192 lines
24 KiB
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192 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83781" ID-Pensoft-Pub="2193-0074-1-99" ID-Pensoft-UUID="2F5D6AE2EEB55A17ACF1623B06B4EA8D" ID-ZooBank="7A16698D4F1848D29D9651A6E0CC15AC" ModsDocID="2193-0074-25-1-99" checkinTime="1667366289781" checkinUser="pensoft" docAuthor="Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew & Moers, Thomas" docDate="2022" docId="09A843FFFAD65A05800819BD091D6844" docLanguage="en" docName="FossilRecord 25(1): 99-145" docOrigin="Fossil Record 25 (1)" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/fr.25.83781" docTitle="Latonia sp." docType="treatment" docVersion="1" id="2F5D6AE2EEB55A17ACF1623B06B4EA8D" lastPageNumber="99" masterDocId="2F5D6AE2EEB55A17ACF1623B06B4EA8D" masterDocTitle="Amphibian and reptilian fauna from the early Miocene of Echzell, Germany" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1667366289781" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Amphibian and reptilian fauna from the early Miocene of Echzell, Germany</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vasilyan, Davit</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8712-0678</mods:nameIdentifier>
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<mods:affiliation>JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">davit.vasilyan@jurassica.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cernansky, Andrej</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8920-2503</mods:nameIdentifier>
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<mods:affiliation>Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szyndlar, Zbigniew</mods:namePart>
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<mods:affiliation>Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moers, Thomas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2268-5824</mods:nameIdentifier>
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<mods:affiliation>Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Record</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-05-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>145</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/fr.25.83781</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83781</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2193-0074-1-99</mods:identifier>
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<mods:identifier type="ZooBank">7A16698D4F1848D29D9651A6E0CC15AC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2F5D6AE2EEB55A17ACF1623B06B4EA8D</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:09A843FFFAD65A05800819BD091D6844" httpUri="http://treatment.plazi.org/id/09A843FFFAD65A05800819BD091D6844" lastPageNumber="99" pageId="0" pageNumber="99">
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<subSubSection pageId="0" pageNumber="99" type="nomenclature">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName LSID="09A843FF-FAD6-5A05-8008-19BD091D6844" authority="sp." authorityName="sp." class="Amphibia" family="Alytidae" genus="Latonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Latonia" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">Latonia sp.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">Fig. 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="materials_examined">
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<paragraph pageId="0" pageNumber="99">Material.</paragraph>
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<paragraph pageId="0" pageNumber="99">76 maxillae HLMD-Ez 2130-2135, seven frontoparietals HLMD-Ez 2141-2144, one prooticoccipital HLMD-Ez 2127, six atlases HLMD-Ez 2116-2117, 23 presacral HLMD-Ez 2118-2120 and 22 sacral vertebrae HLMD-Ez 2121-2123, two costae (ribs) HLMD-Ez 2128, 23 urostyles 2124-2126, 58 ilia HLMD-Ez 2136-2140.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">Description.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Frontoparietals: All bones are very fragmentarily preserved, and all of them possess ornamentation made of a dense network of tubercles (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8A, C, E</figureCitation>
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). In ventral view, a number of ridges and structures are visible (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8B, D, F</figureCitation>
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); however, due to their preservation, any further description/anatomical identification is impossible.
