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<document id="DCBDFDDB71730F187340756042296295" ID-DOI="http://dx.doi.org/10.3897/zookeys.594.8768" ID-GBIF-Dataset="56393d29-ddfc-497f-be95-070616865240" ID-PMC="PMC4926701" ID-Pensoft-Pub="1313-2970-594-99" ID-PubMed="27408560" ID-ZooBank="4D2882008FF2422697EC3984E8EF56DD" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-594-99" ModsDocOrigin="ZooKeys 594" ModsDocTitle="A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1464725703941" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="09B11E36AEDDE7AC1803C1EEE7733EC7" docLanguage="en" docName="ZooKeys 594: 99-110" docOrigin="ZooKeys 594" docSource="http://dx.doi.org/10.3897/zookeys.594.8768" docTitle="Orthogonius sarawakensis Tian &amp; Deuve 2006" docType="treatment" docVersion="5" lastPageNumber="102" masterDocId="37113C2EFFB2286AFFB3FFF6FFD9DD1C" masterDocTitle="A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="110" masterPageNumber="99" pageNumber="102" updateTime="1701318027035" updateUser="plazi">
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<mods:title id="A4D42A107F1DF8A1C1AFC94D65351F30">A review of the sarawakensis species group of the ground beetle genus Orthogonius MacLeay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
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<mods:namePart id="BA1F6E30722446235452A85B88A18836">Tian, Mingyi</mods:namePart>
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<mods:namePart id="D4155E7CD1BEE3389DD54FBCA254B154">Deuve, Thierry</mods:namePart>
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<mods:date id="5E8CB8B0307DFF5B49AB0308D893AC9D">2016</mods:date>
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<mods:identifier id="C98346D6C24FFD3192BA28C618F5FEEA" type="DOI">http://dx.doi.org/10.3897/zookeys.594.8768</mods:identifier>
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<treatment id="09B11E36AEDDE7AC1803C1EEE7733EC7" ID-GBIF-Taxon="127901855" LSID="urn:lsid:plazi:treatment:09B11E36AEDDE7AC1803C1EEE7733EC7" httpUri="http://treatment.plazi.org/id/09B11E36AEDDE7AC1803C1EEE7733EC7" lastPageNumber="102" pageId="3" pageNumber="102">
<subSubSection id="62BA87DB48B242C45015027CACB550B5" pageId="3" pageNumber="102" type="multiple">
<paragraph id="8C2C0D1D915E472CE8BC23B4D9F2076B" pageId="3" pageNumber="102">
<pageBreakToken id="7EFF7E5808D6EA9F3EAA4EC07B81CDA6" pageId="3" pageNumber="102" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Carabidae
</paragraph>
</subSubSection>
<subSubSection id="A55F285D87346421B22A4E05F10C19D2" pageId="3" pageNumber="102" type="nomenclature">
<paragraph id="48CD5A3A2077E78E21067984972F2744" pageId="3" pageNumber="102">
<taxonomicName id="9992DBC632F2E16B97F46FE85414FB06" ID-CoL="6SWZZ" authority="Tian &amp; Deuve, 2006" authorityName="Tian &amp; Deuve" authorityYear="2006" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius sarawakensis" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="sarawakensis">Orthogonius sarawakensis Tian &amp; Deuve, 2006</taxonomicName>
Figs 1B, 2
<normalizedToken id="90F3635BA177C27B45D3DC10569EA711" originalValue="AC">A-C</normalizedToken>
</paragraph>
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<subSubSection id="6345C4718B3204307BB2AEAC45F75657" pageId="3" pageNumber="102" type="reference_group">
<paragraph id="DB0E3AC3674D7ADC2FBD4ABA14C36055" pageId="3" pageNumber="102">
<taxonomicName id="49CD008DBBC88943A06EF8399ACE976B" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius sarawakensis" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="sarawakensis">Orthogonius sarawakensis</taxonomicName>
Tian &amp; Deuve, 2006:
<bibRefCitation id="AB8260187A5555D3C863EAB06D724D98" author="Tian, MY" journalOrPublisher="Coleopteres" pageId="7" pageNumber="106" pagination="69 - 154" title="Contribution to the knowledge of the tribe Orthogoniini of the Oriental Region (Coleoptera: Caraboidea) (I, II)." volume="12" year="2006">Tian and Deuve 2006</bibRefCitation>
: 133.
</paragraph>
</subSubSection>
<subSubSection id="26B83C7E67A4C2F09E8C766ED08B04B4" pageId="3" pageNumber="102" type="type material">
<paragraph id="BF134574F3A2C0997B0AFE9575F62750" pageId="3" pageNumber="102">Type material.</paragraph>
<paragraph id="2007B8DE7E8E26A0833E4F0D532E1F8A" pageId="3" pageNumber="102">Length: 15.0 mm; width: 7.0 mm. Habitus as in Fig. 2A.</paragraph>
<caption id="4EA5CD1DEF717CB7216E7013286413AE" pageId="3" pageNumber="102">
<paragraph id="C73B08B14316499B21A143EE80D48A9A" pageId="3" pageNumber="102">
Figure 2.
<taxonomicName id="7FB3164E07FEC0E207CA4992BCA41F36" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius sarawakensis" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="sarawakensis">Orthogonius sarawakensis</taxonomicName>
Tian &amp; Deuve, 2006 A habitus, male, holotype B median lobe, right lateral view C apical lamella, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5F52D922B57A098224850CDDDD34CE8E" pageId="3" pageNumber="102" type="diagnosis">
<paragraph id="4527B544F3793A3CA9FF0E4BEC85E67F" pageId="3" pageNumber="102">Diagnosis.</paragraph>
<paragraph id="47C396DEA5C17A07AD601A77DB175BD8" pageId="3" pageNumber="102">Head dark brown, other parts of the body yellowish brown; head stout, as long as wide; labrum slightly emarginated at frontal margin; clypeus even near base, with two longitudinal furrows along both side, joining to frontal impressions respectively; mentum bisetose; pronotum transverse, PW/PL = 1.55, widest at about middle, strongly convex; lateral expanded margins almost flat, equal wide throughout; elytra elongate ovate, EL/LW = 1.54; base well-bordered; the 3rd interval with three setiferous pores, all are located on the interval; tarsal claws pectinate; prosternal process unbordered at apex, abdominal ventrite VII shallowly emarginated at apical margin in male.</paragraph>
<paragraph id="01BADAB537BB1D56E7EC634D189B2934" pageId="3" pageNumber="102">Male genitalia (Fig. 2B, C): Short and robust, expanded in median portion; in dorsal view, apical lobe elongated, gradually narrowed towards apex, with apical lamella evidently longer than wide.</paragraph>
<paragraph id="D15EBD0B6ADDD40B1D41E6AD2D2E01AE" pageId="3" pageNumber="102">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="0545B4CB7EC4305FBF9A7C2AD351E17F" pageId="3" pageNumber="102" type="remarks">
<paragraph id="6E8D3418D75205756E5810B2CC6F46C2" pageId="3" pageNumber="102">Remarks.</paragraph>
<paragraph id="51E9EF44A9D8CE9667CC511018E0635C" pageId="3" pageNumber="102">
Easily separated from its congeners,
<taxonomicName id="F06B0B2B7A359004474D44587C75D909" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius perakicus" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="perakicus">Orthogonius perakicus</taxonomicName>
Tian &amp; Deuve, 2007 and
<taxonomicName id="99C2ABFD2867BB48816B2C928DE65C4C" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius morvanianus" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="morvanianus">Orthogonius morvanianus</taxonomicName>
sp. n., by its unbordered prosternal process at apex, pointed elytral apex, and more elongated apical lobe of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="1D4F22B4E67C05FEF880692BE7E816ED" pageId="3" pageNumber="102" type="materials_examined">
<paragraph id="A1628FBA6A09D0446CF1DCBC468949CB" pageId="3" pageNumber="102">Material examined.</paragraph>
<paragraph id="827F7011BEBBC3F275EBA45DF2436DCA" pageId="3" pageNumber="102">1 male, the holotype, &quot;Nord Borneo, Mont Kina Balu, 5-8, 1903, John Waterstradt&quot;, in MNHN; 1 male, same data as in holotype, in SCAU.</paragraph>
</subSubSection>
<subSubSection id="B5CF7579A521E22E5B9CE583725E4981" pageId="3" pageNumber="102" type="distribution">
<paragraph id="DC0D642C3CC37BDACE744E17AB537592" pageId="3" pageNumber="102">Distribution.</paragraph>
<paragraph id="81BAB48AC376EE7A5E24BFAD3F06EA41" pageId="3" pageNumber="102">Malaysia (Sarawak).</paragraph>
</subSubSection>
</treatment>
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