275 lines
38 KiB
XML
275 lines
38 KiB
XML
<document id="3FA328174A904ECEAB98EE3BC171717F" ID-DOI="10.11646/zootaxa.5306.5.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="8073323" ID-ZooBank="439834E8-F594-4999-BB36-AE6DAD18C2AE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687504363901" checkinUser="plazi" docAuthor="Fan, Shilv, Gu, Jun-Jie & Cao, Chengquan" docDate="2023" docId="092C5174FFF72931FF61FE78FC01746A" docLanguage="en" docName="zootaxa.5306.5.7.pdf" docOrigin="Zootaxa 5306 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5306.5.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Burmadactylus tenuicerci Fan & Gu & Cao 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="597" masterDocId="F515290CFFF62933FFF6FF87FFD6737D" masterDocTitle="A new species of the genus Burmadactylus Heads, 2009 from mid-Cretaceous amber in north Myanmar (Orthoptera: Caelifera: Tridactyloidea)" masterLastPageNumber="598" masterPageNumber="595" pageNumber="596" updateTime="1687777483202" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2766BEC2D10BE207BB0148E15D415DBF">A new species of the genus Burmadactylus Heads, 2009 from mid-Cretaceous amber in north Myanmar (Orthoptera: Caelifera: Tridactyloidea)</mods:title>
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<mods:name id="B249FC51CDF168CF9606310D2851412F" type="personal">
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<mods:namePart id="0B7F9F4F1CB9C19D7DA8F84AB61D9DDE">Fan, Shilv</mods:namePart>
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<mods:nameIdentifier id="A10D600972AB4AF4C8486CC98313A7B3" type="ORCID">0009-0006-5005-755X</mods:nameIdentifier>
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<mods:affiliation id="ED918A75E92F836513D6922E9AA5BD03">College of Agronomy, Sichuan Agricultural University, Chengdu 611130, China & 13884121235 @ 163. com; https: // orcid. org / 0009 - 0006 - 5005 - 755 X</mods:affiliation>
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<mods:roleTerm id="B2AA46A5F63982E7486C32678B5E63DB">Author</mods:roleTerm>
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<mods:namePart id="68D7CDFBC80C7E32CF51B85E25A6A28F">Gu, Jun-Jie</mods:namePart>
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<mods:nameIdentifier id="8B9222A3575425C5CE6B6D40EB6306FA" type="ORCID">0000-0003-1931-4424</mods:nameIdentifier>
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<mods:affiliation id="38528ED71B14020D5149D04BB013E7F8">College of Agronomy, Sichuan Agricultural University, Chengdu 611130, China & orthoptera _ gu @ aliyun. com; https: // orcid. org / 0000 - 0003 - 1931 - 4424</mods:affiliation>
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<mods:nameIdentifier id="E305E2D32345C5BDDC3DE4C28954B7B0" type="email">orthoptera_gu@aliyun.com</mods:nameIdentifier>
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<mods:name id="B7471490E8F0180DE2512E817AFF6E18" type="personal">
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<mods:namePart id="8A3DD52F6C0BB7B12F76E7E6F216B1D4">Cao, Chengquan</mods:namePart>
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<mods:nameIdentifier id="E919D5EE8CB82B7368D324050C536E7E" type="ORCID">0000-0001-9701-2638</mods:nameIdentifier>
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<mods:affiliation id="1DBA3F84393948ACEEB31302E3CEC018">College of Life Sciences, Leshan Normal University, Leshan 614004, China & chqcao 1314 @ 163. com; https: // orcid. org / 0000 - 0001 - 9701 - 2638</mods:affiliation>
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<mods:typeOfResource id="E11D5C75426DEFF26021A5F1B5DC141E">text</mods:typeOfResource>
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<mods:title id="C5E842FBEE8BD2A4B5961E4B3302AF81">Zootaxa</mods:title>
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<mods:part id="A5C2EC39DAB1AD2B35B059D41D31557A">