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</paragraph>
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<caption doi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="99">
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<emphasis bold="true" pageId="0" pageNumber="99">Figure 8.</emphasis>
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<taxonomicName class="Amphibia" family="Alytidae" genus="Latonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Latonia" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Latonia</emphasis>
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</taxonomicName>
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sp. from Echzell. Frontoparietals HLMD-Ez 2141 (
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<emphasis bold="true" pageId="0" pageNumber="99">A, B</emphasis>
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), HLMD-Ez 2142 (
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<emphasis bold="true" pageId="0" pageNumber="99">C, D</emphasis>
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), HLMD-Ez 2143 (
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<emphasis bold="true" pageId="0" pageNumber="99">E, F</emphasis>
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). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (
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<emphasis bold="true" pageId="0" pageNumber="99">I-L</emphasis>
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). Atlas HLMD-Ez 2116 (
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<emphasis bold="true" pageId="0" pageNumber="99">M, N</emphasis>
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). Presacral vertebrae HLMD-Ez 2118 (
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<emphasis bold="true" pageId="0" pageNumber="99">O, P</emphasis>
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) and HLMD-Ez 2119 (
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<emphasis bold="true" pageId="0" pageNumber="99">Q-T</emphasis>
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). Sacral verterba HLMD-Ez 2122 (
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<emphasis bold="true" pageId="0" pageNumber="99">W-Y</emphasis>
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). Urostyle HLMD-Ez 2124 (
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<emphasis bold="true" pageId="0" pageNumber="99">Z</emphasis>
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) and HLMD-Ez 2125 (
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<emphasis bold="true" pageId="0" pageNumber="99">AA</emphasis>
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). Ilia HLMD-Ez2137 (
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<emphasis bold="true" pageId="0" pageNumber="99">AB</emphasis>
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), HLMD-Ez2138 (
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<emphasis bold="true" pageId="0" pageNumber="99">AC-AE</emphasis>
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), HLMD-Ez 2140 (
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<emphasis bold="true" pageId="0" pageNumber="99">AF</emphasis>
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) and HLMD-Ez2139 (
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<emphasis bold="true" pageId="0" pageNumber="99">AG-AI</emphasis>
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). Bones are figures in (
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<emphasis bold="true" pageId="0" pageNumber="99">A, C, E, L, O, T, X, Z, AA</emphasis>
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) dorsal, (
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<emphasis bold="true" pageId="0" pageNumber="99">B, D, F, K, M, R</emphasis>
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) ventral, (
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<emphasis bold="true" pageId="0" pageNumber="99">G</emphasis>
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) labial, (
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<emphasis bold="true" pageId="0" pageNumber="99">H</emphasis>
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) lingual, (
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<emphasis bold="true" pageId="0" pageNumber="99">I, Q, Y, Ad, Ah</emphasis>
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) posterior, (
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<emphasis bold="true" pageId="0" pageNumber="99">J, N, P, S, W</emphasis>
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) anterior and (
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<emphasis bold="true" pageId="0" pageNumber="99">AB, AC, AE, AF, AG, AI</emphasis>
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) lateral views. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="99">
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Maxillae: All bones are fragmentarily preserved. Their labial surface is smooth and does not possess any ornamentation (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8G</figureCitation>
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). Only in (HLMD-Ez 2132), the labial surface is covered by some irregularities. In lingual view, the horizontal lamina has a rounded surface and reduces height anteriorly (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8H</figureCitation>
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). Posteriorly, it terminates with a medioposteriorly projecting pterygoid process. The posterior depression is moderately developed. The margo orbitalis is slightly concave.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Prooticoccipital: One preserved prooticoccipital (HLMD-Ez 2127) consists of both fused prootic and lateral occipital processes (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8I-L</figureCitation>
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). It displays a distinct supracondylar depression. The prominentia ductus semicircularis posterioris is present as a distinct crest. It starts at the base of the lateral prootic process and projects medioanteriorly until the articulation surface with the frontoparietal. A foramen is present at the ventral base of the lateral prootic process. The sulcus venae jugularis is present as a horizontal groove. The fenestra ovalis is massive (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8K</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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The atlas has a dorsoventrally flattened centrum (HLMD-Ez 2116, Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8N</figureCitation>
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). The neural arch is not preserved. The crista ventralis is well developed. The condyloid fossae are separated from each other by a notch. The opisthocoelous presacral vertebrae have massive, slightly dorsally compressed centrum. The transverse processes project laterally. The neural arch in a small-sized vertebra projects posterodorsally (HLMD-Ez 2118), and the neural spine is clearly visible (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8O, P</figureCitation>
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). The sacral vertebrae have one anterior and two posterior condyles with very rounded external surfaces (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8X, Y</figureCitation>
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). The neural arch is short. It measures as long as the bases of the transverse processes. The latter widens laterally and project lateroposteriorly. The prezygapophyses in both presacral and sacral vertebrae project laterodorsally (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8S, W</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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The urostyle possesses two condyloid fossae and two lateroposteriorly bending transverse processes (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Latonia sp. from Echzell. Frontoparietals HLMD-Ez 2141 (A, B), HLMD-Ez 2142 (C, D), HLMD-Ez 2143 (E, F). Maxilla HLMD-Ez 2133. Prooticoccipital HLMD-Ez 2127 (I-L). Atlas HLMD-Ez 2116 (M, N). Presacral vertebrae HLMD-Ez 2118 (O, P) and HLMD-Ez 2119 (Q-T). Sacral verterba HLMD-Ez 2122 (W-Y). Urostyle HLMD-Ez 2124 (Z) and HLMD-Ez 2125 (AA). Ilia HLMD-Ez 2137 (AB), HLMD-Ez 2138 (AC-AE), HLMD-Ez 2140 (AF) and HLMD-Ez 2139 (AG-AI). Bones are figures in (A, C, E, L, O, T, X, Z, AA) dorsal, (B, D, F, K, M, R) ventral, (G) labial, (H) lingual, (I, Q, Y, Ad, Ah) posterior, (J, N, P, S, W) anterior and (AB, AC, AE, AF, AG, AI) lateral views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure8" httpUri="https://binary.pensoft.net/fig/682228" pageId="0" pageNumber="99">8Z</figureCitation>
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, AA). The opening of the neural canal is rounded. The neural canal opens dorsally behind the transverse processes in the form of a narrow and long strip.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">The preserved ilia have low or moderately developed dorsal prominence. The dorsal protuberance has a flat surface and shows high variation in shape and size. It can be very reduced in the form of a small protuberance (HLMD-Ez 2137, Fig. 8AB) or rather well-developed elongate (HLMD-Ez 2140, Fig. 8AF) or short drop-shaped (HLMD-Ez 2140-2141, Fig. 8AF) structure. It is connected with the lateromedially compressed iliac shaft by a thin lamina which reduces in height anteriorly behind the dorsal protuberance. The acetabular region is well developed. The dorsal acetabular expansion is well-developed and has a triangular outline. The ventral acetabular expansion is rather reduced and widens ventrally, expanding below the ventral margin of the acetabular crest. The ventral portion of the acetabular crest is well developed and projects ventrolaterally. The supraacetabular fossa is absent or, if present, weakly developed. A distinct tubercular fossa is visible in the corner between dorsal prominence and iliac shaft. In both medial (Fig. 8AE, AI) and posterior views (Fig. 8AH), a well-developed interiliac tubercle is visible. In posterior view, the ilioischiatic juncture can be clearly divided into a massive ventral and a slender dorsal portions, which are delimited in the medial surface by a deep groove.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="remarks">
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<paragraph pageId="0" pageNumber="99">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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The herein described remains can be assigned to the genus
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<taxonomicName class="Amphibia" family="Alytidae" genus="Latonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Latonia" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Latonia</emphasis>
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</taxonomicName>
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based on the presence of the following features: 1) frontoparietal with the characteristic tubercular sculpture; 2) prooticooccipital bone with a distinct supracondylar depression; 3) atlas possessing ventral crest; 4) urostyle with lateroposteriorly projecting transverse processes; 5) the general morphology of ilium and presence of the interiliac tubercle etc. (
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<bibRefCitation author="Rocek, Z" journalOrPublisher="Abhandlungen und Berichte fuer Naturkunde" pageId="0" pageNumber="99" pagination="51 - 56" refId="B103" refString="Rocek, Z, 1994. A Review of the fossil Caudata of Europe. Abhandlungen und Berichte fuer Naturkunde 17: 51 - 56" title="A Review of the fossil Caudata of Europe." volume="17" year="1994">
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<normalizedToken originalValue="Roček">Rocek</normalizedToken>
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1994
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</bibRefCitation>
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). Due to the poor preservation of the material, any further taxonomic identification is impossible.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |