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<document id="5CD1413708FB530ED57A5721B19630A9" ID-DOI="10.11646/zootaxa.3691.3.1" ID-GBIF-Dataset="ad034b98-4029-470c-9a6e-4b5d168b46e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="248009" ID-ZooBank="88298B5D-2941-4A52-8C47-0330168EA240" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460650775633" checkinUser="plazi" docAuthor="Walter, David Evans" docDate="2013" docId="094E480B5C20FFCBFF3E49786A53D09B" docLanguage="en" docName="zt03691p323.pdf" docOrigin="Zootaxa 3691 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sejus americanus Banks 1902" docType="treatment" docVersion="9" lastPageNumber="320" masterDocId="F57730735C2EFFD8FFA94C676921D33F" masterDocTitle="A new genus and family of sejine mites (Acari, Parasitiformes, Mesostigmata, Sejoidea) based on new species from Lord Howe Island and Brazil, and a redescription of Sejus americanus (Banks, 1902)" masterLastPageNumber="323" masterPageNumber="301" pageNumber="315" updateTime="1698337942205" updateUser="plazi">
<mods:mods id="FDE3F85161B8E798DE7052803D13C065" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7446E1699908DFC449EA43899474ABB">
<mods:title id="831AC11C9CDB635A6C57BCB8040B9EF0">A new genus and family of sejine mites (Acari, Parasitiformes, Mesostigmata, Sejoidea) based on new species from Lord Howe Island and Brazil, and a redescription of Sejus americanus (Banks, 1902)</mods:title>
</mods:titleInfo>
<mods:name id="0E0A72BF983506708E6358A1ABB39F49" type="personal">
<mods:role id="8A1204F314F73F132F99DBA84C2816FB">
<mods:roleTerm id="79C54DF5E134DFE60E09881A7DB03991">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB8CDA6BA55FEA4CEAEAA2AC502B2FC6">Walter, David Evans</mods:namePart>
</mods:name>
<mods:typeOfResource id="35C676C1650656A185CB168F9CD5D41A">text</mods:typeOfResource>
<mods:relatedItem id="85D380DEAB45452772BBF9F5EFCB07A3" type="host">
<mods:titleInfo id="5D7DF37ECEEAC6473269678009F1D5D3">
<mods:title id="CB3934BB8D62DB08186F7D48C2E8F446">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E33C91AD36E9C8DB1948F66A823A09CB">
<mods:date id="E062F917C99B5E248EA7C0539DEF8E30">2013</mods:date>
<mods:detail id="DD4FF78CA4544F1E5E282139AC290296" type="volume">
<mods:number id="17652F8AED9364F7B39E81783C7AB847">3691</mods:number>
</mods:detail>
<mods:detail id="8663886E65B1D356AD69ADBB7D145A55" type="issue">
<mods:number id="54C417471F37AA97EE9D742C0716F48A">3</mods:number>
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<mods:start id="EB2F8C821A3C52F30D7F8BC4294E96F4">301</mods:start>
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</mods:extent>
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<mods:classification id="3B26A10FC78CF3EFCF4BEFE06CDAA773">journal article</mods:classification>
<mods:identifier id="823C3FA4622B1A4FA2C058ABCAA79819" type="DOI">10.11646/zootaxa.3691.3.1</mods:identifier>
<mods:identifier id="B367E750D3B0646A88E103418F0F8414" type="GBIF-Dataset">ad034b98-4029-470c-9a6e-4b5d168b46e4</mods:identifier>
<mods:identifier id="56E32AE36FDBA638D0165F5C247B2074" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0F34885BE927922090C0263605799445" type="Zenodo-Dep">248009</mods:identifier>
<mods:identifier id="BF582E19DFB5887B756C0AE949342F31" type="ZooBank">88298B5D-2941-4A52-8C47-0330168EA240</mods:identifier>
</mods:mods>
<treatment id="094E480B5C20FFCBFF3E49786A53D09B" ID-DOI="http://doi.org/10.5281/zenodo.6159960" ID-GBIF-Taxon="119572736" ID-Zenodo-Dep="6159960" LSID="urn:lsid:plazi:treatment:094E480B5C20FFCBFF3E49786A53D09B" httpUri="http://treatment.plazi.org/id/094E480B5C20FFCBFF3E49786A53D09B" lastPageId="19" lastPageNumber="320" pageId="14" pageNumber="315">
<subSubSection id="C9FDAA965C20FFD6FF3E49786C7FD5D9" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="8158F91D5C20FFD6FF3E49786B3ED606" blockId="14.[151,543,1311,1370]" box="[151,543,1311,1337]" pageId="14" pageNumber="315">
<heading id="DA104E715C20FFD6FF3E49786B3ED606" bold="true" box="[151,543,1311,1337]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<taxonomicName id="46E7829E5C20FFD6FF3E49786B3ED606" authority="Banks, 1902" authorityName="Banks" authorityYear="1902" box="[151,543,1311,1337]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C20FFD6FF3E49786B3ED606" bold="true" box="[151,543,1311,1337]" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF3E4978684CD606" bold="true" box="[151,365,1311,1337]" italics="true" pageId="14" pageNumber="315">Sejus americanus</emphasis>
(Banks, 1902)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8158F91D5C20FFD6FF3E49256819D665" blockId="14.[151,543,1311,1370]" box="[151,312,1346,1370]" pageId="14" pageNumber="315">
<figureCitation id="19DCE5985C20FFD6FF3E49256819D665" box="[151,312,1346,1370]" captionStart-0="FIGURES 36 – 37" captionStart-1="FIGURES 38 – 40" captionStart-2="FIGURES 41 – 43" captionStart-3="FIGURES 44 – 47" captionStartId-0="15.[151,264,1967,1990]" captionStartId-1="16.[151,264,1630,1653]" captionStartId-2="17.[151,275,1891,1916]" captionStartId-3="18.[151,264,1967,1990]" captionTargetBox-0="[185,1397,761,1938]" captionTargetBox-1="[158,1430,199,1600]" captionTargetBox-2="[168,1421,213,1866]" captionTargetBox-3="[176,1411,206,1934]" captionTargetId-0="figure@15.[176,1412,753,1948]" captionTargetId-1="figure@16.[151,1436,193,1609]" captionTargetId-2="figure@17.[151,1436,193,1870]" captionTargetId-3="figure@18.[176,1412,193,1935]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURES 36 – 37. Sejus americanus (Banks, 1902) adult female (light micrographs). 36 Dorsum of female. 37 Venter of female. Scale bar = 500 µm." captionText-1="FIGURES 38 – 40. Sejus americanus (Banks, 1902) adult female. 38 Venter subcapitulum and sterno-genital region (arrow points to anchor-shaped tritosternum). 39 Legs III, IV, metapodal plate (arrow). 40 Ventri-anal shield with postanal seta (po). Scale bars = 50 µm." captionText-2="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." captionText-3="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri-0="https://zenodo.org/record/248018/files/figure.png" httpUri-1="https://zenodo.org/record/248019/files/figure.png" httpUri-2="https://zenodo.org/record/248020/files/figure.png" httpUri-3="https://zenodo.org/record/248021/files/figure.png" pageId="14" pageNumber="315">Figures 36–47</figureCitation>
</paragraph>
<paragraph id="8158F91D5C20FFD6FF3E49ED6C7FD5D9" blockId="14.[151,1437,1418,2018]" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF3E49ED68B1D69C" bold="true" box="[151,400,1418,1443]" pageId="14" pageNumber="315">Specimens examined</emphasis>
.
<collectingCountry id="F9F0B98D5C20FFD6FE3649ED68DDD69C" box="[415,508,1418,1443]" name="Canada" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FE3649ED68DDD69C" bold="true" box="[415,508,1418,1443]" pageId="14" pageNumber="315">Canada</emphasis>
</collectingCountry>
,
<emphasis id="B393250F5C20FFD6FDA249ED6B46D69C" bold="true" box="[523,615,1418,1443]" pageId="14" pageNumber="315">Alberta</emphasis>
: One female, ex meadow soil and litter in aspen parkland (
<geoCoordinate id="E4D39FDA5C20FFD6FAAE49ED6CB6D69D" box="[1287,1431,1418,1442]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="15" value="53.6575">53°39' 27”N</geoCoordinate>
,
<geoCoordinate id="E4D39FDA5C20FFD6FF3E49C9681BD6F9" box="[151,314,1454,1478]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="15" value="-112.766945">112°46' 01”W</geoCoordinate>
),
<quantity id="461F54F85C20FFD6FEF949C968B6D6F9" box="[336,407,1454,1478]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="315" unit="km" value="11.0">11 km</quantity>
S Lamont, Lamont Co., Alberta
<collectingCountry id="F9F0B98D5C20FFD6FCA649C96A47D6F9" box="[783,870,1454,1478]" name="Canada" pageId="14" pageNumber="315">Canada</collectingCountry>
,
<date id="F559DFDD5C20FFD6FCDA49C96D2CD6F9" box="[883,1037,1454,1479]" pageId="14" pageNumber="315" value="2007-05-27">27 May 2007</date>
;
<specimenCount id="97E132945C20FFD6FBB349C96D50D6F9" box="[1050,1137,1454,1478]" pageId="14" pageNumber="315" type="male">2 males</specimenCount>
same data but litter under shrubs,
<date id="F559DFDD5C20FFD6FF4649B56841D6D5" box="[239,352,1490,1514]" pageId="14" pageNumber="315" value="2007-05-13">13.v.2007</date>
;
<specimenCount id="97E132945C20FFD6FEC549B568F8D6D5" box="[364,473,1490,1514]" pageId="14" pageNumber="315" type="female">3 females</specimenCount>
ex polypore sporocarps on aspen logs (
<geoCoordinate id="E4D39FDA5C20FFD6FC3349B56D08D6D5" box="[922,1065,1490,1514]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="15" value="53.65667">53°39' 24”N</geoCoordinate>
,
<geoCoordinate id="E4D39FDA5C20FFD6FB9F49B56DF8D6D5" box="[1078,1241,1490,1514]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="15" value="-112.759445">112°45' 34”W</geoCoordinate>
),
<date id="F559DFDD5C20FFD6FB4749B56CA3D6D5" box="[1262,1410,1490,1514]" pageId="14" pageNumber="315" value="2011-07-16">16 July 2011</date>
;
<specimenCount id="97E132945C20FFD6FA2749B569C7D531" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
,
<specimenCount id="97E132945C20FFD6FF584991681ED531" box="[241,319,1526,1550]" pageId="14" pageNumber="315" type="male">2 male</specimenCount>
, 1 deutonymph ex litter, moss, rotten wood under brush pile (
<geoCoordinate id="E4D39FDA5C20FFD6FC5A49916DA3D531" box="[1011,1154,1526,1550]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="15" value="53.65667">53°39' 24”N</geoCoordinate>
,
<geoCoordinate id="E4D39FDA5C20FFD6FB2749916C10D531" box="[1166,1329,1526,1550]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="15" value="-112.759445">112°45' 34”W</geoCoordinate>
),
<date id="F559DFDD5C20FFD6FAEC499169F3D50D" pageId="14" pageNumber="315" value="2012-06-15">15 June 2012</date>
, D.E. Walter &amp; H.C. Proctor;
<specimenCount id="97E132945C20FFD6FD904A7D6B8DD50D" box="[569,684,1562,1586]" pageId="14" pageNumber="315" type="female">2 females</specimenCount>
,
<specimenCount id="97E132945C20FFD6FD124A7D6A2BD50D" box="[699,778,1562,1586]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
Ecosystem Management Emulating Natural Disturbance Project (EMEND), NW Peace River (
<geoCoordinate id="E4D39FDA5C20FFD6FD934A596BF2D569" box="[570,723,1598,1622]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="15" value="56.77028">56° 46' 13'' N</geoCoordinate>
,–
<geoCoordinate id="E4D39FDA5C20FFD6FD4E4A596AB2D569" box="[743,915,1598,1622]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="15" value="-118.37444">118° 22' 28'' W</geoCoordinate>
), 25–940, 83–892, 63–889, I. Diaz.
<emphasis id="B393250F5C20FFD6FA8D4A596CA6D568" bold="true" box="[1316,1415,1598,1623]" pageId="14" pageNumber="315">Quebec:</emphasis>
<specimenCount id="97E132945C20FFD6FA274A5969D0D545" pageId="14" pageNumber="315" type="female">2 females</specimenCount>
,
<specimenCount id="97E132945C20FFD6FF544A056868D545" box="[253,329,1634,1658]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
ex white spruce, Harrington, C.I.P. Tree Farm,
<date id="F559DFDD5C20FFD6FCCF4A056AD0D545" box="[870,1009,1634,1658]" pageId="14" pageNumber="315" value="1965-06-09">9 June 1965</date>
, R.M. Emberson.
<emphasis id="B393250F5C20FFD6FB694A046C5FD544" bold="true" box="[1216,1406,1634,1659]" pageId="14" pageNumber="315">New Brunswick</emphasis>
:
<specimenCount id="97E132945C20FFD6FA274A0569C7D5A1" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
,
<specimenCount id="97E132945C20FFD6FF584AE1681DD5A1" box="[241,316,1670,1694]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
ex deciduous litter in depression, Kouchibouguac National Park, Kent Co.,
<date id="F559DFDD5C20FFD6FB384AE16CB9D5A1" box="[1169,1432,1670,1694]" pageId="14" pageNumber="315" value="1977-09-27" valueMax="1977-09-29" valueMin="1977-09-27">27–29 September 1977</date>
, I.M. Smith; one female same data except beneath bark of fallen beech branch,
<date id="F559DFDD5C20FFD6FBBD4ACD6D88D5FD" box="[1044,1193,1706,1730]" pageId="14" pageNumber="315" value="1978-07-26">26 July 1978</date>
.
<emphasis id="B393250F5C20FFD6FB1C4ACD6CA7D5FC" bold="true" box="[1205,1414,1706,1731]" pageId="14" pageNumber="315">
<collectingCountry id="F9F0B98D5C20FFD6FB1C4ACD6DD0D5FC" box="[1205,1265,1706,1731]" name="United States of America" pageId="14" pageNumber="315">USA</collectingCountry>
: Minnesota:
</emphasis>
<specimenCount id="97E132945C20FFD6FA274ACD69D0D5D9" pageId="14" pageNumber="315" type="female">2 females</specimenCount>
, in rotten wood, Anaka Co.,
<date id="F559DFDD5C20FFD6FD9A4AA96BF3D5D9" box="[563,722,1742,1766]" pageId="14" pageNumber="315" value="1941-04-19">19 April 1941</date>
, G. Kretzschmer. All in the Canadian National Collection.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C20FFCBFF6E4A956C3CD223" lastPageId="19" lastPageNumber="320" pageId="14" pageNumber="315" type="diagnosis">
<paragraph id="8158F91D5C20FFD7FF6E4A956D25D1F3" blockId="14.[151,1437,1418,2018]" lastBlockId="15.[151,1436,151,716]" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF6E4A95681BD434" bold="true" box="[199,314,1778,1803]" pageId="14" pageNumber="315">Diagnosis</emphasis>
. Adults large (adult idiosoma 900–1100 long along midline). Cuticle sticky, with adherent soil particles; soft cuticle plicate and with numerous small (5–6) denticles ventrally, laterally; dorsal shields ornamented with large, rounded tubercles (
<figureCitation id="19DCE5985C20FFD6FD264B5D6BCAD46D" box="[655,747,1850,1874]" captionStart="FIGURES 36 – 37" captionStartId="15.[151,264,1967,1990]" captionTargetBox="[185,1397,761,1938]" captionTargetId="figure@15.[176,1412,753,1948]" captionTargetPageId="15" captionText="FIGURES 36 – 37. Sejus americanus (Banks, 1902) adult female (light micrographs). 36 Dorsum of female. 37 Venter of female. Scale bar = 500 µm." httpUri="https://zenodo.org/record/248018/files/figure.png" pageId="14" pageNumber="315">Figs 36</figureCitation>
,
<figureCitation id="19DCE5985C20FFD6FD554B5D6A3DD46D" box="[764,796,1850,1874]" captionStart="FIGURES 44 – 47" captionStartId="18.[151,264,1967,1990]" captionTargetBox="[176,1411,206,1934]" captionTargetId="figure@18.[176,1412,193,1935]" captionTargetPageId="18" captionText="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="14" pageNumber="315">44</figureCitation>
); peritrematal shield broad, joined to podonotal shield anteriorly; peritreme on ventral margin of shield extending from between coxae III–IV to near juncture podonotal shield. Posterior horn-complexes each formed of inner and outer horns with distal seta and separated from other shields by soft cuticle; outer horn conical with long (~100) stalk bearing stout (125 long), densely barbed seta with spatulate, membranous tip (flame seta); ~4 small (20–25) subdistal (midpoint to near base of flame seta) barbed setae; 4 longer (38–56) densely barbed setae on outer margin, dorsal and ventral poroid; inner projection short (~35) with stout (105 long) densely barbed seta with spatulate, membranous tip. Postanal seta on strong apophysis (
<figureCitation id="19DCE5985C21FFD7FF364CB869D3D3C8" box="[159,242,223,247]" captionStart="FIGURES 38 – 40" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[158,1430,199,1600]" captionTargetId="figure@16.[151,1436,193,1609]" captionTargetPageId="16" captionText="FIGURES 38 – 40. Sejus americanus (Banks, 1902) adult female. 38 Venter subcapitulum and sterno-genital region (arrow points to anchor-shaped tritosternum). 39 Legs III, IV, metapodal plate (arrow). 40 Ventri-anal shield with postanal seta (po). Scale bars = 50 µm." httpUri="https://zenodo.org/record/248019/files/figure.png" pageId="15" pageNumber="316">Fig. 40</figureCitation>
<emphasis id="B393250F5C21FFD7FF554C866839D3C7" box="[252,280,225,248]" italics="true" pageId="15" pageNumber="316">po</emphasis>
). Gnathotectum with sparsely denticulate margin and strong median and lateral processes (
<figureCitation id="19DCE5985C21FFD7FA914CB86CAED3C8" box="[1336,1423,223,247]" captionStart="FIGURES 44 – 47" captionStartId="18.[151,264,1967,1990]" captionTargetBox="[176,1411,206,1934]" captionTargetId="figure@18.[176,1412,193,1935]" captionTargetPageId="18" captionText="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="15" pageNumber="316">Fig. 46</figureCitation>
); palpal apotele with two tines and strong basal spur (
<figureCitation id="19DCE5985C21FFD7FD4B4D636A17D223" box="[738,822,260,284]" captionStart="FIGURES 41 – 43" captionStartId="17.[151,275,1891,1916]" captionTargetBox="[168,1421,213,1866]" captionTargetId="figure@17.[151,1436,193,1870]" captionTargetPageId="17" captionText="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="15" pageNumber="316">Fig. 43</figureCitation>
), palpal setation normal, lateral setae of palptibia with subdistal spur; chelicerae serrate (
<figureCitation id="19DCE5985C21FFD7FD894D406B56D200" box="[544,631,295,319]" captionStart="FIGURES 41 – 43" captionStartId="17.[151,275,1891,1916]" captionTargetBox="[168,1421,213,1866]" captionTargetId="figure@17.[151,1436,193,1870]" captionTargetPageId="17" captionText="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="15" pageNumber="316">Fig. 42</figureCitation>
), fixed digit with 9–10 teeth (including Gabelzahn), short (7), simple pilus dentilis; movable digit with double row of numerous small teeth; deutosternal gutter with numerous rows of 4–6 denticles, patches of ~12 denticles on subcapitulum between sparsely barbed palpcoxal setae (~60 long);
<emphasis id="B393250F5C21FFD7FA294D176CBAD2B7" box="[1408,1435,368,392]" italics="true" pageId="15" pageNumber="316">h1</emphasis>
and internal malae membranous (
<figureCitation id="19DCE5985C21FFD7FDB24DF36B53D293" box="[539,626,404,428]" captionStart="FIGURES 41 – 43" captionStartId="17.[151,275,1891,1916]" captionTargetBox="[168,1421,213,1866]" captionTargetId="figure@17.[151,1436,193,1870]" captionTargetPageId="17" captionText="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="15" pageNumber="316">Fig. 41</figureCitation>
); anterior (
<emphasis id="B393250F5C21FFD7FD5F4DF36A33D293" box="[758,786,404,428]" italics="true" pageId="15" pageNumber="316">h3</emphasis>
~60) and posterior (
<emphasis id="B393250F5C21FFD7FBAE4DF36D02D293" box="[1031,1059,404,428]" italics="true" pageId="15" pageNumber="316">h2</emphasis>
~110) hypostomal setae simple; corniculi (30 long) horn-like; tritosternal base (50 long) columnar, laciniae divided distally, base broadened (60) with a pair of lateral conical teeth (
<figureCitation id="19DCE5985C21FFD7FD8D4DBB6B56D2CB" box="[548,631,476,500]" captionStart="FIGURES 38 – 40" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[158,1430,199,1600]" captionTargetId="figure@16.[151,1436,193,1609]" captionTargetPageId="16" captionText="FIGURES 38 – 40. Sejus americanus (Banks, 1902) adult female. 38 Venter subcapitulum and sterno-genital region (arrow points to anchor-shaped tritosternum). 39 Legs III, IV, metapodal plate (arrow). 40 Ventri-anal shield with postanal seta (po). Scale bars = 50 µm." httpUri="https://zenodo.org/record/248019/files/figure.png" pageId="15" pageNumber="316">Fig. 38</figureCitation>
, arrowed). All legs with paired claws, segments ornamented with short rows of denticles; most setae set on short tubercles, strongly barbed, ending in flame-shaped, membranous tips; basitarsi II–IV with ventral gland opening adjacent to peripodal fissure, ventral setae smooth; telotarsi II–III each with ~6 trident setae (
<figureCitation id="19DCE5985C21FFD7FE3A4E2068C6D160" box="[403,487,583,607]" captionStart="FIGURES 44 – 47" captionStartId="18.[151,264,1967,1990]" captionTargetBox="[176,1411,206,1934]" captionTargetId="figure@18.[176,1412,193,1935]" captionTargetPageId="18" captionText="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="15" pageNumber="316">Fig. 47</figureCitation>
) and nude intercalary sclerite; setae of intercalary sclerite of tarsus IV simple,
<emphasis id="B393250F5C21FFD7FADD4E2F6CBDD15F" box="[1396,1436,584,608]" italics="true" pageId="15" pageNumber="316">av4</emphasis>
(65) about 50% longer than
<emphasis id="B393250F5C21FFD7FE664E0B68D6D1BB" box="[463,503,620,644]" italics="true" pageId="15" pageNumber="316">pv4</emphasis>
(40); tarsus I with apicotarsus. Setation of coxae I–IV: 2-2-2-1; trochanters I–IV: 6- 5-5-5; femora I–IV:
<date id="F559DFDD5C21FFD7FE2D4EE868D7D198" box="[388,502,655,679]" pageId="15" pageNumber="316" value="1913-11-06" valueMax="1913-11-06" valueMin="1913-11-06">13-11-6-6</date>
; genua I–IV:
<date id="F559DFDD5C21FFD7FD3E4EE86A03D198" box="[663,802,655,679]" pageId="15" pageNumber="316" value="2011-12-12" valueMax="2011-12-12" valueMin="2011-12-12">12-12-12-11</date>
(2-3/1, 3/1-2; 2-3/1, 3/1-2; 2-3/1, 3/1-2; 2-3/1, 3/1-1); tibiae I–IV:
<date id="F559DFDD5C21FFD7FEB44ED36887D1F3" box="[285,422,692,716]" pageId="15" pageNumber="316" value="1913-11-11" valueMax="1913-11-11" valueMin="1913-11-11">13-11-11-11</date>
(2-3/2, 3/1-2; 2-2/1, 3/1-2; 2-2/1, 3/1-2; 2-2/1, 3/1-2).
</paragraph>
<caption id="D598A9955C21FFD7FF3E4BC86897D4DB" httpUri="https://zenodo.org/record/248018/files/figure.png" pageId="15" pageNumber="316" targetBox="[185,1397,761,1938]" targetPageId="15">
<paragraph id="8158F91D5C21FFD7FF3E4BC86897D4DB" blockId="15.[151,1436,1967,2020]" pageId="15" pageNumber="316">
<emphasis id="B393250F5C21FFD7FF3E4BC8680AD4FA" bold="true" box="[151,299,1967,1990]" pageId="15" pageNumber="316">FIGURES 36</emphasis>
–
<emphasis id="B393250F5C21FFD7FE9E4BD76876D4FA" bold="true" box="[311,343,1968,1989]" pageId="15" pageNumber="316">37.</emphasis>
<taxonomicName id="46E7829E5C21FFD7FEC84BD76B95D4F9" authority="Banks, 1902" authorityName="Banks" authorityYear="1902" box="[353,692,1967,1990]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C21FFD7FEC84BD76B36D4F9" box="[353,535,1968,1990]" italics="true" pageId="15" pageNumber="316">Sejus americanus</emphasis>
(Banks, 1902)
</taxonomicName>
adult female (light micrographs). 36 Dorsum of female. 37 Venter of female. Scale bar = 500 µm.
</paragraph>
</caption>
<caption id="D598A9955C3EFFC8FF3E4A396843D58E" httpUri="https://zenodo.org/record/248019/files/figure.png" pageId="16" pageNumber="317" targetBox="[158,1430,199,1600]" targetPageId="16">
<paragraph id="8158F91D5C3EFFC8FF3E4A396843D58E" blockId="16.[151,1436,1630,1713]" pageId="16" pageNumber="317">
<emphasis id="B393250F5C3EFFC8FF3E4A39680BD54C" bold="true" box="[151,298,1630,1653]" pageId="16" pageNumber="317">FIGURES 38</emphasis>
–
<emphasis id="B393250F5C3EFFC8FE9E4A396876D54C" bold="true" box="[311,343,1630,1651]" pageId="16" pageNumber="317">40.</emphasis>
<taxonomicName id="46E7829E5C3EFFC8FEC94A386B90D54A" authority="Banks, 1902" authorityName="Banks" authorityYear="1902" box="[352,689,1630,1653]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3EFFC8FEC94A386B35D54B" box="[352,532,1631,1652]" italics="true" pageId="16" pageNumber="317">Sejus americanus</emphasis>
(Banks, 1902)
</taxonomicName>
adult female. 38 Venter subcapitulum and sterno-genital region (arrow points to anchor-shaped tritosternum). 39 Legs III, IV, metapodal plate (arrow). 40 Ventri-anal shield with postanal seta (
<emphasis id="B393250F5C3EFFC8FADD4A196CACD5AC" box="[1396,1421,1662,1683]" italics="true" pageId="16" pageNumber="317">po</emphasis>
). Scale bars = 50 µm.
