651 lines
81 KiB
XML
651 lines
81 KiB
XML
<document id="D675B58A478785979A147A856E9329ED" ID-CLB-Dataset="57778" ID-DOI="10.11646/zootaxa.5319.2.6" ID-GBIF-Dataset="49b790fb-5325-46da-aa80-d36d2e1fcc1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182539" ID-ZooBank="065A967F-2832-4BDF-890A-2EA852CACD74" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690278797849" checkinUser="plazi" docAuthor="Chaves, Gerardo, Ryan, Mason J., Bolaños, Federico, Márquez, Cruz, Köhler, Gunther & Poe, Steven" docDate="2023" docId="09619A08FFA9190E8FECF8D09C30FE81" docLanguage="en" docName="zootaxa.5319.2.6.pdf" docOrigin="Zootaxa 5319 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anolis robinsoni Chaves & Ryan & Bolaños & Márquez & Köhler & Poe 2023, new species" docType="treatment" docVersion="6" lastPageNumber="257" masterDocId="F558E270FFAC19068F7BFFD09E22FFE4" masterDocTitle="Two new species of semiaquatic Anolis (Squamata: Dactyloidae) from Costa Rica" masterLastPageNumber="262" masterPageNumber="249" pageNumber="254" updateTime="1699546558579" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="81FB91B9508B622FC26631950A23E20C">Two new species of semiaquatic Anolis (Squamata: Dactyloidae) from Costa Rica</mods:title>
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<mods:namePart id="506B7D602C164836BB03DCC8918100B3">Chaves, Gerardo</mods:namePart>
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<mods:nameIdentifier id="00E2491F440C39EA62DE3C19EDBE6CA2" type="ORCID">0000-0002-4301-6569</mods:nameIdentifier>
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<mods:affiliation id="4D961CD3D5A9F11AA9007E2230E5EE1D">Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & cachi 13 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4301 - 6569</mods:affiliation>
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<mods:nameIdentifier id="65C7856D39B9F5F079E090C32AF7C417" type="email">cachi13@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5AA6770766FB6D0DD20067BCDA1CDC68">Ryan, Mason J.</mods:namePart>
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<mods:affiliation id="7EC104DF98F8B626BD538C7F1760C6EC">Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA & Arizona Game and Fish Department, 5000 W Carefree Hwy, Phoenix, Arizona 85086</mods:affiliation>
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<mods:namePart id="96B9B3103B3030DFE761D6F62FD977CD">Bolaños, Federico</mods:namePart>
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<mods:nameIdentifier id="F9E0D404BECC90A99C0B8593E003E636" type="ORCID">0000-0002-7935-6418</mods:nameIdentifier>
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<mods:affiliation id="E6888F476742B5CA2B86266C75C9D26E">Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & Escuela de Biología, Universidad de Costa Rica, San Pedro, Costa Rica & federico. bolanos @ ucr. ac. cr; https: // orcid. org / 0000 - 0002 - 7935 - 6418</mods:affiliation>
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<mods:nameIdentifier id="CB2DC80FA45876905951D8B358FECC4A" type="email">federico.bolanos@ucr.ac.cr</mods:nameIdentifier>
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<mods:namePart id="A44089CA65937B91102A6BD3E1EA3C7B">Márquez, Cruz</mods:namePart>
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<mods:nameIdentifier id="375F2A448F24AA740F162444CE63D670" type="ORCID">0009-0003-2220-7379</mods:nameIdentifier>
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<mods:affiliation id="331816AC6428548B580D4D41CDA53879">Direccion, Conservacion, Restauracion y Desarrollo Sustentable de Ecosistemas Insulares; Parque Nacional Galapagos, Ecuador cruzkallw @ gmail. com; https: // orcid. org / 0009 - 0003 - 2220 - 7379</mods:affiliation>
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<mods:nameIdentifier id="4FFCAC2C699CFA18317E41A44610E1AA" type="email">cruzkallw@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F5938AC7980C67A8B29F4A09921448B4">Köhler, Gunther</mods:namePart>
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<mods:nameIdentifier id="307F22D0FC30C056156624265D7A5190" type="ORCID">0000-0002-2563-5331</mods:nameIdentifier>
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<mods:affiliation id="34868C3597ED891DDBB2B322F8387243">Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany gkoehler @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 2563 - 5331</mods:affiliation>
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<mods:nameIdentifier id="09BBCEB4A57D9E5C495E1D11F11A39FA" type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="34D47DDE72717CF1830F13964AE39F87">Poe, Steven</mods:namePart>
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<mods:affiliation id="18F06DA3A68F4AF6E9DA598AC572BEBC">Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA</mods:affiliation>
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<mods:title id="CAF6AD0CD869F1C0B85420B526A1680E">Zootaxa</mods:title>
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<mods:part id="3F0EFABF046487ECAC5A2D2B0947ADC0">
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<mods:date id="D9C05B7293AF6F8B736A2C20FCFAF08C">2023</mods:date>
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<mods:number id="DB52D2FAE143785C5A8AE084A874FC99">2023-07-25</mods:number>
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<mods:number id="ABCDD34945022AD019402AD51EAA2E96">5319</mods:number>
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<mods:url id="C4CBF72670CE7D9AA4A787EB28644E31">http://dx.doi.org/10.11646/zootaxa.5319.2.6</mods:url>
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<mods:classification id="0F41253A0F40032038747B6D5D92F37E">journal article</mods:classification>
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<mods:identifier id="98F4BC5886D20E9CBF87F8182908C566" type="CLB-Dataset">57778</mods:identifier>
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<treatment id="09619A08FFA9190E8FECF8D09C30FE81" ID-DOI="http://doi.org/10.5281/zenodo.8184545" ID-GBIF-Taxon="212419559" ID-Zenodo-Dep="8184545" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81" httpUri="http://treatment.plazi.org/id/09619A08FFA9190E8FECF8D09C30FE81" lastPageId="8" lastPageNumber="257" pageId="5" pageNumber="254">
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<heading id="DA3F9C72FFA919038FECF8D09FCDF8FE" bold="true" box="[151,495,1792,1819]" fontSize="11" level="1" pageId="5" pageNumber="254" reason="1">
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<emphasis id="B3BCF70CFFA919038FECF8D09FCDF8FE" bold="true" box="[151,495,1792,1819]" pageId="5" pageNumber="254">
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<taxonomicName id="46C8509DFFA919038FECF8D09F75F8FF" ID-CoL="675PJ" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[151,343,1792,1819]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="254" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
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||
<emphasis id="B3BCF70CFFA919038FECF8D09F75F8FF" bold="true" box="[151,343,1792,1819]" italics="true" pageId="5" pageNumber="254">Anolis robinsoni</emphasis>
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</taxonomicName>
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,
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||
<taxonomicNameLabel id="A88F4A77FFA919038E1EF8D09FCDF8FE" box="[357,495,1792,1818]" pageId="5" pageNumber="254" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C9D27895FFA919008FECF8989EF8FE35" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254" type="materials_examined">
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<paragraph id="81772B1EFFA919038FECF8989B54F862" blockId="5.[151,1436,1864,2035]" pageId="5" pageNumber="254">
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<materialsCitation id="31A02143FFA919038FECF8989B54F862" ID-GBIF-Occurrence="4164459313" collectingDate="1969-03-15" country="Costa Rica" county="Santa Marta de Santiago de Puriscal" elevation="800" latitude="9.7923" location="Oscar Blanco" longLatPrecision="1" longitude="-84.3953" municipality="Palma" pageId="5" pageNumber="254" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
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<emphasis id="B3BCF70CFFA919038FECF8989EDDF886" bold="true" box="[151,255,1864,1890]" pageId="5" pageNumber="254">
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<typeStatus id="5E7395BCFFA919038FECF8989EDDF886" box="[151,255,1864,1890]" pageId="5" pageNumber="254" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
. UCR 2463 (
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<figureCitation id="19F3379BFFA919038EF6F8989FEEF886" box="[397,460,1864,1890]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="5" pageNumber="254">Fig. 4</figureCitation>
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) collected
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<date id="F5760DDEFFA919038D44F8989CC0F886" box="[575,738,1864,1891]" pageId="5" pageNumber="254" value="1969-03-15">
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<collectingDate id="E532F436FFA919038D44F8989CC0F886" box="[575,738,1864,1891]" pageId="5" pageNumber="254" value="1969-03-15">15 March 1969</collectingDate>
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</date>
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by
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<location id="84177DC5FFA919038C73F8989DB8F886" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81:84177DC5FFA919038C73F8989DB8F886" box="[776,922,1864,1890]" country="Costa Rica" county="Santa Marta de Santiago de Puriscal" latitude="9.7923" longLatPrecision="1" longitude="-84.3953" municipality="Palma" name="Oscar Blanco" pageId="5" pageNumber="254" stateProvince="San Jose">Oscar Blanco</location>
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||
, an adult male from
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<collectingMunicipality id="6113B164FFA919038B09F8989A97F886" box="[1138,1205,1864,1890]" pageId="5" pageNumber="254">Palma</collectingMunicipality>
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stream bridge
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<quantity id="463086FBFFA919038A29F8989BBEF886" box="[1362,1436,1864,1891]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="254" unit="km" value="5.1">5.1 km</quantity>
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south of
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<collectingCounty id="68165392FFA919038F8FF8BC9C5AF862" box="[244,632,1900,1927]" pageId="5" pageNumber="254">Santa Marta de Santiago de Puriscal</collectingCounty>
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,
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<geoCoordinate id="E4FC4DD9FFA919038DF8F8BC9CD4F863" box="[643,758,1900,1927]" degrees="9.79230" direction="north" orientation="latitude" pageId="5" pageNumber="254" precision="1" value="9.7923">9.79230 N</geoCoordinate>
|
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, -
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<geoCoordinate id="E4FC4DD9FFA919038C73F8BC9DACF863" box="[776,910,1900,1927]" degrees="84.39530" direction="west" orientation="longitude" pageId="5" pageNumber="254" precision="1" value="-84.3953">84.39530 W</geoCoordinate>
|
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, ca.
