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<document id="7B8EEC70FBABBBFD405F7096EA41FEC6" ID-CLB-Dataset="29933" ID-DOI="10.11646/zootaxa.4429.3.1" ID-GBIF-Dataset="249faa66-1f6a-44a0-b7f2-f12acaf62e5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1285294" ID-ZooBank="004C0DCB-9B63-4A44-A1BB-2D98ADFB1CEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528403104686" checkinUser="plazi" docAuthor="Ferro, Gustavo Borges &amp; Marshall, Stephen A." docDate="2018" docId="0A6A87F2FFF08017FF10FEF7FF0CF803" docLanguage="en" docName="zootaxa.4429.3.1.pdf" docOrigin="Zootaxa 4429 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiacephala umbra Ferro &amp; Marshall 2018, new species" docType="treatment" docUuid="F7B31E66-1FA6-4E9B-B2BB-88876075558F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="452" masterDocId="F653FF8AFFC38024FF87FF98FFFDFFB9" masterDocTitle="A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)" masterLastPageNumber="458" masterPageNumber="401" pageNumber="452" updateTime="1698548227473" updateUser="ExternalLinkService">
<mods:mods id="F4E748ED63258947EAD86C664A6EA0D4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A11A376D2BA0BAF5CBA4487EBD6BE8F3">
<mods:title id="6B0535FEA7BE26AD9905932A1299EC12">A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)</mods:title>
</mods:titleInfo>
<mods:name id="A6F239883BDC0AA69211657EB190DC3F" type="personal">
<mods:role id="BDC599683A92B265599B013069946E6E">
<mods:roleTerm id="15C4F3391C3D15E27AA24892223FFD2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2A760402C44F04FD0955E2C39C646AE">Ferro, Gustavo Borges</mods:namePart>
</mods:name>
<mods:name id="51D06006A4237FE24CF927C0CE755CAF" type="personal">
<mods:role id="FAE8EE1F71641C6C7B2966AB8A27DD0F">
<mods:roleTerm id="9835532391111696B25595FBC7F7D228">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D6663530046DE101CF3C4A61560AA41">Marshall, Stephen A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2EB3077CBEF041029C3830B1217B8DA5">text</mods:typeOfResource>
<mods:relatedItem id="CD7F1EFE46E5B39E155E7E44263AFB2F" type="host">
<mods:titleInfo id="4AE6042326D9F295E5C503F3B89EFB67">
<mods:title id="F3E23E1525DEFFC61234CBB5B6696A7D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="75EF3623717C6C8944CD964FB8BDD1AA">
<mods:date id="EE57183DDD7F2144D46C653A0CA708DC">2018</mods:date>
<mods:detail id="719460B02A17564BDF97CD3DB39361AF" type="pubDate">
<mods:number id="6B72FB9D99E53E29FC289A49571D6D11">2018-06-07</mods:number>
</mods:detail>
<mods:detail id="E3D244C4DF03F5DC1C581B426CB88512" type="volume">
<mods:number id="8746A87F8B36AB9E10E5C2DDFF2D24CB">4429</mods:number>
</mods:detail>
<mods:detail id="4DE5211BBCD091417F5DA8A7B4E8C058" type="issue">
<mods:number id="BE064DE4B6E91B3D3AE4DBE4B3853F3C">3</mods:number>
</mods:detail>
<mods:extent id="13C76B9370FF41F4ACED67E0F22873CD" unit="page">
<mods:start id="099607F5EF23F6D562C35FD36589EA6C">401</mods:start>
<mods:end id="9917E4F46142C350E3DC468D31BA339F">458</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CF19215E9D2A93CBAC4CAC60B7155C60">journal article</mods:classification>
<mods:identifier id="B88568168460051285D7EC2AE0E2A03C" type="CLB-Dataset">29933</mods:identifier>
<mods:identifier id="816B98834BEB26EC8176865B958D80F8" type="DOI">10.11646/zootaxa.4429.3.1</mods:identifier>
<mods:identifier id="1AA3156F699F8ACD8259B759898A57FA" type="GBIF-Dataset">249faa66-1f6a-44a0-b7f2-f12acaf62e5b</mods:identifier>
<mods:identifier id="1D97A9A41F93A1D0A29E8CAFFF452596" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3136BC043BCAE170B964EAD42FADDAFE" type="Zenodo-Dep">1285294</mods:identifier>
<mods:identifier id="E6335EDEA73DE2B6F0ECAB88303D9B88" type="ZooBank">004C0DCB-9B63-4A44-A1BB-2D98ADFB1CEE</mods:identifier>
</mods:mods>
<treatment id="0A6A87F2FFF08017FF10FEF7FF0CF803" ID-DOI="http://doi.org/10.5281/zenodo.5997559" ID-GBIF-Taxon="145927991" ID-Zenodo-Dep="5997559" LSID="urn:lsid:zoobank.org:act:F7B31E66-1FA6-4E9B-B2BB-88876075558F" httpUri="http://treatment.plazi.org/id/0A6A87F2FFF08017FF10FEF7FF0CF803" lastPageNumber="452" pageId="51" pageNumber="452">
<subSubSection id="CAD9656FFFF08017FF10FEF7FED3FE12" pageId="51" pageNumber="452" type="nomenclature">
<paragraph id="827C36E4FFF08017FF10FEF7FDCBFE33" blockId="51.[151,566,367,427]" box="[151,566,367,394]" pageId="51" pageNumber="452">
<heading id="D9348188FFF08017FF10FEF7FDCBFE33" bold="true" box="[151,566,367,394]" fontSize="11" level="1" pageId="51" pageNumber="452" reason="1">
<emphasis id="B0B7EAF6FFF08017FF10FEF7FDCBFE33" bold="true" box="[151,566,367,394]" pageId="51" pageNumber="452">
<taxonomicName id="45C34D67FFF08017FF10FEF7FE5DFE30" ID-CoL="8BBXJ" authority="Ferro &amp; Marshall, 2018" authorityName="Ferro &amp; Marshall" authorityYear="2018" box="[151,416,367,393]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="452" phylum="Arthropoda" rank="species" species="umbra" status="sp. nov.">
<emphasis id="B0B7EAF6FFF08017FF10FEF7FE5DFE30" bold="true" box="[151,416,367,393]" italics="true" pageId="51" pageNumber="452">Cardiacephala umbra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB84578DFFF08017FE20FEE8FDCBFE33" box="[423,566,368,394]" pageId="51" pageNumber="452" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="827C36E4FFF08017FF10FE0AFED3FE12" blockId="51.[151,566,367,427]" box="[151,302,402,427]" pageId="51" pageNumber="452">
(
<figureCitation id="1AF82A61FFF08017FF18FE0AFEDBFE12" box="[159,294,402,427]" captionStart="FIGURES 93–96" captionStartId="52.[151,264,1222,1245]" captionTargetBox="[151,1436,157,1199]" captionTargetId="figure@52.[151,1436,157,1199]" captionTargetPageId="52" captionText="FIGURES 93–96. Cardiacephala umbra. 93, female, dorsolateral view; 94, female genitalia, ventral view; 95, male, dorsolateral view; 96, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285352/files/figure.png" pageId="51" pageNumber="452">Figs. 93–96</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFF08017FF10FE42FE1FFB1B" pageId="51" pageNumber="452" type="description">
<paragraph id="827C36E4FFF08017FF10FE42FAC6FD3A" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF10FE42FED6FE4A" bold="true" box="[151,299,474,499]" pageId="51" pageNumber="452">Description:</emphasis>
Body length
<quantity id="453B9B01FFF08017FE47FE42FDFEFE4B" box="[448,515,474,499]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="51" pageNumber="452" unit="mm" value="6.0">6 mm</quantity>
, wing
<quantity id="453B9B01FFF08017FDCBFE43FD72FE4B" box="[588,655,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="51" pageNumber="452" unit="mm" value="4.0">4 mm</quantity>
. Palpus dark brown. Clypeus dark brown with pale microsetulae. First flagellomere dark brown covered with pale microsetulae. Fronto-orbital plates and frontal vitta brown. Epicephalon and paracephalon dark brown. Row of katepisternal setae white. Mid and hind femora brown and narrowly white basally. Mid and hind femora narrowly white basally. Fore tibia dark brown. Mid and hind tibiae brown. Fore tarsomere one white, 2–5 dark brown. T2–T4 brownish. Oviscape brown. Epandrium yellow.
</paragraph>
<paragraph id="827C36E4FFF08017FF40FD16FDCDFCAA" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Frontal vitta with a narrowed depression centrally, reaching ocellar triangle. Inner vertical seta insertion flat, without tubercle. Paracephalon smooth, without swelling (not extending beyond the vertex). Vertex flattened. Supracervical setae, weak.</paragraph>
<paragraph id="827C36E4FFF08017FF40FC86FD36FC1F" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">Thorax: Mesonotum with white pruinosity. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing apex clear with three preapical black spots. Crossvein r-m with small dark spot. R4+5 and M1+2 ending separately in wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling.</paragraph>
<paragraph id="827C36E4FFF08017FF40FC29FE1FFB1B" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
Abdomen: Tergites with yellowish setulae. Female abdomen: Single spermathecal duct uniformly thick, ending in a cylindrical spermatheca (somewhat damaged on
<figureCitation id="1AF82A61FFF08017FCD9FC4DFC4BFC57" box="[862,950,981,1006]" captionStart="FIGURES 93–96" captionStartId="52.[151,264,1222,1245]" captionTargetBox="[151,1436,157,1199]" captionTargetId="figure@52.[151,1436,157,1199]" captionTargetPageId="52" captionText="FIGURES 93–96. Cardiacephala umbra. 93, female, dorsolateral view; 94, female genitalia, ventral view; 95, male, dorsolateral view; 96, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285352/files/figure.png" pageId="51" pageNumber="452">Fig. 94</figureCitation>
). Paired spermathecal duct divided into a narrow basal half and a broader distal half. Paired spermathecal duct stems with a small rounded swelling. Paired spermathecae striate and cylindrical, deeply invaginated from apex to base. Male abdomen: Ejaculatory apodeme bigger than epandrium and with a broad blade, wider than sperm pump (
<figureCitation id="1AF82A61FFF08017FC74FBD9FBB6FBE3" box="[1011,1099,1089,1114]" captionStart="FIGURES 93–96" captionStartId="52.[151,264,1222,1245]" captionTargetBox="[151,1436,157,1199]" captionTargetId="figure@52.[151,1436,157,1199]" captionTargetPageId="52" captionText="FIGURES 93–96. Cardiacephala umbra. 93, female, dorsolateral view; 94, female genitalia, ventral view; 95, male, dorsolateral view; 96, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285352/files/figure.png" pageId="51" pageNumber="452">Fig. 96</figureCitation>
). Phallapodeme extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium, ending in a phallic bulb, phallic bulb subequal in length and width.
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFF08017FF40FB35FE99F95B" pageId="51" pageNumber="452" type="materials_examined">
<paragraph id="827C36E4FFF08017FF40FB35FD32FAB7" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF40FB35FE84FB7F" bold="true" box="[199,377,1197,1222]" pageId="51" pageNumber="452">
<typeStatus id="5D788846FFF08017FF40FB35FEFFFB7F" box="[199,258,1197,1222]" pageId="51" pageNumber="452">Type</typeStatus>
material:
</emphasis>
<typeStatus id="5D788846FFF08017FE05FB35FE12FB7F" box="[386,495,1197,1222]" pageId="51" pageNumber="452" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="4007F806FFF08017FE7DFB36FD8FFB7F" box="[506,626,1198,1222]" country="Trinidad and Tobago" name="Tobago" pageId="51" pageNumber="452">
<collectingCountry id="FAD47674FFF08017FE7DFB36FD8FFB7F" box="[506,626,1198,1222]" name="Trinidad and Tobago" pageId="51" pageNumber="452">TOBAGO</collectingCountry>
</collectingRegion>
. Roxborough
<quantity id="453B9B01FFF08017FC94FB36FCA1FB7F" box="[787,860,1197,1222]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="452" unit="km" value="10.0">10 km</quantity>
NE. Gilpin trail,
<quantity id="453B9B01FFF08017FBA0FB35FB57FB7C" box="[1063,1194,1197,1222]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="51" pageNumber="452" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
, montane rain forest, FIT,
<date id="F67D1024FFF08017FF53FB49FE7DFB53" box="[212,384,1233,1258]" pageId="51" pageNumber="452" value="1993-06-26" valueMax="1993-06-30" valueMin="1993-06-26">26–30.VI.1993</date>
, S. &amp; J. Peck (
<specimenCount id="94C5FD6DFFF08017FDCDFB4AFD87FB50" box="[586,634,1234,1258]" pageId="51" pageNumber="452" type="female">1 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FD08FB49FD20FB53" box="[655,733,1233,1258]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00138259).
<typeStatus id="5D788846FFF08017FCEEFB4AFC32FB53" box="[873,975,1234,1258]" pageId="51" pageNumber="452" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="FAD47674FFF08017FC66FB4AFB91FB53" box="[993,1132,1234,1258]" name="Trinidad and Tobago" pageId="51" pageNumber="452">TRINIDAD</collectingCountry>
. Maracas, Maracas Falls,
<date id="F67D1024FFF08017FF10FB6EFEDFFAB7" box="[151,290,1270,1294]" pageId="51" pageNumber="452" value="1981-07-12">12.VII.1981</date>
, J. M. Heraty (
<specimenCount id="94C5FD6DFFF08017FE4BFB6EFE0BFAB4" box="[460,502,1270,1294]" pageId="51" pageNumber="452" type="male">1 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FD85FB6DFDADFAB7" box="[514,592,1269,1294]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
01086450).
</paragraph>
<paragraph id="827C36E4FFF08017FF40FA81FE99F95B" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF40FA81FDE9FA8B" bold="true" box="[199,532,1305,1330]" pageId="51" pageNumber="452">Other Material examined:</emphasis>
<collectingRegion id="4007F806FFF08017FDA2FA82FCEBFA8B" box="[549,790,1306,1330]" country="France" name="Guyane" pageId="51" pageNumber="452">
<collectingCountry id="FAD47674FFF08017FDA2FA82FCEBFA8B" box="[549,790,1306,1330]" name="French Guiana" pageId="51" pageNumber="452">FRENCH GUIANA</collectingCountry>
</collectingRegion>
. Montsinery, Carrefour Du Gallion, Emerald Jungle, Gelbschale,
<date id="F67D1024FFF08017FEA5FAA5FE37FAEF" box="[290,458,1341,1366]" pageId="51" pageNumber="452" value="2003-12-05" valueMax="2003-12-16" valueMin="2003-12-05">5–16.XII.2003</date>
, M. Kotrba (
<specimenCount id="94C5FD6DFFF08017FDE7FAA6FD74FAEC" box="[608,649,1342,1366]" pageId="51" pageNumber="452" type="female">1 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FD1FFAA6FD14FAEF" box="[664,745,1342,1366]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
); Nahe St. Jean,
<date id="F67D1024FFF08017FC2FFAA6FBC9FAEF" box="[936,1076,1342,1366]" pageId="51" pageNumber="452" value="2002-12-29">29.XII.2002</date>
, M. Kotrba (1 #f,
<specimenCount id="94C5FD6DFFF08017FA83FAA6FACFFAEC" box="[1284,1330,1342,1366]" pageId="51" pageNumber="452" type="male">1 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FAB8FAA6FA6DFAEF" box="[1343,1424,1342,1366]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
); Nahe Tresor Nat. Reserve, Na umgefallenen Bäumem,
<date id="F67D1024FFF08017FCD5FAFAFC23FAC3" box="[850,990,1378,1402]" pageId="51" pageNumber="452" value="2002-12-24">24.XII.2002</date>
, M. Kotrba (
<specimenCount id="94C5FD6DFFF08017FB16FAFAFB3BFAC0" box="[1169,1222,1378,1402]" pageId="51" pageNumber="452" type="female">8 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FB67FAFAFACCFAC3" box="[1248,1329,1378,1402]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
); Rorota Wasserspeicher, Nahe Cayenne,
<date id="F67D1024FFF08017FD8FFA1DFD69FA27" box="[520,660,1413,1438]" pageId="51" pageNumber="452" value="2002-12-26">26.XII.2002</date>
, M. Kotrba (
<specimenCount id="94C5FD6DFFF08017FCAFFA1EFCA8FA24" box="[808,853,1414,1438]" pageId="51" pageNumber="452" type="female">4 ♀</specimenCount>
,
<specimenCount id="94C5FD6DFFF08017FCE2FA1DFC6FFA24" box="[869,914,1413,1438]" pageId="51" pageNumber="452" type="male">5 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FC27FA1EFC0CFA27" box="[928,1009,1414,1438]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
);
<collectingRegion id="4007F806FFF08017FB81FA1EFB83FA27" box="[1030,1150,1414,1438]" country="Trinidad and Tobago" name="Tobago" pageId="51" pageNumber="452">
<collectingCountry id="FAD47674FFF08017FB81FA1EFB83FA27" box="[1030,1150,1414,1438]" name="Trinidad and Tobago" pageId="51" pageNumber="452">TOBAGO</collectingCountry>
</collectingRegion>
. Roxborough
<quantity id="453B9B01FFF08017FA98FA1EFA95FA27" box="[1311,1384,1413,1438]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="452" unit="km" value="10.0">10 km</quantity>
NE. Gilpin trail,
<quantity id="453B9B01FFF08017FEB5FA31FE41FA78" box="[306,444,1449,1474]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="51" pageNumber="452" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
, montane rain forest, FIT,
<date id="F67D1024FFF08017FC96FA31FC43FA7B" box="[785,958,1449,1474]" pageId="51" pageNumber="452" value="1993-06-26" valueMax="1993-06-30" valueMin="1993-06-26">26–30.VI.1993</date>
, S. &amp; J. Peck (
<specimenCount id="94C5FD6DFFF08017FB08FA32FB3FFA78" box="[1167,1218,1450,1474]" pageId="51" pageNumber="452" type="female">2 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FB5EFA31FADBFA7B" box="[1241,1318,1449,1474]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00138260,
<collectionCode id="E4D2AE21FFF08017FF10FA55FF1BFA5F" box="[151,230,1485,1510]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00138261);
<collectingCountry id="FAD47674FFF08017FEF5FA56FE00FA5F" box="[370,509,1486,1510]" name="Trinidad and Tobago" pageId="51" pageNumber="452">TRINIDAD</collectingCountry>
. Morne Blue, 2700’,
<date id="F67D1024FFF08017FC83FA55FC7CFA5F" box="[772,897,1485,1510]" pageId="51" pageNumber="452" value="1969-07-06">6.VII.1969</date>
, H. &amp; A. Howden (
<specimenCount id="94C5FD6DFFF08017FB07FA56FB53FA5C" box="[1152,1198,1486,1510]" pageId="51" pageNumber="452" type="female">1 ♀</specimenCount>
,
<specimenCount id="94C5FD6DFFF08017FB44FA56FB08FA5C" box="[1219,1269,1486,1510]" pageId="51" pageNumber="452" type="male">2 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FA8FFA55FABEFA5F" box="[1288,1347,1485,1510]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="452">CNC</collectionCode>
503497,
<collectionCode id="E4D2AE21FFF08017FF10FA69FF2FF9B3" box="[151,210,1521,1546]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="452">CNC</collectionCode>
503498,
<collectionCode id="E4D2AE21FFF08017FEB0FA69FE8EF9B3" box="[311,371,1521,1546]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="452">CNC</collectionCode>
503499); Saugre-Grande,
<date id="F67D1024FFF08017FD1AFA69FD04F9B3" box="[669,761,1521,1546]" pageId="51" pageNumber="452" value="1959-09">IX.1959</date>
(
<specimenCount id="94C5FD6DFFF08017FC97FA6AFCC3F9B0" box="[784,830,1522,1546]" pageId="51" pageNumber="452" type="male">1 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FCC8FA69FC76F9B3" box="[847,907,1521,1546]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="51" pageNumber="452">CNC</collectionCode>
503444); Windblow ridge, Simla House N of
<collectingRegion id="4007F806FFF08017FF10F98DFF1FF997" box="[151,226,1557,1582]" country="Trinidad and Tobago" name="Arima" pageId="51" pageNumber="452">Arima</collectingRegion>
,
<quantity id="453B9B01FFF08017FF75F98DFEBCF994" box="[242,321,1557,1582]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="51" pageNumber="452" unit="m" value="610.0">610 m</quantity>
,
<date id="F67D1024FFF08017FED5F98EFE32F997" box="[338,463,1558,1582]" pageId="51" pageNumber="452" value="1981-07-09">9.VII.1981</date>
, J.M. Heraty (
<specimenCount id="94C5FD6DFFF08017FD03F98EFD4FF994" box="[644,690,1558,1582]" pageId="51" pageNumber="452" type="female">1 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FD42F98EFCEEF997" box="[709,787,1558,1582]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00241882);
<collectingCountry id="FAD47674FFF08017FC27F98EFBB7F997" box="[928,1098,1558,1582]" name="Venezuela" pageId="51" pageNumber="452">VENEZUELA</collectingCountry>
.
<collectingRegion id="4007F806FFF08017FBDDF98EFB60F997" box="[1114,1181,1558,1582]" country="Venezuela" name="Sucre" pageId="51" pageNumber="452">Sucre</collectingRegion>
, El Rincón, nr. Road between Carúpano &amp; El Piler, along Rio El Rincón valley,
<date id="F67D1024FFF08017FC21F9A2FBCEF9EB" box="[934,1075,1594,1618]" pageId="51" pageNumber="452" value="1998-04-08" valueMax="1998-04-09" valueMin="1998-04-08">8–9.IV.1998</date>
, M. von Tschirnhaus (
<specimenCount id="94C5FD6DFFF08017FAD8F9A2FA68F9E8" box="[1375,1429,1594,1618]" pageId="51" pageNumber="452" type="female">3 ♀</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FF10F9C5FF19F9CF" box="[151,228,1629,1654]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256939,
<collectionCode id="E4D2AE21FFF08017FEE5F9C5FE52F9CF" box="[354,431,1629,1654]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256940,
<collectionCode id="E4D2AE21FFF08017FDA9F9C5FD80F9CF" box="[558,637,1629,1654]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256941);
<collectingRegion id="4007F806FFF08017FC84F9C6FCBBF9CF" box="[771,838,1630,1654]" country="Venezuela" name="Sucre" pageId="51" pageNumber="452">Sucre</collectingRegion>
, Puy-Puy,
<quantity id="453B9B01FFF08017FC38F9C5FC02F9CF" box="[959,1023,1629,1654]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="51" pageNumber="452" unit="km" value="32.0">32km</quantity>
E Carupano, plantations &amp; 2º forest near beach, sweep,
<date id="F67D1024FFF08017FE08F91AFDB1F923" box="[399,588,1666,1690]" pageId="51" pageNumber="452" value="1988-03-31" valueMax="1988-04-02" valueMin="1988-03-31">31.III–2.IV.1988</date>
, M. von Tschirnhaus (
<specimenCount id="94C5FD6DFFF08017FCEAF91AFC5FF920" box="[877,930,1666,1690]" pageId="51" pageNumber="452" type="female">2 ♀</specimenCount>
,
<specimenCount id="94C5FD6DFFF08017FC3EF91AFC12F920" box="[953,1007,1666,1689]" pageId="51" pageNumber="452" type="male">7 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFF08017FB82F919FBAFF923" box="[1029,1106,1665,1690]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256928,
<collectionCode id="E4D2AE21FFF08017FB5EF919FADBF923" box="[1241,1318,1665,1690]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256929,
<collectionCode id="E4D2AE21FFF08017FF10F93DFF19F907" box="[151,228,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256930,
<collectionCode id="E4D2AE21FFF08017FEF6F93DFE42F907" box="[369,447,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256931,
<collectionCode id="E4D2AE21FFF08017FDCCF93DFD65F907" box="[587,664,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256932,
<collectionCode id="E4D2AE21FFF08017FCA2F93DFC8FF907" box="[805,882,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256933,
<collectionCode id="E4D2AE21FFF08017FC78F93DFBB1F907" box="[1023,1100,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256934,
<collectionCode id="E4D2AE21FFF08017FB5EF93DFADBF907" box="[1241,1318,1701,1726]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256935,
<collectionCode id="E4D2AE21FFF08017FF10F951FF18F95B" box="[151,229,1737,1762]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="51" pageNumber="452">DEBU</collectionCode>
00256936).
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFF08017FF40F975FCA2F8BF" box="[199,863,1773,1798]" pageId="51" pageNumber="452" type="distribution">
<paragraph id="827C36E4FFF08017FF40F975FCA2F8BF" blockId="51.[151,1437,474,1978]" box="[199,863,1773,1798]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF40F975FE9FF8BF" bold="true" box="[199,354,1773,1798]" pageId="51" pageNumber="452">Distribution:</emphasis>
<collectingRegion id="4007F806FFF08017FEEEF975FDEFF8BF" box="[361,530,1773,1798]" country="France" name="Guyane" pageId="51" pageNumber="452">
<collectingCountry id="FAD47674FFF08017FEEEF975FDEFF8BF" box="[361,530,1773,1798]" name="French Guiana" pageId="51" pageNumber="452">French Guiana</collectingCountry>
</collectingRegion>
,
<collectingRegion id="4007F806FFF08017FD9AF975FD89F8BF" box="[541,628,1773,1798]" country="Trinidad and Tobago" name="Tobago" pageId="51" pageNumber="452">
<collectingCountry id="FAD47674FFF08017FD9AF975FD89F8BF" box="[541,628,1773,1798]" name="Trinidad and Tobago" pageId="51" pageNumber="452">Tobago</collectingCountry>
</collectingRegion>
,
<collectingCountry id="FAD47674FFF08017FDF9F975FD1DF8BF" box="[638,736,1773,1798]" name="Trinidad and Tobago" pageId="51" pageNumber="452">Trinidad</collectingCountry>
,
<collectingCountry id="FAD47674FFF08017FD6BF975FCA2F8BF" box="[748,863,1773,1798]" name="Venezuela" pageId="51" pageNumber="452">Venezuela</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFF08017FF40F889FF12F8F7" pageId="51" pageNumber="452" type="etymology">
<paragraph id="827C36E4FFF08017FF40F889FF12F8F7" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF40F889FEACF893" bold="true" box="[199,337,1809,1834]" pageId="51" pageNumber="452">Etymology:</emphasis>
The specific name
<taxonomicName id="45C34D67FFF08017FDC4F88AFD4FF893" authorityName="Ferro &amp; Marshall" authorityYear="2018" box="[579,690,1809,1834]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="452" phylum="Arthropoda" rank="species" species="umbra">
<emphasis id="B0B7EAF6FFF08017FDC4F88AFD4FF893" box="[579,690,1809,1834]" italics="true" pageId="51" pageNumber="452">C. umbra</emphasis>
</taxonomicName>
refers to the overall shadowed and opaque appearance of this species.
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFF08017FF40F8C1FF0CF803" pageId="51" pageNumber="452" type="discussion">
<paragraph id="827C36E4FFF08017FF40F8C1FF0CF803" blockId="51.[151,1437,474,1978]" pageId="51" pageNumber="452">
<emphasis id="B0B7EAF6FFF08017FF40F8C1FEACF8CB" bold="true" box="[199,337,1881,1906]" pageId="51" pageNumber="452">Comments:</emphasis>
This species most closely resembles
<taxonomicName id="45C34D67FFF08017FD78F8C2FC59F8CB" authorityName="Ferro &amp; Marshall" authorityYear="2018" baseAuthorityName="Walker" baseAuthorityYear="1852" box="[767,932,1881,1906]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="452" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B0B7EAF6FFF08017FD78F8C2FC59F8CB" box="[767,932,1881,1906]" italics="true" pageId="51" pageNumber="452">C. brevipennis</emphasis>
</taxonomicName>
, from which it is easily distinguished by its white fore tarsomere one and wing with three preapical black spots. Female and male genitalia are also very distinct.
</paragraph>
</subSubSection>
</treatment>
</document>
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