</paragraph>
</caption>
<paragraph id="8158F91D5C3EFFCBFF6E4ABE6C3CD223" blockId="16.[151,1436,1753,2029]" lastBlockId="19.[151,1436,151,932]" lastPageId="19" lastPageNumber="320" pageId="16" pageNumber="317">
<emphasis id="B393250F5C3EFFC8FF6E4ABE683FD5CE" bold="true" box="[199,286,1753,1777]" pageId="16" pageNumber="317">Female</emphasis>
. Sternal setae (
<figureCitation id="19DCE5985C3EFFC8FE654ABE6B3ED5CE" box="[460,543,1753,1777]" captionStart="FIGURES 38 – 40" captionStartId="16.[151,264,1630,1653]" captionTargetBox="[158,1430,199,1600]" captionTargetId="figure@16.[151,1436,193,1609]" captionTargetPageId="16" captionText="FIGURES 38 – 40. Sejus americanus (Banks, 1902) adult female. 38 Venter subcapitulum and sterno-genital region (arrow points to anchor-shaped tritosternum). 39 Legs III, IV, metapodal plate (arrow). 40 Ventri-anal shield with postanal seta (po). Scale bars = 50 µm." httpUri="https://zenodo.org/record/248019/files/figure.png" pageId="16" pageNumber="317">Fig. 38</figureCitation>
) simple,
<emphasis id="B393250F5C3EFFC8FD204ABE6B8BD5CE" box="[649,682,1753,1777]" italics="true" pageId="16" pageNumber="317">st1</emphasis>
and
<emphasis id="B393250F5C3EFFC8FD4B4ABE6A23D5CE" box="[738,770,1753,1777]" italics="true" pageId="16" pageNumber="317">st2</emphasis>
each on small platelet with lyrifissure on posterior margin (
<emphasis id="B393250F5C3EFFC8FF094A9A69EED42A" box="[160,207,1789,1813]" italics="true" pageId="16" pageNumber="317">stp1</emphasis>
–
<emphasis id="B393250F5C3EFFC8FF744A9A69CAD42A" box="[221,235,1789,1813]" italics="true" pageId="16" pageNumber="317">2</emphasis>
, respectively);
<emphasis id="B393250F5C3EFFC8FE0A4A9A68E5D42A" box="[419,452,1789,1813]" italics="true" pageId="16" pageNumber="317">st2</emphasis>
and
<emphasis id="B393250F5C3EFFC8FDAD4A9A6B04D42A" box="[516,549,1789,1813]" italics="true" pageId="16" pageNumber="317">st3</emphasis>
–
<emphasis id="B393250F5C3EFFC8FD9A4A9A6B60D42A" box="[563,577,1789,1813]" italics="true" pageId="16" pageNumber="317">4</emphasis>
inserted on small, indistinct platelets. Endopodal elements III–IV fused, narrow, bearing
<emphasis id="B393250F5C3EFFC8FEF84B46685ED406" box="[337,383,1825,1849]" italics="true" pageId="16" pageNumber="317">stp3</emphasis>
anteriorly. Genital shield microtuberculate, rounded (180 long, 200 wide) with four pairs of simple marginal setae, one pair of lyrifissures in posterior corners, anterior notch shallow; internal median flaskshaped sclerotisation (
<figureCitation id="19DCE5985C3EFFC8FE3D4B0E68C6D4BE" box="[404,487,1897,1921]" captionStart="FIGURES 36 – 37" captionStartId="15.[151,264,1967,1990]" captionTargetBox="[185,1397,761,1938]" captionTargetId="figure@15.[176,1412,753,1948]" captionTargetPageId="15" captionText="FIGURES 36 – 37. Sejus americanus (Banks, 1902) adult female (light micrographs). 36 Dorsum of female. 37 Venter of female. Scale bar = 500 µm." httpUri="https://zenodo.org/record/248018/files/figure.png" pageId="16" pageNumber="317">Fig. 37</figureCitation>
) supporting spermatheca. Ventral setae on small, raised platelets or larger platelets with 1–2 dermal gland openings, simple anteriorly, becoming longer and densely barbed posteriorly; metapodal plates (115 long, 95 with) rounded-subtriangular, tuberculate, and with median dermal gland opening; ventri-anal shield projecting anal opening ventrally, postanal seta (55) on short tubercle (~10), densely barbed and with modified tip. Podonotal shield (450 along midline) truncate posteriorly, joined to peritrematal shields anteriorly with large, raised median region (
<figureCitation id="19DCE5985C3DFFCBFD894CDB6B56D3EB" box="[544,631,188,212]" captionStart="FIGURES 36 – 37" captionStartId="15.[151,264,1967,1990]" captionTargetBox="[185,1397,761,1938]" captionTargetId="figure@15.[176,1412,753,1948]" captionTargetPageId="15" captionText="FIGURES 36 – 37. Sejus americanus (Banks, 1902) adult female (light micrographs). 36 Dorsum of female. 37 Venter of female. Scale bar = 500 µm." httpUri="https://zenodo.org/record/248018/files/figure.png" pageId="19" pageNumber="320">Fig. 36</figureCitation>
); two pairs of mesonotal scutellae, anterior longer and narrower than posterior; pygidial shield oval (340 across) with ~ten pairs of setae, tuberculate anteriorly and medially, extended anteriorly to between posterior mesonotal scutellae by irregular field of dark, lightly sclerotised cuticle.
</paragraph>
</subSubSection>
<caption id="D598A9955C3FFFC9FF3E4B0469CFD481" httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="17" pageNumber="318" targetBox="[168,1421,213,1866]" targetPageId="17">
<paragraph id="8158F91D5C3FFFC9FF3E4B0469CFD481" blockId="17.[151,1436,1891,1982]" pageId="17" pageNumber="318">
<emphasis id="B393250F5C3FFFC9FF3E4B046817D443" bold="true" box="[151,310,1891,1916]" pageId="17" pageNumber="318">FIGURES 41</emphasis>
–
<emphasis id="B393250F5C3FFFC9FEED4B046846D443" bold="true" box="[324,359,1891,1916]" pageId="17" pageNumber="318">43.</emphasis>
<taxonomicName id="46E7829E5C3FFFC9FEC74B026B95D444" authority="Banks, 1902" authorityName="Banks" authorityYear="1902" box="[366,692,1893,1915]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="americanus">Sejus americanus (Banks, 1902)</taxonomicName>
adult female. 41 Hypostome showing membranous seta h1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~25 µm long). Scale bars 41 = 10 µm, 42 =50 µm.
</paragraph>
</caption>
<caption id="D598A9955C3CFFCAFF3E4BC86A9FD4D6" httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="18" pageNumber="319" targetBox="[176,1411,206,1934]" targetPageId="18">
<paragraph id="8158F91D5C3CFFCAFF3E4BC86A9FD4D6" blockId="18.[151,1436,1967,2025]" pageId="18" pageNumber="319">
<emphasis id="B393250F5C3CFFCAFF3E4BC86809D4FA" bold="true" box="[151,296,1967,1990]" pageId="18" pageNumber="319">FIGURES 44</emphasis>
–
<emphasis id="B393250F5C3CFFCAFE9C4BD76874D4FA" bold="true" box="[309,341,1968,1989]" pageId="18" pageNumber="319">47.</emphasis>
<taxonomicName id="46E7829E5C3CFFCAFEF54BD66B86D4F9" authority="(Banks, 1902)" authorityName="Banks" authorityYear="1902" baseAuthorityName="Banks" baseAuthorityYear="1902" box="[348,679,1968,1990]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3CFFCAFEF54BD66B2ED4F9" box="[348,527,1969,1990]" italics="true" pageId="18" pageNumber="319">Sejus americanus</emphasis>
(Banks, 1902)
</taxonomicName>
adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44–45 = 500 µm; 46–47 = 50 µm.
</paragraph>
</caption>
<subSubSection id="C9FDAA965C3DFFCBFF6E4D4F6C08D293" pageId="19" pageNumber="320" type="description">
<paragraph id="8158F91D5C3DFFCBFF6E4D4F6C08D293" blockId="19.[151,1436,151,932]" pageId="19" pageNumber="320">
<emphasis id="B393250F5C3DFFCBFF6E4D4F6822D27F" bold="true" box="[199,259,296,320]" pageId="19" pageNumber="320">Male</emphasis>
. Similar to female except smaller and more extensively sclerotised, opisthonotal shield (
<figureCitation id="19DCE5985C3DFFCBFB724D406C0DD200" box="[1243,1324,295,319]" captionStart="FIGURES 44 – 47" captionStartId="18.[151,264,1967,1990]" captionTargetBox="[176,1411,206,1934]" captionTargetId="figure@18.[176,1412,193,1935]" captionTargetPageId="18" captionText="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="19" pageNumber="320">Fig. 44</figureCitation>
) covering region, without separate scutellae or pygidial shield; ventri-anal shield with irregular anterior margin, extended laterally to capture metapodal plates (
<figureCitation id="19DCE5985C3DFFCBFDE94D086BB4D2B8" box="[576,661,367,391]" captionStart="FIGURES 44 – 47" captionStartId="18.[151,264,1967,1990]" captionTargetBox="[176,1411,206,1934]" captionTargetId="figure@18.[176,1412,193,1935]" captionTargetPageId="18" captionText="FIGURES 44 – 47. Sejus americanus (Banks, 1902) adult male. 44 Dorsum of male. 45 Venter of male. 46 Gnathotectum. 47 Leg II with trident setae on tarsus. Scale bars 44 – 45 = 500 µm; 46 – 47 = 50 µm." httpUri="https://zenodo.org/record/248021/files/figure.png" pageId="19" pageNumber="320">Fig. 45</figureCitation>
); sternitigenital shield (260) bearing four pairs of simple setae, three pairs of lyrifissures; genital opening subcircular (55 long), covered by two nude valves, between coxae II.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C3DFFCBFF6E4DDF6A53D09B" pageId="19" pageNumber="320" type="discussion">
<paragraph id="8158F91D5C3DFFCBFF6E4DDF6B5CD1BB" blockId="19.[151,1436,151,932]" pageId="19" pageNumber="320">
<emphasis id="B393250F5C3DFFCBFF6E4DDF6815D2EF" bold="true" box="[199,308,440,464]" pageId="19" pageNumber="320">Remarks</emphasis>
. Banks’ (1902) description and illustration are sketchy, but the large size (Banks says 1.5 mm, but presumably this includes the horns and gnathosoma), the distinctive median ridge on the podonotal shield, and the northern distribution (Olympia, Washington, St. Croix Falls, Wisconsin) all suggest that the Alberta mites are
<taxonomicName id="46E7829E5C3DFFCBFA2E4E67683BD103" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3DFFCBFA2E4E67683BD103" italics="true" pageId="19" pageNumber="320">S. americanus</emphasis>
</taxonomicName>
. The
<typeStatus id="5E5C47BF5C3DFFCBFEF24E4268AAD102" box="[347,395,549,573]" pageId="19" pageNumber="320">type</typeStatus>
specimen appears to be lost (on loan from the Museum of Comparative Zoology at Harvard to Ohio State University), but specimens identified as
<taxonomicName id="46E7829E5C3DFFCBFD5A4E2F6AB3D15F" box="[755,914,584,608]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3DFFCBFD5A4E2F6AB3D15F" box="[755,914,584,608]" italics="true" pageId="19" pageNumber="320">S. americanus</emphasis>
</taxonomicName>
from the Canadian National Collection appear to be the same species as the Alberta mites.
</paragraph>
<paragraph id="8158F91D5C3DFFCBFF6E4EE86A53D09B" blockId="19.[151,1436,151,932]" pageId="19" pageNumber="320">
The adults of
<taxonomicName id="46E7829E5C3DFFCBFECD4EF76B0AD197" box="[356,555,656,680]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3DFFCBFECD4EF76B0AD197" box="[356,555,656,680]" italics="true" pageId="19" pageNumber="320">Sejus americanus</emphasis>
</taxonomicName>
are easily distinguished from the only other described species of the genus in North
<collectingCountry id="F9F0B98D5C3DFFCBFF4A4ED36867D1F3" box="[227,326,692,716]" name="United States of America" pageId="19" pageNumber="320">America</collectingCountry>
,
<taxonomicName id="46E7829E5C3DFFCBFEFC4ED26AA3D1F3" authority="Lekveishvili &amp; Klompen (2004)" authorityName="Lekveishvili &amp; Klompen" authorityYear="2004" box="[341,898,692,716]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis id="B393250F5C3DFFCBFEFC4ED268DCD1F3" box="[341,509,692,716]" italics="true" pageId="19" pageNumber="320">S. carolinensis</emphasis>
Lekveishvili &amp; Klompen (2004)
</taxonomicName>
. For example,
<taxonomicName id="46E7829E5C3DFFCBFB9A4ED26DF7D1F3" box="[1075,1238,693,716]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3DFFCBFB9A4ED26DF7D1F3" box="[1075,1238,693,716]" italics="true" pageId="19" pageNumber="320">S. americanus</emphasis>
</taxonomicName>
has a 3-pronged tectum (vs triangular in
<taxonomicName id="46E7829E5C3DFFCBFE0F4EBF6B0FD1CF" box="[422,558,728,752]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis id="B393250F5C3DFFCBFE0F4EBF6B0FD1CF" box="[422,558,728,752]" italics="true" pageId="19" pageNumber="320">carolinensis</emphasis>
</taxonomicName>
), a double row of teeth on the moveable digit (vs one row), an anchor-shaped tritosternum (vs sickle-shaped), a female genital shield with four pairs of setae and a pair of pores in the posterior corners (vs three pairs and pores on midline), a much broader ventri-anal shield, and the posterior horns are separated from the pygidial and ventri-anal shields by soft cuticle. However,
<taxonomicName id="46E7829E5C3DFFCBFC5D4F226DB5D063" box="[1012,1172,837,860]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C3DFFCBFC5D4F226DB5D063" box="[1012,1172,837,860]" italics="true" pageId="19" pageNumber="320">S. americanus</emphasis>
</taxonomicName>
is closely related to the
<typeStatus id="5E5C47BF5C3DFFCBFF3E4F0E69E6D0BE" box="[151,199,873,897]" pageId="19" pageNumber="320">type</typeStatus>
species of the genus,
<taxonomicName id="46E7829E5C3DFFCBFE134F0F6BCDD040" authority="C.L. Koch, 1836" authorityName="C.L. Koch" authorityYear="1836" box="[442,748,871,896]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="togatus">
<emphasis id="B393250F5C3DFFCBFE134F0F6B06D0BF" box="[442,551,872,896]" italics="true" pageId="19" pageNumber="320">S. togatus</emphasis>
C.L. Koch, 1836
</taxonomicName>
, a somewhat variable species (Gwiazdowicz &amp; Gulvik 2009), and further comparisons between the species might be profitable.
</paragraph>
</subSubSection>
</treatment>
</document>
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