|
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<quantity id="463086FBFFA919038CC6F8BC9A02F862" box="[957,1056,1900,1926]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="254" unit="m" value="800.0">
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<elevation id="0AE5CC2DFFA919038CC6F8BC9A02F862" box="[957,1056,1900,1926]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="254" unit="m" value="800.0">800 masl</elevation>
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||
</quantity>
|
||
,
|
||
<collectingRegion id="430CE5FCFFA919038B50F8BC9AD2F862" box="[1067,1264,1900,1926]" country="Costa Rica" name="San Jose" pageId="5" pageNumber="254">San José Province</collectingRegion>
|
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,
|
||
<collectingCountry id="F9DF6B8EFFA919038B81F8BC9B50F862" box="[1274,1394,1900,1926]" name="Costa Rica" pageId="5" pageNumber="254">Costa Rica</collectingCountry>
|
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.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA919008FBCF8409EF8FE35" blockId="5.[151,1436,1864,2035]" lastBlockId="6.[151,1437,151,465]" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254">
|
||
<materialsCitation id="31A02143FFA919038FBCF8409B32F82A" ID-GBIF-Occurrence="4164459307" collectingDate="2010-03-27" collectionCode="SMF" country="Costa Rica" elevation="825" latitude="9.90392" location="San Jose Province" longLatPrecision="1" longitude="-84.28169" pageId="5" pageNumber="254" specimenCode="SMF 92437" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<emphasis id="B3BCF70CFFA919038FBCF8409F6AF84E" bold="true" box="[199,328,1936,1962]" pageId="5" pageNumber="254">
|
||
<typeStatus id="5E7395BCFFA919038FBCF8409F18F84E" box="[199,314,1936,1962]" pageId="5" pageNumber="254" type="paratype">Paratypes</typeStatus>
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(
|
||
</emphasis>
|
||
all from
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||
<collectingCountry id="F9DF6B8EFFA919038EDAF8409C3AF84E" box="[417,536,1936,1962]" name="Costa Rica" pageId="5" pageNumber="254">Costa Rica</collectingCountry>
|
||
:
|
||
<collectingRegion id="430CE5FCFFA919038D5AF8409CC3F84E" box="[545,737,1936,1962]" country="Costa Rica" name="San Jose" pageId="5" pageNumber="254">San José Province</collectingRegion>
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||
<emphasis id="B3BCF70CFFA919038D9AF8409CC8F84E" bold="true" box="[737,746,1936,1962]" pageId="5" pageNumber="254">)</emphasis>
|
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.
|
||
<emphasis id="B3BCF70CFFA919038D8EF8409D5CF84E" bold="true" box="[757,894,1936,1962]" pageId="5" pageNumber="254">
|
||
<collectorName id="2C3D4EC8FFA919038D8EF8409D15F84E" box="[757,823,1936,1962]" pageId="5" pageNumber="254">Adult</collectorName>
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||
males
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</emphasis>
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||
:
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<specimenCode id="D16E8365FFA919038CF0F8409A2BF84E" box="[907,1033,1936,1962]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="254">SMF 92437</specimenCode>
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||
(collected
|
||
<date id="F5760DDEFFA919038B06F8409B3DF84E" box="[1149,1311,1936,1962]" pageId="5" pageNumber="254" value="2010-03-27">
|
||
<collectingDate id="E532F436FFA919038B06F8409B3DF84E" box="[1149,1311,1936,1962]" pageId="5" pageNumber="254" value="2010-03-27">27 March 2010</collectingDate>
|
||
</date>
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||
by
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||
<collectorName id="2C3D4EC8FFA919038A3FF8409EC1F82A" pageId="5" pageNumber="254">Gunther Köhler</collectorName>
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) from
|
||
<collectorName id="2C3D4EC8FFA919038E56F8659C67F82A" box="[301,581,1972,1999]" pageId="5" pageNumber="254">Zona Protectora El Rodeo</collectorName>
|
||
(
|
||
<collectorName id="2C3D4EC8FFA919038D2FF8649CFAF82A" box="[596,728,1972,1998]" pageId="5" pageNumber="254">Universidad</collectorName>
|
||
para la
|
||
<collectorName id="2C3D4EC8FFA919038C54F8649D7BF82A" box="[815,857,1972,1998]" pageId="5" pageNumber="254">Paz</collectorName>
|
||
),
|
||
<geoCoordinate id="E4FC4DD9FFA919038C17F8649DC2F82B" box="[876,992,1972,1999]" degrees="9.90392" direction="north" orientation="latitude" pageId="5" pageNumber="254" precision="1" value="9.90392">9.90392 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFA919038C88F8649A58F82B" box="[1011,1146,1972,1999]" degrees="84.28169" direction="west" orientation="longitude" pageId="5" pageNumber="254" precision="1" value="-84.28169">84.28169 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFA919038BD1F8649B32F82A" box="[1194,1296,1972,1999]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="5" pageNumber="254" unit="m" value="825.0">
|
||
<elevation id="0AE5CC2DFFA919038BD1F8649B32F82A" box="[1194,1296,1972,1999]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="5" pageNumber="254" unit="m" value="825.0">825 masl</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFA919008A60F8649F9EFF55" ID-GBIF-Occurrence="4164459311" collectingDate="2010-03-27" collectionCode="SMF" country="Costa Rica" elevation="780" lastPageId="6" lastPageNumber="255" latitude="9.90355" longLatPrecision="1" longitude="-84.28192" pageId="5" pageNumber="254" specimenCode="SMF 92438" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFA919038A60F8649BBEF82A" box="[1307,1436,1972,1998]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="254">SMF 92438</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFA919038E72F8089F93F816" box="[265,433,2008,2034]" pageId="5" pageNumber="254" value="2010-03-27">
|
||
<collectingDate id="E532F436FFA919038E72F8089F93F816" box="[265,433,2008,2034]" pageId="5" pageNumber="254" value="2010-03-27">27 March 2010</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFA919038EA6F8089CA8F816" box="[477,650,2008,2034]" pageId="5" pageNumber="254">Gunther Köhler</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFA919038DACF8099DD1F816" box="[727,1011,2008,2035]" pageId="5" pageNumber="254">Zona Protectora El Rodeo</collectorName>
|
||
(
|
||
<collectorName id="2C3D4EC8FFA919038B7FF8089AAAF816" box="[1028,1160,2008,2034]" pageId="5" pageNumber="254">Universidad</collectorName>
|
||
para la
|
||
<collectorName id="2C3D4EC8FFA919038B98F8089B2FF816" box="[1251,1293,2008,2034]" pageId="5" pageNumber="254">Paz</collectorName>
|
||
),
|
||
<geoCoordinate id="E4FC4DD9FFA919038A59F8089BBAF817" box="[1314,1432,2008,2035]" degrees="9.90355" direction="north" orientation="latitude" pageId="5" pageNumber="254" precision="1" value="9.90355">9.90355 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFAA19008FE5FF479F07FF55" box="[158,293,151,177]" degrees="84.28192" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="1" value="-84.28192">84.28192 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFAA19008E2DFF479F9EFF55" box="[342,444,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="255" unit="m" value="780.0">
|
||
<elevation id="0AE5CC2DFFAA19008E2DFF479F9EFF55" box="[342,444,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="255" unit="m" value="780.0">780 masl</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFAA19008EBCFF479A56FF1D" ID-GBIF-Occurrence="4164459317" collectingDate="2001-10-28" collectionCode="SMF" country="Costa Rica" county="Cerros de Escazu" elevation="780" latitude="9.8846" location="Zona Protectora El Rodeo" longLatPrecision="1" longitude="-84.2765" municipality="Rio Jaris" pageId="6" pageNumber="255" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
|
||
UCR 2464 (same data of the
|
||
<typeStatus id="5E7395BCFFAA19008C7FFF479D44FF55" box="[772,870,151,177]" pageId="6" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
); UCR 16048 (collected
|
||
<date id="F5760DDEFFAA19008B08FF479B0BFF55" box="[1139,1321,151,177]" pageId="6" pageNumber="255" value="2001-10-28">
|
||
<collectingDate id="E532F436FFAA19008B08FF479B0BFF55" box="[1139,1321,151,177]" pageId="6" pageNumber="255" value="2001-10-28">28 October 2001</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFAA19008A29FF479EC6FF31" pageId="6" pageNumber="255">Mason J Ryan</collectorName>
|
||
,
|
||
<collectorName id="2C3D4EC8FFAA19008F8AFF6B9FEAFF31" box="[241,456,187,213]" pageId="6" pageNumber="255">Robert Puschendorf</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFAA19008D7BFF6B9CBEFF31" box="[512,668,187,213]" pageId="6" pageNumber="255">Brian Kubicki</collectorName>
|
||
) from the west slope of the
|
||
<collectingCounty id="68165392FFAA19008CA8FF6B9AB1FF31" box="[979,1171,187,213]" pageId="6" pageNumber="255">Cerros de Escazú</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6113B164FFAA19008BDBFF6B9B22FF31" box="[1184,1280,187,213]" pageId="6" pageNumber="255">Río Jaris</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFAA19008A74FF6B9BA4FF31" box="[1295,1414,187,213]" degrees="9.88460" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="1" value="9.8846">9.88460 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFAA19008FECFF0E9F3EFF1D" box="[151,284,222,249]" degrees="84.27650" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="1" value="-84.2765">84.27650 W</geoCoordinate>
|
||
, 500 ca. masl), close to the
|
||
<location id="84177DC5FFAA19008D38FF0F9D78FF1D" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81:84177DC5FFAA19008D38FF0F9D78FF1D" box="[579,858,223,249]" county="Cerros de Escazu" latitude="9.8846" longLatPrecision="1" longitude="-84.2765" municipality="Rio Jaris" name="Zona Protectora El Rodeo" pageId="6" pageNumber="255">Zona Protectora El Rodeo</location>
|
||
(
|
||
<location id="84177DC5FFAA19008C13FF0F9DCEFF1D" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81:84177DC5FFAA19008C13FF0F9DCEFF1D" box="[872,1004,223,249]" county="Cerros de Escazu" latitude="9.8846" longLatPrecision="1" longitude="-84.2765" municipality="Rio Jaris" name="Universidad" pageId="6" pageNumber="255">Universidad</location>
|
||
para
|
||
<collectorName id="2C3D4EC8FFAA19008B53FF0F9A4EFF1D" box="[1064,1132,223,249]" pageId="6" pageNumber="255">la Paz</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="31A02143FFAA19008B04FF0F9C53FEA4" ID-GBIF-Occurrence="4164459301" collectingDate="2007-12-27" collectionCode="MCZ" country="Costa Rica" elevation="1000" latitude="9.7889" location="Adult" longLatPrecision="1" longitude="-84.3944" municipality="Quebrada La Palma in Alto Palma de Puriscal" pageId="6" pageNumber="255" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectionCode id="E7D9B3DBFFAA19008B04FF0F9A98FF1D" box="[1151,1210,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="6" pageNumber="255" type="Museum">MCZ</collectionCode>
|
||
R-186162 (collected
|
||
<date id="F5760DDEFFAA19008FECFED39F44FEF9" box="[151,358,259,285]" pageId="6" pageNumber="255" value="2007-12-27">
|
||
<collectingDate id="E532F436FFAA19008FECFED39F44FEF9" box="[151,358,259,285]" pageId="6" pageNumber="255" value="2007-12-27">27 December 2007</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFAA19008EEBFED39FD9FEF9" box="[400,507,259,285]" pageId="6" pageNumber="255">Steve Poe</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFAA19008D49FED39C59FEF9" box="[562,635,259,285]" pageId="6" pageNumber="255">Mason</collectorName>
|
||
J.
|
||
<collectorName id="2C3D4EC8FFAA19008DE0FED39CF6FEF9" box="[667,724,259,285]" pageId="6" pageNumber="255">Ryan</collectorName>
|
||
) from
|
||
<collectingMunicipality id="6113B164FFAA19008C64FED39B30FEF9" box="[799,1298,259,285]" pageId="6" pageNumber="255">Quebrada La Palma in Alto Palma de Puriscal</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFAA19008A5AFED39BB5FEF9" box="[1313,1431,259,285]" degrees="9.78890" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="1" value="9.7889">9.78890 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFAA19008FE5FEF79F06FEA5" box="[158,292,295,321]" degrees="84.39440" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="1" value="-84.3944">84.39440 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFAA19008E2AFEF79FE1FEA5" box="[337,451,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="255" unit="m" value="1000.0">
|
||
<elevation id="0AE5CC2DFFAA19008E2AFEF79FE1FEA5" box="[337,451,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="255" unit="m" value="1000.0">1000 masl</elevation>
|
||
</quantity>
|
||
).
|
||
<emphasis id="B3BCF70CFFAA19008EAFFEF69C53FEA4" bold="true" box="[468,625,294,320]" pageId="6" pageNumber="255">
|
||
<location id="84177DC5FFAA19008EAFFEF69C34FEA4" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81:84177DC5FFAA19008EAFFEF69C34FEA4" box="[468,534,294,320]" country="Costa Rica" latitude="9.7889" longLatPrecision="1" longitude="-84.3944" municipality="Quebrada La Palma in Alto Palma de Puriscal" name="Adult" pageId="6" pageNumber="255" stateProvince="San Jose">Adult</location>
|
||
females
|
||
</emphasis>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="31A02143FFAA19008D05FEF79AA4FE81" ID-GBIF-Occurrence="4164459306" collectingDate="2010-03-27" collectionCode="SMF" country="Costa Rica" elevation="820" latitude="9.90364" longLatPrecision="1" longitude="-84.28182" pageId="6" pageNumber="255" specimenCode="SMF 92439, 92450-51" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<specimenCode id="D16E8365FFAA19008D05FEF79D22FEA5" box="[638,768,295,321]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="255">SMF 92439</specimenCode>
|
||
,
|
||
<specimenCode id="D16E8365FFAA19008C72FEF69D57FEA5" box="[777,885,294,321]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="255">92450–51</specimenCode>
|
||
(collected
|
||
<date id="F5760DDEFFAA19008C92FEF79AA9FEA5" box="[1001,1163,295,321]" pageId="6" pageNumber="255" value="2010-03-27">
|
||
<collectingDate id="E532F436FFAA19008C92FEF79AA9FEA5" box="[1001,1163,295,321]" pageId="6" pageNumber="255" value="2010-03-27">27 March 2010</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFAA19008BCAFEF79B79FEA5" box="[1201,1371,295,321]" pageId="6" pageNumber="255">Gunther Köhler</collectorName>
|
||
) from
|
||
<collectorName id="2C3D4EC8FFAA19008FECFE9B9F90FE81" box="[151,434,331,357]" pageId="6" pageNumber="255">Zona Protectora El Rodeo</collectorName>
|
||
(
|
||
<collectorName id="2C3D4EC8FFAA19008EBAFE9B9C67FE81" box="[449,581,331,357]" pageId="6" pageNumber="255">Universidad</collectorName>
|
||
para la
|
||
<collectorName id="2C3D4EC8FFAA19008DE4FE9B9CEBFE81" box="[671,713,331,357]" pageId="6" pageNumber="255">Paz</collectorName>
|
||
),
|
||
<geoCoordinate id="E4FC4DD9FFAA19008DA6FE9B9D70FE81" box="[733,850,331,357]" degrees="9.90364" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="1" value="9.90364">9.90364 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFAA19008C1DFE9B9DCFFE81" box="[870,1005,331,357]" degrees="84.28182" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="1" value="-84.28182">84.28182 W</geoCoordinate>
|
||
, ca.
|
||
<quantity id="463086FBFFAA19008B5BFE9B9AA4FE81" box="[1056,1158,331,357]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="255" unit="m" value="820.0">
|
||
<elevation id="0AE5CC2DFFAA19008B5BFE9B9AA4FE81" box="[1056,1158,331,357]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="255" unit="m" value="820.0">820 masl</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
; UCR 2558 (collected
|
||
<date id="F5760DDEFFAA19008AFAFE9A9F39FE6D" pageId="6" pageNumber="255" value="1969-03-15">15 March 1969</date>
|
||
by Oscar Blanco and Douglas Robinson) and
|
||
<materialsCitation id="31A02143FFAA19008C68FEBF9EF4FE35" ID-GBIF-Occurrence="4164459312" collectingDate="2007-12-27" collectionCode="MCZ" country="Costa Rica" elevation="1000" latitude="9.7889" location="Quebrada La Palma in Alto Palma de Puriscal" longLatPrecision="1" longitude="-84.3944" pageId="6" pageNumber="255" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectionCode id="E7D9B3DBFFAA19008C68FEBF9D6CFE6D" box="[787,846,367,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="6" pageNumber="255" type="Museum">MCZ</collectionCode>
|
||
R-186161 (collected
|
||
<date id="F5760DDEFFAA19008B43FEBF9B25FE6D" box="[1080,1287,367,393]" pageId="6" pageNumber="255" value="2007-12-27">
|
||
<collectingDate id="E532F436FFAA19008B43FEBF9B25FE6D" box="[1080,1287,367,393]" pageId="6" pageNumber="255" value="2007-12-27">27 December 2007</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="2C3D4EC8FFAA19008A4AFEBF9BBFFE6D" box="[1329,1437,367,393]" pageId="6" pageNumber="255">Steve Poe</collectorName>
|
||
and
|
||
<collectorName id="2C3D4EC8FFAA19008FBFFE439F2FFE49" box="[196,269,403,429]" pageId="6" pageNumber="255">Mason</collectorName>
|
||
J.
|
||
<collectorName id="2C3D4EC8FFAA19008E50FE439F46FE49" box="[299,356,403,429]" pageId="6" pageNumber="255">Ryan</collectorName>
|
||
) both of them from
|
||
<location id="84177DC5FFAA19008D40FE439A06FE49" LSID="urn:lsid:plazi:treatment:09619A08FFA9190E8FECF8D09C30FE81:84177DC5FFAA19008D40FE439A06FE49" box="[571,1060,403,429]" country="Costa Rica" latitude="9.7889" longLatPrecision="1" longitude="-84.3944" name="Quebrada La Palma in Alto Palma de Puriscal" pageId="6" pageNumber="255" stateProvince="San Jose">Quebrada La Palma in Alto Palma de Puriscal</location>
|
||
(
|
||
<geoCoordinate id="E4FC4DD9FFAA19008B4AFE439A84FE49" box="[1073,1190,403,429]" degrees="9.78890" direction="north" orientation="latitude" pageId="6" pageNumber="255" precision="1" value="9.7889">9.78890 N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="E4FC4DD9FFAA19008BCCFE439B1CFE49" box="[1207,1342,403,429]" degrees="84.39440" direction="west" orientation="longitude" pageId="6" pageNumber="255" precision="1" value="-84.3944">84.39440 W</geoCoordinate>
|
||
, ca
|
||
<quantity id="463086FBFFAA19008A1DFE439EECFE35" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="255" unit="m" value="1000.0">
|
||
<elevation id="0AE5CC2DFFAA19008A1DFE439EECFE35" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="255" unit="m" value="1000.0">1000 masl</elevation>
|
||
</quantity>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D5B77B96FFAA19008FE3FA109B5DF9C4" ID-DOI="http://doi.org/10.5281/zenodo.8182547" ID-Zenodo-Dep="8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="6" pageNumber="255" startId="6.[152,255,1472,1496]" targetBox="[157,1429,489,1449]" targetPageId="6" targetType="figure">
|
||
<paragraph id="81772B1EFFAA19008FE3FA109B5DF9C4" blockId="6.[151,1436,1472,1569]" pageId="6" pageNumber="255">
|
||
<emphasis id="B3BCF70CFFAA19008FE3FA109F35FA3C" bold="true" box="[152,279,1472,1496]" pageId="6" pageNumber="255">FIGURE 4.</emphasis>
|
||
A)
|
||
<taxonomicName id="46C8509DFFAA19008E40FA109FC3FA3C" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[315,481,1472,1496]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni">
|
||
<emphasis id="B3BCF70CFFAA19008E40FA109FC3FA3C" box="[315,481,1472,1496]" italics="true" pageId="6" pageNumber="255">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
in life; B) Male dewlap of
|
||
<taxonomicName id="46C8509DFFAA19008D96FA109DB1FA3C" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[749,915,1472,1496]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni">
|
||
<emphasis id="B3BCF70CFFAA19008D96FA109DB1FA3C" box="[749,915,1472,1496]" italics="true" pageId="6" pageNumber="255">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
(SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of
|
||
<taxonomicName id="46C8509DFFAA19008C09FA359DCCFA18" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[882,1006,1508,1532]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni">
|
||
<emphasis id="B3BCF70CFFAA19008C09FA359DCCFA18" box="[882,1006,1508,1532]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C9D27895FFAA19018FBCF9A29A70FEF9" lastPageId="7" lastPageNumber="256" pageId="6" pageNumber="255" type="diagnosis">
|
||
<paragraph id="81772B1EFFAA19018FBCF9A29A70FEF9" blockId="6.[151,1437,1650,2037]" lastBlockId="7.[151,1437,150,2013]" lastPageId="7" lastPageNumber="256" pageId="6" pageNumber="255">
|
||
<emphasis id="B3BCF70CFFAA19008FBCF9A29F17F968" bold="true" box="[199,309,1650,1676]" pageId="6" pageNumber="255">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFAA19008E3EF9A39FDAF969" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[325,504,1651,1677]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008E3EF9A39FDAF969" box="[325,504,1651,1677]" italics="true" pageId="6" pageNumber="255">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008D7AF9A39C7BF969" bold="true" box="[513,601,1651,1677]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008D7AF9A39C7BF969" box="[513,601,1651,1677]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a semiaquatic anole that is distinguished from all other
|
||
<taxonomicName id="46C8509DFFAA19008B9BF9A39B07F969" box="[1248,1317,1651,1677]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="genus">
|
||
<emphasis id="B3BCF70CFFAA19008B9BF9A39B07F969" box="[1248,1317,1651,1677]" italics="true" pageId="6" pageNumber="255">Anolis</emphasis>
|
||
</taxonomicName>
|
||
of Central America by male dewlap coloration (chocolate brown with ill-defined brick red horizontal streaks,
|
||
<figureCitation id="19F3379BFFAA19008BAAF9479B07F955" box="[1233,1317,1687,1713]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="6" pageNumber="255">Fig. 4B</figureCitation>
|
||
). The only other species with which
|
||
<taxonomicName id="46C8509DFFAA19008EDDF96B9C0BF931" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[422,553,1723,1749]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008EDDF96B9C0BF931" box="[422,553,1723,1749]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008D54F96B9CA6F931" bold="true" box="[559,644,1723,1749]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008D54F96B9CA6F931" box="[559,644,1723,1749]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
might be confused in the field are the other Middle American semiaquatic anoles
|
||
<taxonomicName id="46C8509DFFAA19008F99F90F9F45F91C" authorityName="Taylor" authorityYear="1956" box="[226,359,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFAA19008F99F90F9F45F91C" box="[226,359,1759,1785]" italics="true" pageId="6" pageNumber="255">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="46C8509DFFAA19008E0FF90F9FC5F91D" authorityName="Cope" authorityYear="1861" box="[372,487,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFAA19008E0FF90F9FC5F91D" box="[372,487,1759,1785]" italics="true" pageId="6" pageNumber="255">A. lionotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="46C8509DFFAA19008E8FF90F9CA7F91D" authorityName="Cope" authorityYear="1862" box="[500,645,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFAA19008E8FF90F9CA7F91D" box="[500,645,1759,1785]" italics="true" pageId="6" pageNumber="255">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="46C8509DFFAA19008DC4F90F9D09F91D" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[703,811,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFAA19008DC4F90F9D09F91D" box="[703,811,1759,1785]" italics="true" pageId="6" pageNumber="255">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="46C8509DFFAA19008C4CF90F9DFAF91D" authorityName="Cope" authorityYear="1861" box="[823,984,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="lionotus">
|
||
<emphasis id="B3BCF70CFFAA19008C4CF90F9DFAF91D" box="[823,984,1759,1785]" italics="true" pageId="6" pageNumber="255">Anolis lionotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="46C8509DFFAA19008B70F90F9ABEF91D" authorityName="Cope" authorityYear="1862" box="[1035,1180,1759,1785]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="poecilopus">
|
||
<emphasis id="B3BCF70CFFAA19008B70F90F9ABEF91D" box="[1035,1180,1759,1785]" italics="true" pageId="6" pageNumber="255">A. poecilopus</emphasis>
|
||
</taxonomicName>
|
||
are easily distinguished from
|
||
<taxonomicName id="46C8509DFFAA19008FA8F8D39F75F8F9" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[211,343,1795,1821]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008FA8F8D39F75F8F9" box="[211,343,1795,1821]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008E25F8D39F97F8F9" bold="true" box="[350,437,1795,1821]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008E25F8D39F97F8F9" box="[350,437,1795,1821]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by male dewlap color (dull orange-yellow in these species, chocolate brown with ill-defined brick red horizontal streaks in
|
||
<taxonomicName id="46C8509DFFAA19008EA7F8F79C42F8A5" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[476,608,1831,1857]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008EA7F8F79C42F8A5" box="[476,608,1831,1857]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008D1DF8F79C9EF8A5" bold="true" box="[614,700,1831,1857]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008D1DF8F79C9EF8A5" box="[614,700,1831,1857]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), presence of enlarged postcloacal scales in most males (absent in
|
||
<taxonomicName id="46C8509DFFAA19008AFEF8F79EDFF881" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008AFEF8F79EDFF881" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008E7EF89B9F79F881" bold="true" box="[261,347,1867,1893]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008E7EF89B9F79F881" box="[261,347,1867,1893]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), and presence of a longitudinal zone of 10–24 enlarged middorsal scales (0–3 scales enlarged in
|
||
<taxonomicName id="46C8509DFFAA19008AFEF89B9EDFF86D" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008AFEF89B9EDFF86D" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008E78F8BF9F7BF86D" bold="true" box="[259,345,1903,1929]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008E78F8BF9F7BF86D" box="[259,345,1903,1929]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
<taxonomicName id="46C8509DFFAA19008E15F8BF9C03F86C" authorityName="Taylor" authorityYear="1956" box="[366,545,1903,1929]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFAA19008E15F8BF9C03F86C" box="[366,545,1903,1929]" italics="true" pageId="6" pageNumber="255">Anolis aquaticus</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from
|
||
<taxonomicName id="46C8509DFFAA19008C37F8BF9DF2F86D" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[844,976,1903,1929]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008C37F8BF9DF2F86D" box="[844,976,1903,1929]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008CADF8BF9A0EF86D" bold="true" box="[982,1068,1903,1929]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008CADF8BF9A0EF86D" box="[982,1068,1903,1929]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by male dewlap color (orange-red with yellow in
|
||
<taxonomicName id="46C8509DFFAA19008E4FF8439F9BF848" authorityName="Taylor" authorityYear="1956" box="[308,441,1939,1965]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFAA19008E4FF8439F9BF848" box="[308,441,1939,1965]" italics="true" pageId="6" pageNumber="255">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="19F3379BFFAA19008EBFF8439C37F849" box="[452,533,1938,1965]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="6" pageNumber="255">Fig. 5A</figureCitation>
|
||
) and possession of larger dorsal scales of the head and body (e.g., 15–20 scales across the snout between second canthals in
|
||
<taxonomicName id="46C8509DFFAA19008D5CF8679C88F835" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[551,682,1975,2001]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008D5CF8679C88F835" box="[551,682,1975,2001]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008DCBF8679D24F835" bold="true" box="[688,774,1975,2001]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008DCBF8679D24F835" box="[688,774,1975,2001]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="19F3379BFFAA19008C6FF8679D45F835" box="[788,871,1975,2001]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="6" pageNumber="255">Fig. 4C</figureCitation>
|
||
),
|
||
<quantity id="463086FBFFAA19008C01F8679DE9F835" box="[890,971,1975,2001]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.556" metricValueMin="1.778" pageId="6" pageNumber="255" unit="in" value="10.5" valueMax="14.0" valueMin="7.0">7–14 in</quantity>
|
||
<taxonomicName id="46C8509DFFAA19008CAAF8679A75F834" authorityName="Taylor" authorityYear="1956" box="[977,1111,1975,2001]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFAA19008CAAF8679A75F834" box="[977,1111,1975,2001]" italics="true" pageId="6" pageNumber="255">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="19F3379BFFAA19008B1FF8679A94F835" box="[1124,1206,1974,2001]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="6" pageNumber="255">Fig. 5B</figureCitation>
|
||
,
|
||
<tableCitation id="CC4A1EA5FFAA19008BBBF8679B2CF835" box="[1216,1294,1975,2001]" captionStart="TABLE 1" captionStartId="4.[151,239,512,536]" captionTargetPageId="4" captionText="TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value" httpUri="http://table.plazi.org/id/D5B77B96FFA819028FECFDD09BBEFDFC" pageId="6" pageNumber="255" tableUuid="D5B77B96FFA819028FECFDD09BBEFDFC">Table 1</tableCitation>
|
||
) and the size of the middorsal caudal row scales relative to adjacent scales (no more than two times in
|
||
<taxonomicName id="46C8509DFFAA19008B36F80B9AF2F811" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1101,1232,2011,2037]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="255" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAA19008B36F80B9AF2F811" box="[1101,1232,2011,2037]" italics="true" pageId="6" pageNumber="255">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAA19008BADF80B9B0EF811" bold="true" box="[1238,1324,2011,2037]" pageId="6" pageNumber="255">
|
||
<taxonomicNameLabel id="A88F4A77FFAA19008BADF80B9B0EF811" box="[1238,1324,2011,2037]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="19F3379BFFAA19008A41F80B9BADF811" box="[1338,1423,2011,2037]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="6" pageNumber="255">Fig. 4D</figureCitation>
|
||
), more than three times in
|
||
<taxonomicName id="46C8509DFFAB19018EDAFF479C04FF54" authorityName="Taylor" authorityYear="1956" box="[417,550,151,177]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="aquaticus">
|
||
<emphasis id="B3BCF70CFFAB19018EDAFF479C04FF54" box="[417,550,151,177]" italics="true" pageId="7" pageNumber="256">A. aquaticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="19F3379BFFAB19018D4FFF479CA1FF55" box="[564,643,150,177]" captionStart="FIGURE 5" captionStartId="8.[152,255,1364,1388]" captionTargetBox="[260,1328,381,1341]" captionTargetId="figure-113@8.[260,1328,381,1341]" captionTargetPageId="8" captionText="FIGURE 5. A) Lateral and dewlap view of Anolis aquaticus in life. B) Dorsal view of head and C) Small middorsal scales in the base of the tail of A. aquaticus (UCR 5880) locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Steven Poe and B and C by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182549" httpUri="https://zenodo.org/record/8182549/files/figure.png" pageId="7" pageNumber="256">Fig 5C</figureCitation>
|
||
).
|
||
<taxonomicName id="46C8509DFFAB19018DEEFF479D0DFF55" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[661,815,151,177]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFAB19018DEEFF479D0DFF55" box="[661,815,151,177]" italics="true" pageId="7" pageNumber="256">Anolis barkeri</emphasis>
|
||
</taxonomicName>
|
||
is most easily distinguished from
|
||
<taxonomicName id="46C8509DFFAB19018BE7FF479B3DFF55" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[1180,1311,151,177]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018BE7FF479B3DFF55" box="[1180,1311,151,177]" italics="true" pageId="7" pageNumber="256">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018A5EFF479B59FF55" bold="true" box="[1317,1403,151,177]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018A5EFF479B59FF55" box="[1317,1403,151,177]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its larger size (maximum SVL
|
||
<quantity id="463086FBFFAB19018EA5FF6B9C0AFF31" box="[478,552,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="256" unit="mm" value="91.0">91 mm</quantity>
|
||
in
|
||
<taxonomicName id="46C8509DFFAB19018D31FF6B9C97FF31" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[586,693,187,213]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFAB19018D31FF6B9C97FF31" box="[586,693,187,213]" italics="true" pageId="7" pageNumber="256">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="463086FBFFAB19018DB9FF6B9D2EFF31" box="[706,780,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="256" unit="mm" value="74.0">74 mm</quantity>
|
||
in
|
||
<taxonomicName id="46C8509DFFAB19018C55FF6B9D93FF31" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[814,945,187,213]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018C55FF6B9D93FF31" box="[814,945,187,213]" italics="true" pageId="7" pageNumber="256">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018CCDFF6B9A29FF31" bold="true" box="[950,1035,187,213]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018CCDFF6B9A29FF31" box="[950,1035,187,213]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), lack of distinctly expanded toe pads (ratio width of expanded pads/width of distal phalanx
|
||
<quantity id="463086FBFFAB19018DA7FF0F9D6AFF1D" box="[732,840,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.588" metricValueMin="4.572" pageId="7" pageNumber="256" unit="in" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 in</quantity>
|
||
<taxonomicName id="46C8509DFFAB19018C35FF0F9DF0FF1D" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[846,978,223,249]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018C35FF0F9DF0FF1D" box="[846,978,223,249]" italics="true" pageId="7" pageNumber="256">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018CA3FF0F9A0CFF1D" bold="true" box="[984,1070,223,249]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018CA3FF0F9A0CFF1D" box="[984,1070,223,249]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
versus
|
||
<quantity id="463086FBFFAB19018BFBFF0F9ACEFF1D" box="[1152,1260,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="7" pageNumber="256" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
|
||
<taxonomicName id="46C8509DFFAB19018B89FF0F9B7CFF1D" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[1266,1374,223,249]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFAB19018B89FF0F9B7CFF1D" box="[1266,1374,223,249]" italics="true" pageId="7" pageNumber="256">A. barkeri</emphasis>
|
||
</taxonomicName>
|
||
), and presence of a double row of middorsal caudal scales (single row in
|
||
<taxonomicName id="46C8509DFFAB19018C1FFED39DC5FEF9" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[868,999,259,285]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018C1FFED39DC5FEF9" box="[868,999,259,285]" italics="true" pageId="7" pageNumber="256">A. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018C96FED39A60FEF9" bold="true" box="[1005,1090,259,285]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018C96FED39A60FEF9" box="[1005,1090,259,285]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFAB19018FBCFEF69AAFF9ED" pageId="7" pageNumber="256" type="description">
|
||
<paragraph id="81772B1EFFAB19018FBCFEF69D60FBF5" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCFEF69C7FFEA4" bold="true" box="[199,605,294,321]" pageId="7" pageNumber="256">
|
||
Description of the
|
||
<typeStatus id="5E7395BCFFAB19018EE4FEF79C23FEA5" box="[415,513,295,321]" pageId="7" pageNumber="256" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="19F3379BFFAB19018D6AFEF69C77FEA4" box="[529,597,294,320]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="7" pageNumber="256">Fig. 4</figureCitation>
|
||
)
|
||
</emphasis>
|
||
. Adult male as indicated by dewlap and everted hemipenes; SVL
|
||
<quantity id="463086FBFFAB19018A48FEF79BBAFEA5" box="[1331,1432,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="6.39" pageId="7" pageNumber="256" unit="mm" value="63.9">63.9 mm</quantity>
|
||
; HL
|
||
<quantity id="463086FBFFAB19018FBAFE9A9F07FE81" box="[193,293,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="7" pageNumber="256" unit="mm" value="17.5">17.5 mm</quantity>
|
||
; HL/SVL 0.28; HW
|
||
<quantity id="463086FBFFAB19018D7EFE9B9C4BFE81" box="[517,617,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="7" pageNumber="256" unit="mm" value="10.1">10.1 mm</quantity>
|
||
; EOH
|
||
<quantity id="463086FBFFAB19018DC9FE9B9D34FE81" box="[690,790,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="7" pageNumber="256" unit="mm" value="2.36">2.36 mm</quantity>
|
||
; IL
|
||
<quantity id="463086FBFFAB19018C44FE9B9D81FE81" box="[831,931,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="7" pageNumber="256" unit="mm" value="1.18">1.18 mm</quantity>
|
||
; IL/EOH 0.50; SLS-I
|
||
<quantity id="463086FBFFAB19018BEAFE9B9AD3FE81" box="[1169,1265,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="7" pageNumber="256" unit="mm" value="0.18">0.18 mm</quantity>
|
||
mm; AGL
|
||
<quantity id="463086FBFFAB19018A16FE9B9EE7FE6D" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="7" pageNumber="256" unit="mm" value="23.1">23.1 mm</quantity>
|
||
; FL
|
||
<quantity id="463086FBFFAB19018F83FEBF9F7CFE6D" box="[248,350,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="7" pageNumber="256" unit="mm" value="16.6">16.6 mm</quantity>
|
||
; T4L
|
||
<quantity id="463086FBFFAB19018EE4FEBE9C26FE6D" box="[415,516,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="7" pageNumber="256" unit="mm" value="15.1">15.1 mm</quantity>
|
||
; T4W
|
||
<quantity id="463086FBFFAB19018D2BFEBF9C97FE6D" box="[592,693,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="7" pageNumber="256" unit="mm" value="1.42">1.42 mm</quantity>
|
||
; TL
|
||
<quantity id="463086FBFFAB19018D92FEBF9D6FFE6D" box="[745,845,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="7" pageNumber="256" unit="mm" value="10.4">10.4 mm</quantity>
|
||
, tail complete; TL/SVL 0.38; T4L
|
||
<quantity id="463086FBFFAB19018BB7FEBE9B0FFE6D" box="[1228,1325,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="7" pageNumber="256" unit="mm" value="15.1">15.1 mm</quantity>
|
||
and T4W
|
||
<quantity id="463086FBFFAB19018FECFE439ED8FE49" box="[151,250,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="7" pageNumber="256" unit="mm" value="1.42">1.42 mm</quantity>
|
||
. 18 scales across the snout between second canthals; two scales separate the nasal opening from rostral scale; nine postrostral scales between supralabials; nine supralabial scales from rostral scale to level of middle of eye; six scales separating supraorbital semicircles; one rows of scales separate subocular scales from supralabials; two slightly elongate superciliary scale; eight scales separating interparietal scale from supraorbital semicircles; 12 enlarged scales in supraocular disk; 11 loreal scales in column just anterior to eye; 11 postmental scales posteriorly in contact with mental scale between infralabials; 16 scales in the loreal region; 19 rows of single scales in the dewlap (10–42 scales per row, rows somewhat irregular); 2 middorsal scale rows slightly larger than adjacent scales; 16 dorsal scales in 5% of SVL; 12 ventral scales in 5% of SVL; 220 scales around midbody; 23 expanded lamellae under fourth toe; 26 small middorsal caudal scales before the first large caudal scale (
|
||
<figureCitation id="19F3379BFFAB19018B46FD639AB6FD29" box="[1085,1172,691,717]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="7" pageNumber="256">Fig. 4D</figureCitation>
|
||
). Preoccipital, enlarged infralabial, tail crest, middorsal, and postcloacal scales absent; supraocular disc presents small scales. Frontal region of head concave; scales in supraocular disc unicarinate; rostral scale with weak cleft and overlaps mental scale; ear opening vertically ovoid; dorsal edge of ear lacks ornamentation; mental scale partially divided posteriorly and with its posterior edge convex; dewlap extends well on to chest with anterior insertion at level of anterior portion of the eye and without protruding scales at distal edge; dorsal scales small, keeled with two middorsal scale rows slightly larger than adjacent scales; ventral scales in diagonal rows, keeled and larger than dorsal scales; lateral scales homogeneous; tail is compressed and triangular in cross-section with the base taller than wide; middorsal caudal scales in single row and its size is less than three times than the adjacent scales (
|
||
<figureCitation id="19F3379BFFAB19018CF9FC039DF5FC09" box="[898,983,979,1005]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="7" pageNumber="256">Fig. 4D</figureCitation>
|
||
); the longest toe of adpressed limb reaches anterior to eye and, supradigital scales keeled and multicarinate.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFAB19018FBCFBCA9BACFB0D" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCFBCA9F77FBD0" bold="true" box="[199,341,1050,1076]" pageId="7" pageNumber="256">Color in life</emphasis>
|
||
(male based on field notes and color photographs;
|
||
<figureCitation id="19F3379BFFAB19018CF6FBCB9A19FBD1" box="[909,1083,1051,1077]" captionStart="FIGURE 4" captionStartId="6.[152,255,1472,1496]" captionTargetBox="[157,1429,489,1449]" captionTargetId="figure-181@6.[157,1429,489,1449]" captionTargetPageId="6" captionText="FIGURE 4. A) Anolis robinsoni in life; B) Male dewlap of Anolis robinsoni (SMF 92437) in life. C) Dorsal view of head and D) Small middorsal scales in the base of the tail of paratype (UCR 2464) of A. robinsoni locate between breakpoint (a) and the first enlarge middorsal caudal scale (b). Photo A by Robert Puschendorf, B by Gunther Köhler, and C and D by Gerardo Chaves." figureDoi="http://doi.org/10.5281/zenodo.8182547" httpUri="https://zenodo.org/record/8182547/files/figure.png" pageId="7" pageNumber="256">Fig. 4A and 4B</figureCitation>
|
||
). The dewlap of males in life is chocolate brown with lighter orange-brown bars located centrally. The dorsal ground color is olive to chestnut brown with transverse olive–green bars across dorsum down flanks extending to tail. The head is uniformly dark or light brown. A cream-colored line extends posteriorly from the labial scales down the side of the body. Scattered small greenish spots are present on the dorsum. Scales on the tip of the snout with black pigmentation. The dorsum and tail are marked with olive–green transverse bars and the limbs marked with pale green to yellowish transverse bars with small punctuations.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFAB19018FBCFB229DB4FA79" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCFB229F10FAE8" bold="true" box="[199,306,1266,1292]" pageId="7" pageNumber="256">Variation</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFAB19018E45FB239FCCFAE9" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[318,494,1267,1293]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018E45FB239FCCFAE9" box="[318,494,1267,1293]" italics="true" pageId="7" pageNumber="256">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018E89FB239C65FAE9" bold="true" box="[498,583,1267,1293]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018E89FB239C65FAE9" box="[498,583,1267,1293]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showed variation in the SVL of reproductive males (58.6 ± 11.0,
|
||
<quantity id="463086FBFFAB19018B89FB239BB3FAE9" box="[1266,1425,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="5.795" metricValueMax="7.38" metricValueMin="4.21" pageId="7" pageNumber="256" unit="mm" value="57.95" valueMax="73.8" valueMin="42.1">42.1–73.8 mm</quantity>
|
||
).
|
||
<tableCitation id="CC4A1EA5FFAB19018FECFAC79EC4FAD5" box="[151,230,1303,1329]" captionStart="TABLE 1" captionStartId="4.[151,239,512,536]" captionTargetPageId="4" captionText="TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value" httpUri="http://table.plazi.org/id/D5B77B96FFA819028FECFDD09BBEFDFC" pageId="7" pageNumber="256" tableUuid="D5B77B96FFA819028FECFDD09BBEFDFC">Table 1</tableCitation>
|
||
shows the variation in lepidosis. The gular region of females is white with brown streaks. Females have a small brown to black dewlap. Transverse, dorsal olive-green bands of males varied between 7–8, with 10–12 tail bands; females had 7–8 olive–green dorsal transverse bars with 9–11 tail bands. In life, the dorsal coloration of both sexes may vary with transverse bands darker when cold or stressed (e.g., when held in hand).
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFAB19018FBCFA769AAFF9ED" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCFA769FFAFA25" bold="true" box="[199,472,1446,1473]" pageId="7" pageNumber="256">Hemipenis morphology.</emphasis>
|
||
The almost completely everted hemipenis of SMF 92438 (
|
||
<figureCitation id="19F3379BFFAB19018B38FA779AB7FA25" box="[1091,1173,1447,1473]" captionStart="FIGURE 7" captionStartId="11.[152,255,1295,1319]" captionTargetBox="[356,1232,345,1266]" captionTargetId="figure-125@11.[356,1232,345,1272]" captionTargetPageId="11" captionText="FIGURE 7. Hemipenes of A. robinsoni (A), A. riparius (B), and Anolis aquaticus (C). The notable differences in appearance are due to preservation artifacts (i.e., the hemipenes of A. riparius and A. robinsoni are not completely everted). Scale bar = 1 mm for all photographs. Photos by Gunther Köhler." figureDoi="http://doi.org/10.5281/zenodo.8182555" httpUri="https://zenodo.org/record/8182555/files/figure.png" pageId="7" pageNumber="256">Fig. 7A</figureCitation>
|
||
) is a stout bilobed organ. The sulcus spermaticus is bordered by well-developed sulcal lips opening into two broad concave areas, one on each lobe. A small asulcate ridge is present. The apex is strongly calyculate, truncus with transverse folds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFAB19018FBCF9C39F15F97D" pageId="7" pageNumber="256" type="etymology">
|
||
<paragraph id="81772B1EFFAB19018FBCF9C39F15F97D" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCF9C39F61F9C9" bold="true" box="[199,323,1555,1581]" pageId="7" pageNumber="256">Etymology</emphasis>
|
||
. The specific epithet is a patronym in honor of the late Douglas C. Robinson, who contributed enormously to the herpetological knowledge of
|
||
<collectingCountry id="F9DF6B8EFFAB19018D69F9E79CA5F9B5" box="[530,647,1591,1617]" name="Costa Rica" pageId="7" pageNumber="256">Costa Rica</collectingCountry>
|
||
through extensive field collections and by inspiring a generation of Costa Rican biologists. He received his Ph.D. from Texas A&M University in 1968 and studied the Mexican semiaquatic anole,
|
||
<taxonomicName id="46C8509DFFAB19018FECF9AF9F12F97D" baseAuthorityName="Schmidt" baseAuthorityYear="1939" box="[151,304,1663,1689]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="barkeri">
|
||
<emphasis id="B3BCF70CFFAB19018FECF9AF9F12F97D" box="[151,304,1663,1689]" italics="true" pageId="7" pageNumber="256">Anolis barkeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFAB19018FBCF9729BA7F8CD" pageId="7" pageNumber="256" type="distribution">
|
||
<paragraph id="81772B1EFFAB19018FBCF9729BA7F8CD" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCF9729F71F958" bold="true" box="[199,339,1698,1724]" pageId="7" pageNumber="256">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="46C8509DFFAB19018E25F9739C2CF959" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[350,526,1699,1725]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018E25F9739C2CF959" box="[350,526,1699,1725]" italics="true" pageId="7" pageNumber="256">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018D69F9739C45F959" bold="true" box="[530,615,1699,1725]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018D69F9739C45F959" box="[530,615,1699,1725]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from the riparian gallery forests of the Río Candelaria basin (
|
||
<figureCitation id="19F3379BFFAB19018A34F9739BA9F959" box="[1359,1419,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[152,255,1647,1671]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure-155@2.[151,1436,453,1623]" captionTargetPageId="2" captionText="FIGURE 1. Map of Costa Rica showing collecting sites for Anolis aquaticus group. Caribbean individuals appear to be A. aquaticus, but we consider their status uncertain on geographic grounds." figureDoi="http://doi.org/10.5281/zenodo.8182541" httpUri="https://zenodo.org/record/8182541/files/figure.png" pageId="7" pageNumber="256">Fig. 1</figureCitation>
|
||
), the foothills of the Montañas de Turrubares in the northwestern section of the Cordillera de Talamanca, and Río Jaris along the western border of the Zona Protectora El Rodeo. These areas are within the Tropical Wet and Humid Premontane forest life zones (
|
||
<bibRefCitation id="E55956EFFFAB19018E35F8DF9FDEF8CD" author="Holdridge, L. R." box="[334,508,1807,1833]" pageId="7" pageNumber="256" refId="ref9012" refString="Holdridge, L. R. (1967) Life zone ecology. Tropical Science Center, San Jose, Costa Rica, 206 pp." type="book" year="1967">Holdridge 1967</bibRefCitation>
|
||
).
|
||
<taxonomicName id="46C8509DFFAB19018D75F8DF9C9DF8CD" authority="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe, 2023" authorityName="Chaves & Ryan & Bolaños & Márquez & Köhler & Poe" authorityYear="2023" box="[526,703,1807,1833]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="256" phylum="Chordata" rank="species" species="robinsoni" status="sp. nov.">
|
||
<emphasis id="B3BCF70CFFAB19018D75F8DF9C9DF8CD" box="[526,703,1807,1833]" italics="true" pageId="7" pageNumber="256">Anolis robinsoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3BCF70CFFAB19018DBFF8DF9D38F8CD" bold="true" box="[708,794,1807,1833]" pageId="7" pageNumber="256">
|
||
<taxonomicNameLabel id="A88F4A77FFAB19018DBFF8DF9D38F8CD" box="[708,794,1807,1833]" pageId="7" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been found at elevations between 500 and
|
||
<quantity id="463086FBFFAB19018A6BF8DF9BA3F8CD" box="[1296,1409,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="256" unit="m" value="1100.0">1100 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9D27895FFAB190E8FBCF8E39C30FE81" lastPageId="8" lastPageNumber="257" pageId="7" pageNumber="256" type="biology_ecology">
|
||
<paragraph id="81772B1EFFAB190E8FBCF8E39CF7FF31" blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1437,151,357]" lastPageId="8" lastPageNumber="257" pageId="7" pageNumber="256">
|
||
<emphasis id="B3BCF70CFFAB19018FBCF8E39F00F8A9" bold="true" box="[199,290,1843,1869]" pageId="7" pageNumber="256">Ecology</emphasis>
|
||
. Most of the following observations are from Márquez–Baltán (1994) and
|
||
<bibRefCitation id="E55956EFFFAB19018B11F8E39B7CF8A9" author="Marquez, C. & Mora, J. M. & Bolanos, F. & Rea, S." box="[1130,1374,1842,1869]" pageId="7" pageNumber="256" pagination="59 - 69" refId="ref9391" refString="Marquez, C., Mora, J. M., Bolanos, F. & Rea, S. (2005) Aspectos de la biologia poblacional en el campo de Anolis aquaticus, Sauria: Polychridae en Costa Rica. Ecologia Aplicada, 4, 59 - 69. https: // doi. org / 10.21704 / rea. v 4 i 1 - 2.299" type="journal article" year="2005">
|
||
Márquez
|
||
<emphasis id="B3BCF70CFFAB19018BAFF8E39B2FF8A9" box="[1236,1293,1843,1869]" italics="true" pageId="7" pageNumber="256">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
from Quebrada La Palma. During the day this species is found on bare rocks and boulders along streams with dense canopy cover. At night it is found sleeping on moss and vegetation on boulders near the splash zones of small and medium waterfalls. Both sexes and juveniles were found no more than two meters from the edge of a stream. The operational sex ratio is approximately equal proportions of males to females (
|
||
<bibRefCitation id="E55956EFFFAB19018C57F8139A28F839" author="Marquez, C. & Mora, J. M. & Bolanos, F. & Rea, S." box="[812,1034,1986,2013]" pageId="7" pageNumber="256" pagination="59 - 69" refId="ref9391" refString="Marquez, C., Mora, J. M., Bolanos, F. & Rea, S. (2005) Aspectos de la biologia poblacional en el campo de Anolis aquaticus, Sauria: Polychridae en Costa Rica. Ecologia Aplicada, 4, 59 - 69. https: // doi. org / 10.21704 / rea. v 4 i 1 - 2.299" type="journal article" year="2005">
|
||
Márquez
|
||
<emphasis id="B3BCF70CFFAB19018CEEF8139DE9F839" box="[917,971,1987,2013]" italics="true" pageId="7" pageNumber="256">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
), and males share territories with one to
|
||
<specimenCount id="97CEE097FFA4190E8FCFFF479F6AFF55" box="[180,328,151,177]" pageId="8" pageNumber="257" type="female">three females</specimenCount>
|
||
. The estimated population density varied from 100 to 200 individuals per hectare and population size fluctuated monthly with a peak during the dry season.
|
||
</paragraph>
|
||
<paragraph id="81772B1EFFA4190E8FBCFF0F9C30FE81" blockId="8.[151,1437,151,357]" pageId="8" pageNumber="257">
|
||
Reproduction occurs throughout the year with a lull in the wet season (May to November). Females lay
|
||
<specimenCount id="97CEE097FFA4190E8A4DFF0F9B50FF1D" box="[1334,1394,223,249]" pageId="8" pageNumber="257" type="egg">1 egg</specimenCount>
|
||
per clutch in cracks and interstitial spaces of boulders in streams. Up to
|
||
<specimenCount id="97CEE097FFA4190E8C16FED39D90FEF9" box="[877,946,259,285]" pageId="8" pageNumber="257" type="egg">6 eggs</specimenCount>
|
||
have been found together suggesting that this species lays eggs in communal nest sites. Incubation times ranged between 75–82 days, and hatchlings are approximately
|
||
<quantity id="463086FBFFA4190E8FECFE9A9EC1FE81" box="[151,227,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="257" unit="mm" value="25.0">25 mm</quantity>
|
||
SVL (
|
||
<bibRefCitation id="E55956EFFFA4190E8E52FE9B9C24FE81" author="Marquez, C. & Mora, J. M. & Bolanos, F. & Rea, S." box="[297,518,330,357]" pageId="8" pageNumber="257" pagination="59 - 69" refId="ref9391" refString="Marquez, C., Mora, J. M., Bolanos, F. & Rea, S. (2005) Aspectos de la biologia poblacional en el campo de Anolis aquaticus, Sauria: Polychridae en Costa Rica. Ecologia Aplicada, 4, 59 - 69. https: // doi. org / 10.21704 / rea. v 4 i 1 - 2.299" type="journal article" year="2005">
|
||
Márquez
|
||
<emphasis id="B3BCF70CFFA4190E8EEAFE9B9FE4FE81" box="[401,454,331,357]" italics="true" pageId="8" pageNumber="257">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |