">1 ♂</specimenCount>
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,
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<typeStatus pageId="2" pageNumber="59">paratype</typeStatus>
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, same dates as holotype
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</materialsCitation>
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,
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<materialsCitation ID-GBIF-Occurrence="2433522098" collectingDate="1989-02-19" collectingDateMax="1989-02-21" collectingDateMin="1989-02-19" collectionCode="MIZA" collectorName="J. DeMarmels & A. Chacon" country="Venezuela" elevation="1300" latitude="1.2166667" location="Tamacuari (Sierra de Tapirapeco)" longLatPrecision="21" longitude="-64.691666" pageId="2" pageNumber="59" specimenCount="4" specimenCount-male="4" stateProvince="Amazonas" typeStatus="paratype">
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<specimenCount box="[860,913,3010,3051]" pageId="2" pageNumber="59" type="male">4 ♂</specimenCount>
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,
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<typeStatus box="[961,1144,3013,3051]" pageId="2" pageNumber="59">paratypes</typeStatus>
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, Tamacuari
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="59" type="description">
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<paragraph blockId="2.[194,1153,2693,3273]" lastBlockId="2.[1269,2238,414,3274]" pageId="2" pageNumber="59">
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Male (
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<typeStatus box="[370,537,3130,3168]" pageId="2" pageNumber="59">holotype</typeStatus>
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). Labium dark brown, face brilliant black, a white transverse band across antedypeus and base of labrum; occiput and top of head black, a rufous streak between each lateral ocellus and corresponding antenna; frons angulate. Prothorax dorsally brassy black, its hind margin convexly rounded; pterothorax dark brown to black with metallic blue or green reflections; mesepisternum and mesepimeron densely covered with bluish pruinescence; some pruinosity also along sutures and ventrally. Legs dark brown to black. Wings slightly smoky, pterostigma dark brown. 23 (25) postnodal cross·veins (px) in fore wings 19 (20) in hind wings. 2 cubito-anal cross. veins (Cux) in three wings, 3 in left fore wing. 2 antenodal cells in discoidal field in all but in left fore wing (3). R4 originating very little before subnodus, IR, at first px. R3 branching after px 6 in fore wings, at px 6 in hind wings; IR2 originating at px 11 in fore wings, in hind wings at or after px 9. Abdomen dark brown to black; segments 3-6 with a brown basal ring; segments 1,2 and 9 with blue pruinescence dorsally. Superior anal appendages slightly shorter than segment 10 and triangularly pointed in lateral view (Fig. 8) in dorsal view tips convergent (Fig. 9). Penis similar to
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<taxonomicName authority="DeMarmels" authorityName="DeMarmels" box="[1444,2008,1632,1670]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="mesembrinum">
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<emphasis box="[1444,1478,1633,1670]" italics="true" pageId="2" pageNumber="59">D</emphasis>
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.
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<emphasis box="[1508,1772,1632,1670]" italics="true" pageId="2" pageNumber="59">mesembrinum</emphasis>
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DeMarmels
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</taxonomicName>
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(
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<bibRefCitation author="DEMARMELS, J." journalOrPublisher="Boletin de la Academia de las Ciencias Fisicas, Matematicas y Naturales, Caracas" pageId="2" pageNumber="59" pagination="81 - 87" part="25" refId="ref10274" refString="DEMARMELS, J. 1989 b. - Odonata or dragonflies from Cerro de la Neblina and the adjacent lowland between the Rio Baria, the Casiquiare and the Rio Negro (Venezuela). II. Additions to the adults. Boletin de la Academia de las Ciencias Fisicas, Matematicas y Naturales, Caracas, 25: 81 - 87." title="Odonata or dragonflies from Cerro de la Neblina and the adjacent lowland between the Rio Baria, the Casiquiare and the Rio Negro (Venezuela). II. Additions to the adults" type="journal article" year="1989">
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DEMARMELS, 1989b:
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<figureCitation box="[1573,1744,1685,1723]" captionStart="Plate I" captionStartId="13.[201,296,3134,3168]" captionTargetBox="[266,2132,441,3065]" captionTargetPageId="13" captionText="Plate I: Figures 1-6. Iridictyon trebbaui, larva. (1) male larva. ultimate instar, dorsal view; (2) right antenna; (3) labium, dorsal view; (4) abdominal segments 9 (with gonapophyses) and 10, ventral view (all same specimen); (5) female larva ¿.ante?) penultimate instar, detached lateral gill; (6) median gill, left lateral view (same specimen)." httpUri="https://zenodo.org/record/3266067/files/figure.png" pageId="2" pageNumber="59">Figs. 4-5</figureCitation>
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</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="2.[1269,2238,414,3274]" pageId="2" pageNumber="59">Measurements. Tctal length (excl. anal appendages) 38.0; abdomen (excl. anal app.) 30.5; anal appendages 0.8; hind wing 35.2; pterostigma (costal side, front wing) 1.0; radial side 2.0.</paragraph>
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<paragraph blockId="2.[1269,2238,414,3274]" pageId="2" pageNumber="59">
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The
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<typeStatus box="[1409,1572,1973,2011]" pageId="2" pageNumber="59">paratype</typeStatus>
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males do not differ substantially from the
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<typeStatus box="[1458,1631,2026,2064]" pageId="2" pageNumber="59">holotype</typeStatus>
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: 3-4 antenodal cells in the discoidal field; minor variations in the branching of IR2 and R3; R4 often originating slightly more proximally. Measurements. Total length (excl. appendages) 40.5-43.5.
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</paragraph>
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<paragraph blockId="2.[1269,2238,414,3274]" pageId="2" pageNumber="59">
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Female (paraytpe). Labium pale; pale transverse band across face also including mandibles and genae. Prothorax as in male. Pterothorax brown whith cooppery reflections and yellowish stripes along humeral and lateral sutures. Wings smoky, pterostigma brown. 24(25) px in fore wings, 21 (22) in hind wings. Only 2 antenodal cells in the discoidal field; 2-3 cux. R3 originating after px 7 in three wings, after px 8 in left fore wing; IR2 branching at px 12 (10) in fore wings, at px 9(11) in hind wings. R4 originating more proximally from subnodus than in the
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<typeStatus box="[1867,2031,2877,2915]" pageId="2" pageNumber="59">holotype</typeStatus>
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. Abdomen brown; cerci two times the length of segment 10
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<figureCitation box="[1280,1450,2994,3032]" captionStart="Plate II" captionStartId="14.[193,288,3057,3091]" captionTargetBox="[212,2070,564,2898]" captionTargetPageId="14" captionText="Plate II: Figure 7. Chalcopteryx scintillans, female ultimate instar exuviae, labium, ventral view. Figures 8-10. Dimeragrion unturanense sp.n. (8) holotype male, abdominal segment 10 with anal appendages, left lateral view; (9) same, dorsal view; (10) female, paratype, tip of abdomen, left lateral view. Figure3 11-12. Megapodagrion megalopus, male. (11) penis, ri.ght lateral view; (12) same, ventral view. Figures 13-14. Allopodagrion oscillans (Venezuela, Tachira, Betania). male. (13) penis, right lateral view; (14) same, ventral view." httpUri="https://zenodo.org/record/3266069/files/figure.png" pageId="2" pageNumber="59">(Fig. 10)</figureCitation>
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.
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</paragraph>
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<paragraph blockId="2.[1269,2238,414,3274]" pageId="2" pageNumber="59">Measurements. Total length (excl. anal appendages) 31.5; abdomen (excl. app.) 25.0.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59" type="discussion">
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<paragraph blockId="2.[1269,2238,414,3274]" lastBlockId="3.[185,1151,413,3274]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
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Remarks. Male
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<taxonomicName authority="DeMarmels, 1992" authorityName="DeMarmels" authorityYear="1992" box="[1627,1917,3180,3219]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="unturanense" status="sp.n.">
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<emphasis box="[1627,1917,3180,3219]" italics="true" pageId="2" pageNumber="59">D. unturanense</emphasis>
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</taxonomicName>
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differ from male
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<emphasis box="[1334,1652,3235,3273]" italics="true" pageId="2" pageNumber="59">
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<taxonomicName box="[1334,1644,3235,3273]" pageId="2" pageNumber="59">D.mesembrinum</taxonomicName>
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,
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</emphasis>
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<taxonomicName authority="Calvert" authorityName="Calvert" authorityYear="1913" box="[1688,2135,3235,3274]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="percubitale">
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<emphasis box="[1688,1726,3236,3274]" italics="true" pageId="2" pageNumber="59">D.</emphasis>
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<emphasis box="[1757,1961,3235,3273]" italics="true" pageId="2" pageNumber="59">percubitale</emphasis>
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Calvert
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</taxonomicName>
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and
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<taxonomicName authority="Needham" authorityName="Needham" authorityYear="1933" box="[185,631,413,450]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="secundum">
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<emphasis box="[185,217,413,450]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[249,434,413,450]" italics="true" pageId="3" pageNumber="60">secundum</emphasis>
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Needham
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</taxonomicName>
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in having shorter, triangularly pointed superior anal appendages (lateral view) with the basal dorsal tubercle higher than tergal surface of segment 10. Tips of inferior appendages strongly curved upwards and slightly cephalad. The female of
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<taxonomicName authorityName="Needham" authorityYear="1933" box="[702,966,679,716]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="secundum">
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<emphasis box="[702,735,679,716]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[780,966,679,716]" italics="true" pageId="3" pageNumber="60">secundum</emphasis>
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</taxonomicName>
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is unknown, but probably' differs from
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<taxonomicName authorityName="DeMarmels" authorityYear="1992" box="[854,1146,732,769]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="unturanense">
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<emphasis box="[854,887,732,769]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[920,1146,732,769]" italics="true" pageId="3" pageNumber="60">unturanense</emphasis>
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</taxonomicName>
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in color pattern of head and in having only one cux. Female
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<taxonomicName authorityName="DeMarmels" authorityYear="1992" box="[440,734,839,876]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="unturanense">
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<emphasis box="[440,734,839,876]" italics="true" pageId="3" pageNumber="60">D. unturanense</emphasis>
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</taxonomicName>
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have the dorsobasal tubercle of the cerci slightly better developed female
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<taxonomicName authorityName="Calvert" authorityYear="1913" box="[297,569,944,981]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="percubitale">
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<emphasis box="[297,331,944,981]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[363,569,944,981]" italics="true" pageId="3" pageNumber="60">percubitale</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[694,1038,944,981]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="mesembrinum">
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<emphasis box="[694,727,944,981]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[770,1038,944,981]" italics="true" pageId="3" pageNumber="60">mesembrinum</emphasis>
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</taxonomicName>
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<emphasis box="[1038,1044,944,981]" italics="true" pageId="3" pageNumber="60">.</emphasis>
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The latter has only one cux in all wings. At both localities
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<taxonomicName authorityName="DeMarmels" authorityYear="1992" box="[347,649,1052,1089]" class="Insecta" family="Megapodagrionidae" genus="Dimeragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="unturanense">
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<emphasis box="[347,385,1052,1089]" italics="true" pageId="3" pageNumber="60">D</emphasis>
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.
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<emphasis box="[422,649,1052,1089]" italics="true" pageId="3" pageNumber="60">unturanense</emphasis>
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</taxonomicName>
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was taken along small creeks in the forest together with
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|
<taxonomicName authorityName="Racenis" authorityYear="1968" class="Insecta" family="Calopterygidae" genus="Iridictyon" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trebbaui">
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|
<emphasis box="[856,1039,1108,1145]" pageId="3" pageNumber="60">Iridictyon</emphasis>
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|
<emphasis italics="true" pageId="3" pageNumber="60">trebbaui</emphasis>
|
|
</taxonomicName>
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|
and (larvae of)
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|
<taxonomicName authority="Racenis" authorityName="Racenis" box="[600,1145,1162,1200]" class="Insecta" family="Corduliidae" genus="Neocordulia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="biancoi">
|
|
<emphasis box="[600,829,1162,1200]" italics="true" pageId="3" pageNumber="60">Neocordulia</emphasis>
|
|
<emphasis box="[849,984,1163,1200]" italics="true" pageId="3" pageNumber="60">biancoi</emphasis>
|
|
Racenis
|
|
</taxonomicName>
|
|
(see below).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |