302 lines
24 KiB
XML
302 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1111.77371" ID-Pensoft-Pub="1313-2970-1111-425" ID-Pensoft-UUID="028D93AD33835AD589E06F29DAE7580C" ID-ZooBank="8474125F447546C3A6DCC46F7D038BDC" ModsDocID="1313-2970-1111-425" checkinTime="1657577134577" checkinUser="pensoft" docAuthor="Thomson, Robin E., Armitage, Brian J. & Harris, Steven C." docDate="2022" docId="0AE63BE727355777955239625A0FE4E0" docLanguage="en" docName="ZooKeys 1111: 425-466" docOrigin="ZooKeys 1111" docPubDate="2022-07-11" docSource="http://dx.doi.org/10.3897/zookeys.1111.77371" docTitle="Leucotrichia ruiteri Thomson & Armitage & Harris 2022, sp. nov." docType="treatment" docUuid="F7961E34-1751-4095-A5D1-27D5E29B698D" docUuidSource="ZooBank" docVersion="1" id="028D93AD33835AD589E06F29DAE7580C" lastPageNumber="425" masterDocId="028D93AD33835AD589E06F29DAE7580C" masterDocTitle="The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama" masterLastPageNumber="466" masterPageNumber="425" pageNumber="425" updateTime="1657577436089" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Trichoptera of Panama. XIX. Additions to and a review of the genus Leucotrichia (Trichoptera, Hydroptilidae) in Panama</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Thomson, Robin E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5883-2040</mods:nameIdentifier>
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<mods:affiliation>University of Minnesota, Department of Entomology, 1980 Folwell Ave., St. Paul, MN 55108, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Armitage, Brian J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3182-1533</mods:nameIdentifier>
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<mods:affiliation>Museo de Peces de Agua Dulce e Invertebrados, Universidad Autonoma de Chiriqui, David, Panama</mods:affiliation>
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<mods:nameIdentifier type="email">tobikera89@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harris, Steven C.</mods:namePart>
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<mods:affiliation>Department of Biology and Geosciences, Clarion University, Clarion, PA, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1111</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>425</mods:start>
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<mods:end>466</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1111.77371</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1111.77371</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1111-425</mods:identifier>
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<mods:identifier type="ZooBank">8474125F447546C3A6DCC46F7D038BDC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">028D93AD33835AD589E06F29DAE7580C</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:F7961E34-1751-4095-A5D1-27D5E29B698D" httpUri="http://treatment.plazi.org/id/0AE63BE727355777955239625A0FE4E0" lastPageNumber="425" pageId="0" pageNumber="425">
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<subSubSection pageId="0" pageNumber="425" type="nomenclature">
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<paragraph pageId="0" pageNumber="425">
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<taxonomicName LSID="http://zoobank.org/F7961E34-1751-4095-A5D1-27D5E29B698D" authority="Thomson & Armitage & Harris, 2022" authorityName="Thomson & Armitage & Harris" authorityYear="2022" class="Insecta" family="Hydroptilidae" genus="Leucotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucotrichia ruiteri" order="Trichoptera" pageId="0" pageNumber="425" phylum="Arthropoda" rank="species" species="ruiteri" status="sp. nov.">Leucotrichia ruiteri</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="425">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="description">
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<paragraph pageId="0" pageNumber="425">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Modifications A Forewing, Leucotrichia ruiteri sp. nov. B head and scape, Leucotrichia chiriquiensis Flint, 1970 C head and antennae, Leucotrichia fairchildi Flint, 1970. Fig. 2 B and 2 C modified from Thomson and Holzenthal (2015)." figureDoi="10.3897/zookeys.1111.77371.figure2" httpUri="https://binary.pensoft.net/fig/713490" pageId="0" pageNumber="425">Figs 2A</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">, 14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="type locality">
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<paragraph pageId="0" pageNumber="425">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="425">
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<emphasis bold="true" pageId="0" pageNumber="425">Panama: Chiriqui Province</emphasis>
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: Cuenca 108; Boquete District; Quebrada Jaramillo, Finca Monterey;
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<geoCoordinate degrees="8.7632" direction="north" orientation="latitude" precision="5" value="8.7632">8.7632°N</geoCoordinate>
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,
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<geoCoordinate degrees="82.41383" direction="west" orientation="longitude" precision="1" value="-82.41383">82.41383°W</geoCoordinate>
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; 1,214 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="type material">
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<paragraph pageId="0" pageNumber="425">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="425">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="425">Holotype</emphasis>
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: male,
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<emphasis bold="true" pageId="0" pageNumber="425">Panama: Chiriqui Province</emphasis>
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: Cuenca 108; Boquete District; Quebrada Jaramillo, Finca Monterey;
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<geoCoordinate degrees="8.7632" direction="north" orientation="latitude" precision="5" value="8.7632">8.7632°N</geoCoordinate>
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,
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<geoCoordinate degrees="82.41383" direction="west" orientation="longitude" precision="1" value="-82.41383">82.41383°W</geoCoordinate>
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; 1,214 m a.s.l.; 8-12 Jun. 2018, K. Collier, leg.; Malaise trap; in alcohol; MIUP-004-T-2021.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="425">Paratype</emphasis>
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: same data as for holotype; 1 male; UMSP.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="materials_examined">
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<paragraph pageId="0" pageNumber="425">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="425">
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<materialsCitation collectingDate="2017-09-01" collectingDateMax="2017-09-05" collectingDateMin="2017-09-01" collectingMethod="Malaise trap" collectorName="La Amistad International Park, E. Alvarez, T. Rios, E. Perez" country="Panama" county="Cuenca" elevation="1799" latitude="8.90614" location="La Amistad International Park" longLatPrecision="1" longitude="-82.72882" municipality="Renacimiento District" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui Province">
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<emphasis bold="true" pageId="0" pageNumber="425">
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<collectingCountry name="Panama">Panama</collectingCountry>
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:
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<collectingRegion country="Panama" name="Chiriqui">Chiriqui Province</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">1 male</specimenCount>
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;
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<collectingCounty>Cuenca</collectingCounty>
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102,
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<collectingMunicipality>Renacimiento District</collectingMunicipality>
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;
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<collectorName>
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<location LSID="urn:lsid:plazi:treatment:0AE63BE727355777955239625A0FE4E0:AAD5A990865F4488545829BCF732A431" country="Panama" county="Cuenca" latitude="8.90614" longLatPrecision="1" longitude="-82.72882" municipality="Renacimiento District" name="La Amistad International Park" stateProvince="Chiriqui Province">La Amistad International Park</location>
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</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:0AE63BE727355777955239625A0FE4E0:4AADD2D131C4A5B71CEB397B2A1D98DF" country="Panama" county="Cuenca" latitude="8.90614" longLatPrecision="1" longitude="-82.72882" municipality="Renacimiento District" name="Rio Candela" stateProvince="Chiriqui Province">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Candela
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</location>
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,
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<location LSID="urn:lsid:plazi:treatment:0AE63BE727355777955239625A0FE4E0:E7D0B803F9A305176C2957FA5C72E508" country="Panama" county="Cuenca" latitude="8.90614" longLatPrecision="1" longitude="-82.72882" municipality="Renacimiento District" name="Finca Felix" stateProvince="Chiriqui Province">Finca Felix</location>
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, PSPSCB-PILA-C102-2017-021;
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<geoCoordinate degrees="8.90614" direction="north" orientation="latitude" precision="1" value="8.90614">8.90614°N</geoCoordinate>
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,
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<geoCoordinate degrees="82.72882" direction="west" orientation="longitude" precision="1" value="-82.72882">82.72882°W</geoCoordinate>
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;
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.799" unit="m" value="1799.0">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.799" unit="m" value="1799.0">1,799 m</quantity>
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a.s.l.
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</elevation>
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,
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<collectingDate value="2017-09-01" valueMax="2017-09-05" valueMin="2017-09-01">1-5 Sep. 2017</collectingDate>
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;
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<collectorName>
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E.
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<normalizedToken originalValue="Álvarez">Alvarez</normalizedToken>
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</collectorName>
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,
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<collectorName>
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T.
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<normalizedToken originalValue="Ríos">Rios</normalizedToken>
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</collectorName>
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,
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<collectorName>
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E.
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<normalizedToken originalValue="Pérez">Perez</normalizedToken>
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</collectorName>
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, leg.;
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<collectingMethod>Malaise trap</collectingMethod>
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; in alcohol; COZEM
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</materialsCitation>
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.
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<materialsCitation collectingDate="2017-04-19" collectingDateMax="2017-04-23" collectingDateMin="2017-04-19" collectingMethod="Malaise trap" collectorName="A. Cornejo, T. Rios, E. Alvarez, C. Nieto" country="Panama" county="Cuenca" elevation="623" latitude="8.52577" location="Rio Mulaba" longLatPrecision="1" longitude="-81.13045" municipality="Santa Fe National Park" specimenCount="1" specimenCount-male="1" stateProvince="Veraguas Province">
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<emphasis bold="true" pageId="0" pageNumber="425">
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<collectingRegion country="Panama" name="Veraguas">Veraguas Province</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">1 male</specimenCount>
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;
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<collectingCounty>Cuenca</collectingCounty>
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132,
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<collectingMunicipality>Santa Fe National Park</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:0AE63BE727355777955239625A0FE4E0:719B857EBF46AC25CABF70C6EC4A3122" country="Panama" county="Cuenca" latitude="8.52577" longLatPrecision="1" longitude="-81.13045" municipality="Santa Fe National Park" name="Rio Mulaba" stateProvince="Veraguas Province">
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Mulaba
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</location>
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, 2do
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<location LSID="urn:lsid:plazi:treatment:0AE63BE727355777955239625A0FE4E0:591D9F00ED041CC5C16891A964EE4D45" country="Panama" county="Cuenca" latitude="8.52577" longLatPrecision="1" longitude="-81.13045" municipality="Santa Fe National Park" name="Brazo" stateProvince="Veraguas Province">Brazo</location>
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, PSPSCB-NPSF-C-097-2017-007;
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<geoCoordinate degrees="8.52577" direction="north" orientation="latitude" precision="1" value="8.52577">8.52577°N</geoCoordinate>
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,
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<geoCoordinate degrees="81.13045" direction="west" orientation="longitude" precision="1" value="-81.13045">81.13045°W</geoCoordinate>
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;
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<elevation metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">
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<quantity metricMagnitude="2" metricUnit="m" metricValue="6.2299999999999995" unit="m" value="623.0">623 m</quantity>
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a.s.l.
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</elevation>
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;
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<collectingDate value="2017-04-19" valueMax="2017-04-23" valueMin="2017-04-19">19-23 Apr. 2017</collectingDate>
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;
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<collectorName>A. Cornejo</collectorName>
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,
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<collectorName>
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T.
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<normalizedToken originalValue="Ríos">Rios</normalizedToken>
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</collectorName>
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,
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<collectorName>
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E.
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<normalizedToken originalValue="Álvarez">Alvarez</normalizedToken>
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</collectorName>
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,
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<collectorName>C. Nieto</collectorName>
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, leg.;
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<collectingMethod>Malaise trap</collectingMethod>
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; in alcohol; MUPADI
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="diagnosis">
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<paragraph pageId="0" pageNumber="425">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="425">
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<taxonomicName authorityName="Thomson & Armitage & Harris" authorityYear="2022" class="Insecta" family="Hydroptilidae" genus="Leucotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucotrichia ruiteri" order="Trichoptera" pageId="0" pageNumber="425" phylum="Arthropoda" rank="species" species="ruiteri">
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<emphasis italics="true" pageId="0" pageNumber="425">Leucotrichia ruiteri</emphasis>
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</taxonomicName>
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sp. nov. is most similar to
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<taxonomicName genus="L." lsidName="L. hispida" pageId="0" pageNumber="425" rank="species" species="hispida">
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<emphasis italics="true" pageId="0" pageNumber="425">L. hispida</emphasis>
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</taxonomicName>
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, as both species bear prominent setae on the posteroventral production of sternum VIII, a bilobed phallus apex, and a similar shaped inferior appendage.
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<taxonomicName authorityName="Thomson & Armitage & Harris" authorityYear="2022" class="Insecta" family="Hydroptilidae" genus="Leucotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucotrichia ruiteri" order="Trichoptera" pageId="0" pageNumber="425" phylum="Arthropoda" rank="species" species="ruiteri">
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<emphasis italics="true" pageId="0" pageNumber="425">Leucotrichia ruiteri</emphasis>
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</taxonomicName>
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can be separated by the single, elongate seta on sternum VIII compared to the cluster of setae present on
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<taxonomicName genus="L." lsidName="L. hispida" pageId="0" pageNumber="425" rank="species" species="hispida">
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<emphasis italics="true" pageId="0" pageNumber="425">L. hispida</emphasis>
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</taxonomicName>
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. Additionally, the forewings of
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<taxonomicName genus="L." lsidName="L. ruiteri" pageId="0" pageNumber="425" rank="species" species="ruiteri">
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<emphasis italics="true" pageId="0" pageNumber="425">L. ruiteri</emphasis>
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</taxonomicName>
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are modified with a large setae-filled pocket, while those of
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<taxonomicName genus="L." lsidName="L. hispida" pageId="0" pageNumber="425" rank="species" species="hispida">
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<emphasis italics="true" pageId="0" pageNumber="425">L. hispida</emphasis>
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</taxonomicName>
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are unmodified.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="description">
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<paragraph pageId="0" pageNumber="425">Description.</paragraph>
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<paragraph pageId="0" pageNumber="425">
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<emphasis bold="true" pageId="0" pageNumber="425">Male.</emphasis>
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Length of forewing 2.0-2.1 mm (
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<emphasis italics="true" pageId="0" pageNumber="425">n</emphasis>
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= 4). Forewing with large pocket filled with scales (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Modifications A Forewing, Leucotrichia ruiteri sp. nov. B head and scape, Leucotrichia chiriquiensis Flint, 1970 C head and antennae, Leucotrichia fairchildi Flint, 1970. Fig. 2 B and 2 C modified from Thomson and Holzenthal (2015)." figureDoi="10.3897/zookeys.1111.77371.figure2" httpUri="https://binary.pensoft.net/fig/713490" pageId="0" pageNumber="425">2A</figureCitation>
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); hindwing unmodified. Head unmodified, with three ocelli; antennae unmodified. Tibial spur count 1, 3, 4. Color in alcohol brown.
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<emphasis bold="true" pageId="0" pageNumber="425">
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<emphasis italics="true" pageId="0" pageNumber="425">Genitalia</emphasis>
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.
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</emphasis>
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Abdominal sternum VII slender, elongate (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14A, D</figureCitation>
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). Sternum VIII with posteroventral production bearing prominent apical seta (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14A</figureCitation>
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); in ventral view posterior margin concave (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14D</figureCitation>
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). Segment IX anterolateral margin convex, posterolateral margin straight with slight irregularity (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14B</figureCitation>
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); dorsally, anterior margin concave, posterior margin broadly convex (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14C</figureCitation>
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). Tergum X with dorsal sclerite with irregular dorsal margin; ventral sclerite semi-elliptic with knoblike projection mesally on posterior margin; membranous apex subquadrate (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14B, C</figureCitation>
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). Subgenital plate with dorsal arm simple, with slight preapical emargination on dorsal margin, apex truncate (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14B</figureCitation>
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); ventral arm narrowing apically, ventral margin slightly sinuate, in ventral view subovate with narrow basal projection and small rounded apical emargination (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14B, D</figureCitation>
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). Inferior appendage broadest mesally, with small dorsal subapical peg-like seta, apex rounded (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14B</figureCitation>
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); ventrally slender, with digitate basal projections, apex slightly hooked on inner margin (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14D</figureCitation>
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). Phallus tubular basally, constricted at midlength with typical median complex bearing basal loop and pair of spherical
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<normalizedToken originalValue="“windows”">"windows"</normalizedToken>
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; apex membranous with internal sclerotized structures and two apical lobes extending dorsad (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Leucotrichia ruiteri sp. nov. Male genitalia: A segments VII-VIII and segment IX margin, lateral B segments IX-X, lateral (base of phallus crosshatched) C segments IX-X, dorsal D segments VII-IX, ventral E phallus, lateral F phallus, dorsal." figureDoi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425">14E, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1111.77371.figure14" httpUri="https://binary.pensoft.net/fig/713502" pageId="0" pageNumber="425" start="Figure 14" startId="F14">
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<paragraph pageId="0" pageNumber="425">
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<emphasis bold="true" pageId="0" pageNumber="425">Figure 14.</emphasis>
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<taxonomicName authorityName="Thomson & Armitage & Harris" authorityYear="2022" class="Insecta" family="Hydroptilidae" genus="Leucotrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucotrichia ruiteri" order="Trichoptera" pageId="0" pageNumber="425" phylum="Arthropoda" rank="species" species="ruiteri">
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<emphasis italics="true" pageId="0" pageNumber="425">Leucotrichia ruiteri</emphasis>
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</taxonomicName>
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sp. nov. Male genitalia:
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<emphasis bold="true" pageId="0" pageNumber="425">A</emphasis>
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segments VII-VIII and segment IX margin, lateral
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<emphasis bold="true" pageId="0" pageNumber="425">B</emphasis>
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segments IX-X, lateral (base of phallus crosshatched)
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<emphasis bold="true" pageId="0" pageNumber="425">C</emphasis>
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segments IX-X, dorsal
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<emphasis bold="true" pageId="0" pageNumber="425">D</emphasis>
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segments VII-IX, ventral
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<emphasis bold="true" pageId="0" pageNumber="425">E</emphasis>
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phallus, lateral
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<emphasis bold="true" pageId="0" pageNumber="425">F</emphasis>
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phallus, dorsal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="distribution">
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<paragraph pageId="0" pageNumber="425">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="425">Panama.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="425" type="etymology">
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<paragraph pageId="0" pageNumber="425">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="425">This species is named in honor and memory of Dave Ruiter, a passionate and enthusiastic caddisfly researcher and good friend, who recently passed away.</paragraph>
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</subSubSection>
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</treatment>
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</document> |