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<document id="DCAF891128DEB42F21ADF66B48889824" ID-CLB-Dataset="29933" ID-DOI="10.11646/zootaxa.4429.3.1" ID-GBIF-Dataset="249faa66-1f6a-44a0-b7f2-f12acaf62e5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1285294" ID-ZooBank="004C0DCB-9B63-4A44-A1BB-2D98ADFB1CEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528403104686" checkinUser="plazi" docAuthor="Ferro, Gustavo Borges &amp; Marshall, Stephen A." docDate="2018" docId="0A6A87F2FFCA802EFF10FB4DFA89FDD9" docLanguage="en" docName="zootaxa.4429.3.1.pdf" docOrigin="Zootaxa 4429 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiacephala aeruginosa Ferro &amp; Marshall 2018, new species" docType="treatment" docUuid="F1E125A7-F5A5-4B7F-8340-8960E9710232" docUuidSource="ZooBank" docVersion="10" lastPageNumber="411" masterDocId="F653FF8AFFC38024FF87FF98FFFDFFB9" masterDocTitle="A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)" masterLastPageNumber="458" masterPageNumber="401" pageNumber="410" updateTime="1698548227473" updateUser="ExternalLinkService">
<mods:mods id="5D905A7C2434482E21852687A7D2BBFB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B8A1EA3C0696A80FEB2FBBCBEECE2D28">A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae)</mods:title>
</mods:titleInfo>
<mods:name id="569A75C8ED5C7196439D771CAAC59B0E" type="personal">
<mods:role id="0444D21210F17176F74A6987DE56B51D">
<mods:roleTerm id="374607C0371AAE8952CF3A85F1CBF6D6">Author</mods:roleTerm>
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<mods:namePart id="2CDDAEF15811A858077BBF8EE6171A2F">Ferro, Gustavo Borges</mods:namePart>
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<mods:namePart id="3BAB21566521B11EBE3B952BE43BEB7A">Marshall, Stephen A.</mods:namePart>
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<mods:title id="B8AC99C42A222E073AAFC04DF12B3C29">Zootaxa</mods:title>
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<mods:part id="9754E3416E2654292232D0FD7C533E00">
<mods:date id="5EDBE38E98B83AF81E86FB6DB8B6B40D">2018</mods:date>
<mods:detail id="A7A21075FEA3650490EAB7FBA2B626DD" type="pubDate">
<mods:number id="27FB1C2C606A7255679F8D35979561C4">2018-06-07</mods:number>
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<mods:number id="7FE7BDBF19C13E111FCC06F376D8EE7D">4429</mods:number>
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<mods:number id="D45B83FC39838CA0FE2AD8C1D9B98910">3</mods:number>
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<mods:classification id="426049003F65DB83C190016A9B47A3DC">journal article</mods:classification>
<mods:identifier id="8D9CF0E6BE0123464107E08DD51753B3" type="CLB-Dataset">29933</mods:identifier>
<mods:identifier id="7E1E96AF380334D23109D80076E20781" type="DOI">10.11646/zootaxa.4429.3.1</mods:identifier>
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<treatment id="0A6A87F2FFCA802EFF10FB4DFA89FDD9" ID-DOI="http://doi.org/10.5281/zenodo.5997481" ID-GBIF-Taxon="145928016" ID-Zenodo-Dep="5997481" LSID="urn:lsid:zoobank.org:act:F1E125A7-F5A5-4B7F-8340-8960E9710232" httpUri="http://treatment.plazi.org/id/0A6A87F2FFCA802EFF10FB4DFA89FDD9" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
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<paragraph id="827C36E4FFCA802DFF10FB4DFD93FB56" blockId="9.[151,622,1237,1297]" box="[151,622,1237,1263]" pageId="9" pageNumber="410">
<heading id="D9348188FFCA802DFF10FB4DFD93FB56" bold="true" box="[151,622,1237,1263]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="B0B7EAF6FFCA802DFF10FB4DFD93FB56" bold="true" box="[151,622,1237,1263]" pageId="9" pageNumber="410">
<taxonomicName id="45C34D67FFCA802DFF10FB4DFE2AFB56" ID-CoL="8BBWJ" authority="Ferro &amp; Marshall, 2018" authorityName="Ferro &amp; Marshall" authorityYear="2018" box="[151,471,1237,1263]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="aeruginosa" status="sp. nov.">
<emphasis id="B0B7EAF6FFCA802DFF10FB4DFE2AFB56" bold="true" box="[151,471,1237,1263]" italics="true" pageId="9" pageNumber="410">Cardiacephala aeruginosa</emphasis>
</taxonomicName>
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</emphasis>
</heading>
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(
<figureCitation id="1AF82A61FFCA802DFF18FB60FEF4FAA8" box="[159,265,1272,1297]" captionStart="FIGURES 58" captionStartId="10.[151,264,1875,1898]" captionTargetBox="[151,1436,663,1853]" captionTargetId="figure@10.[151,1436,663,1853]" captionTargetPageId="10" captionText="FIGURES 58. Cardiacephala aeruginosa. 5, female, lateral view; 6, female genitalia, ventral view; 7, male, dorsal view; 8, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285300/files/figure.png" pageId="9" pageNumber="410">Figs. 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFCA802EFF10FAD8FBD3FF41" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="description">
<paragraph id="827C36E4FFCA802DFF10FAD8FD65F9B4" blockId="9.[151,1437,1344,2020]" pageId="9" pageNumber="410">
<emphasis id="B0B7EAF6FFCA802DFF10FAD8FED6FAE0" bold="true" box="[151,299,1344,1369]" pageId="9" pageNumber="410">Description:</emphasis>
Body length
<quantity id="453B9B01FFCA802DFE5DFAD8FDD8FAE1" box="[474,549,1344,1368]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="410" unit="mm" value="8.0">8 mm</quantity>
, wing
<quantity id="453B9B01FFCA802DFDF8FAD8FD31FAE1" box="[639,716,1344,1369]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="410" unit="mm" value="5.0">5 mm</quantity>
. Palpus dark brown. Clypeus dark brown with sparse pale microsetulae. First flagellomere black. Frontal vitta brown. Fronto-orbital plates, paracephalon and epicephalon darkened. Thorax brownish and slightly ferruginous, opaque. Row of katepisternal setae black. Femora brown and shiny. Fore tibia dark brown. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. T1 anterior half dark brown, posterior half and T2 pale. T3T5 pale brown to orange. Oviscape brownish and shiny, apex dark brown. Epandrium rusty red.
</paragraph>
<paragraph id="827C36E4FFCA802DFF49F980FDF4F924" blockId="9.[151,1437,1344,2020]" pageId="9" pageNumber="410">Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. First flagellomere covered by golden microsetulae. Two frontal setae. Vertex flattened. Paracephalon smooth, without swelling (not extending beyond the vertex). Inner vertical setae insertions flat, without tubercles. Black supracervical setae sparse, weak.</paragraph>
<paragraph id="827C36E4FFCA802DFF40F930FE47F8ED" blockId="9.[151,1437,1344,2020]" pageId="9" pageNumber="410">
Thorax: Mesonotum with golden pruinosity. Scutellum upturned, triangular. Femora covered with sparse black microsetulae. Mid and hind tibia flattened, sulcus on outer face weak but somewhat curved (not extremely dilated as in
<taxonomicName id="45C34D67FFCA802DFF52F969FEC9F8B1" box="[213,308,1777,1800]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B0B7EAF6FFCA802DFF52F969FEC9F8B1" box="[213,308,1777,1800]" italics="true" pageId="9" pageNumber="410">C. nigra</emphasis>
</taxonomicName>
). Wing with stigmatal and discal bands, apex with three preapical black spots. Crossvein r-m with small dark spot. R4+5 and M1+2 ending separately in wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling.
</paragraph>
<paragraph id="827C36E4FFCA802EFF40F8C7FBD3FF41" blockId="9.[151,1437,1344,2020]" lastBlockId="10.[151,1437,151,608]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
Abdomen: Tergites with small brownish setulae. Female abdomen: Single spermathecal duct uniformly thick, ending in a cylindrical spermatheca (
<figureCitation id="1AF82A61FFCA802DFDCEF81BFD6FF825" box="[585,658,1923,1948]" captionStart="FIGURES 58" captionStartId="10.[151,264,1875,1898]" captionTargetBox="[151,1436,663,1853]" captionTargetId="figure@10.[151,1436,663,1853]" captionTargetPageId="10" captionText="FIGURES 58. Cardiacephala aeruginosa. 5, female, lateral view; 6, female genitalia, ventral view; 7, male, dorsal view; 8, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285300/files/figure.png" pageId="9" pageNumber="410">Fig. 6</figureCitation>
). Paired spermathecal duct divided into a narrow basal half and a broader distal half. Paired spermathecal duct stems thin and smooth in basal third, followed by a large rounded swelling and deeply invaginated sac-like (distally swollen and tapered basally) paired spermathecae. Male abdomen: Ejaculatory apodeme somewhat thin and about same length as epandrium, blade with a moderate size, as wide as sperm pump (
<figureCitation id="1AF82A61FFC9802EFE13FF24FE24FF6D" box="[404,473,188,213]" captionStart="FIGURES 58" captionStartId="10.[151,264,1875,1898]" captionTargetBox="[151,1436,663,1853]" captionTargetId="figure@10.[151,1436,663,1853]" captionTargetPageId="10" captionText="FIGURES 58. Cardiacephala aeruginosa. 5, female, lateral view; 6, female genitalia, ventral view; 7, male, dorsal view; 8, male genitalia, lateral view." httpUri="https://zenodo.org/record/1285300/files/figure.png" pageId="10" pageNumber="411">Fig. 8</figureCitation>
). Phallapodeme extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium, ending in phallic bulb, phallic bulb subequal in length and width.
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFC9802EFF40FE9CFAD0FE14" pageId="10" pageNumber="411" type="materials_examined">
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<typeStatus id="5D788846FFC9802EFF40FE9CFEFFFEA4" box="[199,258,260,285]" pageId="10" pageNumber="411">Type</typeStatus>
material:
</emphasis>
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:
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.
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,
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,
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,
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,
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,
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,
<collectorName id="2F365332FFC9802EFDC1FEB0FD10FEF9" box="[582,749,295,320]" pageId="10" pageNumber="411">S. A. Marshall</collectorName>
(
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,
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,
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, sequence
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).
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<typeStatus id="5D788846FFC9802EFF10FED5FEF4FEDC" box="[151,265,333,357]" pageId="10" pageNumber="411" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="FAD47674FFC9802EFE93FED4FE62FEDD" box="[276,415,332,356]" name="Ecuador" pageId="10" pageNumber="411">ECUADOR</collectingCountry>
.
<collectingRegion id="4007F806FFC9802EFE2DFED5FE14FEDD" box="[426,489,333,356]" country="Ecuador" name="Napo" pageId="10" pageNumber="411">Napo</collectingRegion>
, Yasuní National Park, Yasuní Research station,
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,
<date id="F67D1024FFC9802EFB7DFED4FA65FEDD" box="[1274,1432,332,356]" pageId="10" pageNumber="411" value="1998-11-03" valueMax="1998-11-20" valueMin="1998-11-03">320.XI.1998</date>
, T. Pape &amp; B. Viklund (
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,
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,
<collectionCode id="E4D2AE21FFC9802EFDB5FEE8FD75FE31" box="[562,648,368,392]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="411">ZMUC</collectionCode>
); Prov. Orellano, Yasuni Natl. Pk, Yasuni research Stn,
<geoCoordinate id="E7F75023FFC9802EFA9AFEF7FA66FE31" box="[1309,1435,367,392]" direction="south" orientation="latitude" pageId="10" pageNumber="411" precision="15" value="-0.6805556">0º4050” S</geoCoordinate>
<geoCoordinate id="E7F75023FFC9802EFF10FE0CFEE0FE15" box="[151,285,404,429]" direction="west" orientation="longitude" pageId="10" pageNumber="411" precision="15" value="-76.40056">76º242” W</geoCoordinate>
,
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,
<date id="F67D1024FFC9802EFEFAFE0CFDCEFE15" box="[381,563,404,428]" pageId="10" pageNumber="411" value="2009-04-28" valueMax="2009-05-08" valueMin="2009-04-28">28.IV8.V.2009</date>
. S. A. Marshall (
<specimenCount id="94C5FD6DFFC9802EFD73FE0CFCE3FE15" box="[756,798,404,428]" pageId="10" pageNumber="411" type="male">1 ♂</specimenCount>
,
<collectionCode id="E4D2AE21FFC9802EFCADFE0CFC8AFE14" box="[810,887,404,429]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="10" pageNumber="411">DEBU</collectionCode>
01015590, sequence
<collectionCode id="E4D2AE21FFC9802EFBE5FE0CFB33FE14" box="[1122,1230,404,429]" pageId="10" pageNumber="411">MYCRO</collectionCode>
26616).
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<emphasis id="B0B7EAF6FFC9802EFF40FE2FFE9FFE69" bold="true" box="[199,354,439,464]" pageId="10" pageNumber="411">Distribution:</emphasis>
<collectingCountry id="FAD47674FFC9802EFEEEFE2FFE34FE69" box="[361,457,439,464]" name="Ecuador" pageId="10" pageNumber="411">Ecuador</collectingCountry>
.
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</subSubSection>
<subSubSection id="CAD9656FFFC9802EFF40FE44FA66FE4C" box="[199,1435,476,501]" pageId="10" pageNumber="411" type="etymology">
<paragraph id="827C36E4FFC9802EFF40FE44FA66FE4C" blockId="10.[151,1437,151,608]" box="[199,1435,476,501]" pageId="10" pageNumber="411">
<emphasis id="B0B7EAF6FFC9802EFF40FE44FEACFE4C" bold="true" box="[199,337,476,501]" pageId="10" pageNumber="411">Etymology:</emphasis>
The specific name
<taxonomicName id="45C34D67FFC9802EFDABFE45FD36FE4D" authorityName="Ferro &amp; Marshall" authorityYear="2018" box="[556,715,477,500]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="aeruginosa">
<emphasis id="B0B7EAF6FFC9802EFDABFE45FD36FE4D" box="[556,715,477,500]" italics="true" pageId="10" pageNumber="411">C. aeruginosa</emphasis>
</taxonomicName>
refers to the mostly rusty red head and abdomen of this species.
</paragraph>
</subSubSection>
<subSubSection id="CAD9656FFFC9802EFF40FE67FA89FDD9" pageId="10" pageNumber="411" type="discussion">
<paragraph id="827C36E4FFC9802EFF40FE67FA89FDD9" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="411">
<emphasis id="B0B7EAF6FFC9802EFF40FE67FEACFDA1" bold="true" box="[199,337,511,536]" pageId="10" pageNumber="411">Comments:</emphasis>
This Ecuadorian species is distinct for its rusty red appearance and dorsally flattened head. The sulcus on the outer face of the mid and hind tibia is somewhat curved (but not extremely dilated as in
<taxonomicName id="45C34D67FFC9802EFA91FDBDFA8EFD85" box="[1302,1395,549,572]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B0B7EAF6FFC9802EFA91FDBDFA8EFD85" box="[1302,1395,549,572]" italics="true" pageId="10" pageNumber="411">C. nigra</emphasis>
</taxonomicName>
) as in
<taxonomicName id="45C34D67FFC9802EFF33FDD0FEAFFDD9" authorityName="Ferro &amp; Marshall" authorityYear="2018" box="[180,338,583,608]" class="Insecta" family="Micropezidae" genus="Cardiacephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="planivertex">
<emphasis id="B0B7EAF6FFC9802EFF33FDD0FEAFFDD9" box="[180,338,583,608]" italics="true" pageId="10" pageNumber="411">C. planivertex</emphasis>
</taxonomicName>
. Although similar, these species differ by their colour and genitalia (both in male and female).
</paragraph>
</subSubSection>
</treatment>
</document>