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<mods:date id="AC5D47856CCFD6EAACE30D51CF90FD8F">2023</mods:date>
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<mods:number id="3456E284C3FDB31C6F5A3605AEDFB1F4">2023-06-23</mods:number>
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<mods:number id="1251685D9A8C83E211FF9D30601BE57B">5306</mods:number>
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<mods:number id="9E1098AB55FC915532BA8B73F9367FF0">5</mods:number>
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<mods:url id="D9FD8E0B76243CE681311D1260877110">http://dx.doi.org/10.11646/zootaxa.5306.5.7</mods:url>
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<mods:classification id="36DD6400E5BFB38DFF5463D5B74ED831">journal article</mods:classification>
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<mods:identifier id="1ADBC712C9ACBDB0F08494E89BC42E60" type="DOI">10.11646/zootaxa.5306.5.7</mods:identifier>
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<treatment id="092C5174FFF72931FF61FE78FC01746A" ID-DOI="http://doi.org/10.5281/zenodo.8073321" ID-Zenodo-Dep="8073321" LSID="urn:lsid:plazi:treatment:092C5174FFF72931FF61FE78FC01746A" httpUri="http://treatment.plazi.org/id/092C5174FFF72931FF61FE78FC01746A" lastPageId="2" lastPageNumber="597" pageId="1" pageNumber="596">
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<subSubSection id="C99FB3E9FFF72932FF61FE78FDE27164" box="[151,564,511,538]" pageId="1" pageNumber="596" type="nomenclature">
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<paragraph id="813AE062FFF72932FF61FE78FDE27164" blockId="1.[151,956,511,574]" box="[151,564,511,538]" pageId="1" pageNumber="596">
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<heading id="DA72570EFFF72932FF61FE78FDE27164" bold="true" box="[151,564,511,538]" fontSize="11" level="1" pageId="1" pageNumber="596" reason="1">
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<emphasis id="B3F13C70FFF72932FF61FE78FDE27164" bold="true" box="[151,564,511,538]" pageId="1" pageNumber="596">
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<taxonomicName id="46859BE1FFF72932FF61FE78FE067164" authority="Fan & Gu & Cao, 2023" authorityName="Fan & Gu & Cao" authorityYear="2023" box="[151,464,511,538]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="tenuicerci" status="sp. nov.">
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<emphasis id="B3F13C70FFF72932FF61FE78FE067164" bold="true" box="[151,464,511,538]" italics="true" pageId="1" pageNumber="596">Burmadactylus tenuicerci</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A8C2810BFFF72932FE21FE78FDE27164" box="[471,564,511,537]" pageId="1" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C99FB3E9FFF72932FF61FDA4FC6A7143" box="[151,956,547,574]" pageId="1" pageNumber="596" type="description">
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<paragraph id="813AE062FFF72932FF61FDA4FC6A7143" blockId="1.[151,956,511,574]" box="[151,956,547,574]" pageId="1" pageNumber="596">
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<uri id="F514EC60FFF72932FF61FDA4FC6A7143" box="[151,956,547,574]" pageId="1" pageNumber="596">
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urn:lsid:zoobank.org:act:
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<uuid id="F523DAB7FFF72932FE5DFDA4FC6A7143" box="[427,956,547,574]" pageId="1" pageNumber="596">0DF8451E-7121-4B2D-9328-64E458B68B8B</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C99FB3E9FFF72932FF61FDECFD7F71B0" pageId="1" pageNumber="596" type="materials_examined">
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<paragraph id="813AE062FFF72932FF61FDECFD7F71B0" blockId="1.[151,1436,619,754]" pageId="1" pageNumber="596">
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<emphasis id="B3F13C70FFF72932FF61FDECFED071F8" bold="true" box="[151,262,619,645]" pageId="1" pageNumber="596">Material.</emphasis>
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<materialsCitation id="31EDEA3FFFF72932FEF9FDECFE3E71F8" box="[271,488,619,645]" collectionCode="LNU" pageId="1" pageNumber="596" specimenCode="LNU-22" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5E3E5EC0FFF72932FEF9FDECFEAC71F8" box="[271,378,619,645]" pageId="1" pageNumber="596">Holotype</typeStatus>
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,
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<specimenCode id="D1234819FFF72932FE70FDECFE3E71F8" box="[390,488,619,645]" pageId="1" pageNumber="596">LNU-22</specimenCode>
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</materialsCitation>
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, gender unknown, a nearly complete specimen, including the head, prothoracic leg, mesothoracic leg, metathoracic leg and terminal abdomen. The specimen is in fair condition, covered by a large crack from the head to the end of the abdomen.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C99FB3E9FFF72932FF31FD50FAAB718C" box="[199,1405,727,754]" pageId="1" pageNumber="596" type="discussion">
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<paragraph id="813AE062FFF72932FF31FD50FAAB718C" blockId="1.[151,1436,619,754]" box="[199,1405,727,754]" pageId="1" pageNumber="596">
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||
<emphasis id="B3F13C70FFF72932FF31FD50FE6F718C" bold="true" box="[199,441,727,753]" pageId="1" pageNumber="596">Locality and horizon.</emphasis>
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Hukawng Valley,
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<collectingRegion id="43412E80FFF72932FD8DFD50FCF8718C" box="[635,814,727,753]" country="Myanmar" name="Kachin" pageId="1" pageNumber="596">Kachin Province</collectingRegion>
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,
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<collectingCountry id="F992A0F2FFF72932FCCEFD5FFC75718F" box="[824,931,728,754]" name="Myanmar" pageId="1" pageNumber="596">Myanmar</collectingCountry>
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; lowermost Cenomanian, Upper Cretaceous.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="D5FAB0EAFFF72932FF61F96CFDE474CA" ID-DOI="http://doi.org/10.5281/zenodo.8073325" ID-Zenodo-Dep="8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="1" pageNumber="596" startId="1.[151,250,1771,1795]" targetBox="[177,1411,775,1737]" targetPageId="1" targetType="figure">
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<paragraph id="813AE062FFF72932FF61F96CFDE474CA" blockId="1.[151,1437,1771,1975]" pageId="1" pageNumber="596">
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||
<emphasis id="B3F13C70FFF72932FF61F96CFEC3747E" bold="true" box="[151,277,1771,1795]" pageId="1" pageNumber="596">FIGURE 1.</emphasis>
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Holotype (LNU-22) of
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<taxonomicName id="46859BE1FFF72932FDF9F96CFCC3747E" authority="Fan & Gu & Cao, 2023" authorityName="Fan & Gu & Cao" authorityYear="2023" box="[527,789,1771,1795]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="tenuicerci" status="sp. nov.">
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<emphasis id="B3F13C70FFF72932FDF9F96CFCC3747E" box="[527,789,1771,1795]" italics="true" pageId="1" pageNumber="596">Burmadactylus tenuicerci</emphasis>
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</taxonomicName>
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||
<emphasis id="B3F13C70FFF72932FCEBF96CFCB9747E" bold="true" box="[797,879,1771,1795]" pageId="1" pageNumber="596">
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||
<taxonomicNameLabel id="A8C2810BFFF72932FCEBF96CFCB9747E" box="[797,879,1771,1795]" pageId="1" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent.
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</paragraph>
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||
</caption>
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||
<subSubSection id="C99FB3E9FFF42931FF31FF11FDD1776C" pageId="2" pageNumber="597" type="description">
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||
<paragraph id="813AE062FFF42931FF31FF11FC5D7260" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
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<emphasis id="B3F13C70FFF42931FF31FF11FE8F73CD" bold="true" box="[199,345,150,176]" pageId="2" pageNumber="597">Description.</emphasis>
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<typeStatus id="5E3E5EC0FFF42931FE96FF10FE1D73CC" box="[352,459,151,177]" pageId="2" pageNumber="597">Holotype</typeStatus>
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, LNU-22 (
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||
<figureCitation id="19BEFCE7FFF42931FDB2FF10FD5373CC" box="[580,645,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1</figureCitation>
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), gender unknown, body
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<quantity id="467D4D87FFF42931FC50FF10FBDC73CC" box="[934,1034,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.12" pageId="2" pageNumber="597" unit="mm" value="3.12">3.12 mm</quantity>
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long measured from the head to the abdominal apex; pronotum
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<quantity id="467D4D87FFF42931FE39FF3CFDF173A8" box="[463,551,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="597" unit="mm" value="0.7">0.7 mm</quantity>
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||
long at midline; profemur
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||
<quantity id="467D4D87FFF42931FCADFF3CFC6473A8" box="[859,946,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="597" unit="mm" value="0.6">0.6 mm</quantity>
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long; protibia
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<quantity id="467D4D87FFF42931FBADFF3DFB6473A8" box="[1115,1202,186,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="597" unit="mm" value="0.5">0.5 mm</quantity>
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||
long; mesofemur
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<quantity id="467D4D87FFF42931FA88FF3CFF157384" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="597" unit="mm" value="1.1">1.1 mm</quantity>
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long; mesotibia1.0 mm long; metafemur
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<quantity id="467D4D87FFF42931FD7AFF58FD377384" box="[652,737,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="597" unit="mm" value="2.1">2.1 mm</quantity>
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||
long; metatibia
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||
<quantity id="467D4D87FFF42931FC65FF58FC3E7384" box="[915,1000,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="597" unit="mm" value="1.8">1.8 mm</quantity>
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long. The specimen is in fair condition, covered by a large fissure from the head to the end of the abdomen.
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||
</paragraph>
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||
<paragraph id="813AE062FFF42931FF31FEA0FEFF72F4" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
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||
Head. Hypognathous, capsulate; compound eyes large and well developed; ocelli invisible; antennae moniliform, 7 segments visible, flagellomere widening towards apex, inserted beneath the lower margin of the compound eyes (
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<figureCitation id="19BEFCE7FFF42931FF69FEE8FECA72F4" box="[159,284,367,393]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1A, D</figureCitation>
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).
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</paragraph>
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<paragraph id="813AE062FFF42931FF31FE14FAC172AC" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
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Thorax. The thorax dark brown; pronotum large, shield-like, extending posteriorly to entirely cover the metanotum; posterior margin of pronotum broadly rounded; tegmen and hindwing absent (
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<figureCitation id="19BEFCE7FFF42931FB78FE30FADD72AC" box="[1166,1291,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1A, D</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="813AE062FFF42931FF31FE5CFC427020" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
|
||
Leg. Prothoracic leg brown; profemur slender, distinctly shorter than mesothoracic leg; protibia robust, apex slightly inflated, with a sparse covering of thick setae and four strong teeth; protarsus two-segmented, slender, with second segment longer than first; basitarsus short; apical tarsomere elongate, slightly curved, with two claws. Mesothoracic leg brown; mesofemur slender, curved, over twice as long as the profemur, basally narrow and apically broad, with sparse setae on the ventral margins; mesotibia almost as long as mesofemur, middle inflated, with dorsal and ventral setae; mesotarsus almost identical to the protarsus. Metathoracic leg dark, saltatorial; metafemur slightly longer than abdomen and greatly inflated along its entire length, with prominent dorsal carina; metafemur mostly obscured by cracks, ventral with several irregularly spaced setae (
|
||
<figureCitation id="19BEFCE7FFF42931FC34FD50FBED718C" box="[962,1083,727,753]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1A, D</figureCitation>
|
||
); metatibia slightly shorter than metafemur, very slender, with tiny spines on the dorsal margins, without setae, lacking swimming plates, with two small subapical spurs and longer apical spurs; metatarsus slightly over twice as long as apical spurs, with a subapical denticular process; metatarsus ventral with sparse setae (
|
||
<figureCitation id="19BEFCE7FFF42931FCE7FCC4FC5E7020" box="[785,904,835,861]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1C, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="813AE062FFF42931FF31FCE0FDD1776C" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
|
||
Abdomen. The abdomen dark brown, with sparse and long setae; dorsal of abdomen obscured by cracks; subgenital plate broad, without styli, bearing a marginal fringe of long setae; cercus long, with numerous long setae, two-segmented, the second segment longer and distinctly slender than the first; paraproctal lobe cone-shaped, relatively small, almost the same as the first segment of cercus, covered with sparse setae, one of the setae near the apex thick and long (
|
||
<figureCitation id="19BEFCE7FFF42931FE75FC70FE2D776C" box="[387,507,1015,1041]" captionStart="FIGURE 1" captionStartId="1.[151,250,1771,1795]" captionTargetBox="[177,1411,775,1737]" captionTargetId="figure-103@1.[177,1411,775,1745]" captionTargetPageId="1" captionText="FIGURE 1. Holotype (LNU-22) of Burmadactylus tenuicerci sp. nov. holotype; (A) photograph of the habitus in the dorsal view, (B) photograph of the terminal abdomen (part of the setae near paraproctal lobe in this view are the setae of the cerci in the opposite side), (C) photograph of the left metatarsus, (D) line drawing of the habitus, (E) line drawing of the terminal abdomen, (F) line drawing of the left metatarsus. The abbreviations of the characteristics are as follows: prothoracic leg, pro; mesothoracic leg, meso; metafemur, mf; cercus, ce; paraproctal lobe, pptl; metatarsus, mt; apical spur, asp; subapical spur, sasp; subapical denticular process of metatarsus, sadent." figureDoi="http://doi.org/10.5281/zenodo.8073325" httpUri="https://zenodo.org/record/8073325/files/figure.png" pageId="2" pageNumber="597">Fig. 1B, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C99FB3E9FFF42931FF31FB9DFEDB7724" pageId="2" pageNumber="597" type="diagnosis">
|
||
<paragraph id="813AE062FFF42931FF31FB9DFEDB7724" blockId="2.[151,1437,150,1113]" pageId="2" pageNumber="597">
|
||
<emphasis id="B3F13C70FFF42931FF31FB9DFE947749" bold="true" box="[199,322,1050,1076]" pageId="2" pageNumber="597">Diagnosis.</emphasis>
|
||
The new species is similar to
|
||
<taxonomicName id="46859BE1FFF42931FD78FB9CFBE57748" authority="Heads, 2009" authorityName="Heads" authorityYear="2009" box="[654,1075,1051,1077]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="grimaldi">
|
||
<emphasis id="B3F13C70FFF42931FD78FB9CFC4B7748" box="[654,925,1051,1077]" italics="true" pageId="2" pageNumber="597">Burmadactylus grimaldi</emphasis>
|
||
<bibRefCitation id="E5149D93FFF42931FC52FB9CFBE57748" author="Heads, S. W." box="[932,1075,1051,1077]" pageId="2" pageNumber="597" pagination="75 - 82" refId="ref1726" refString="Heads, S. W. (2009) A new pygmy mole cricket in cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia, 26, 75 - 82." type="journal article" year="2009">Heads, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
. The major differences are listed in
|
||
<tableCitation id="CC07D5D9FFF42931FF45FBB8FED17724" box="[179,263,1087,1113]" captionStart="TABLE 1" captionStartId="2.[151,239,1160,1184]" captionTargetPageId="2" captionText="TABLE 1. Comparison of Burmadactylus tenuicerci sp. nov. and B. grimaldi Heads, 2009" httpUri="http://table.plazi.org/id/D5FAB0EAFFF42931FF61FB0FFB8B77DC" pageId="2" pageNumber="597" tableUuid="D5FAB0EAFFF42931FF61FB0FFB8B77DC">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D5FAB0EAFFF42931FF61FB0FFB8B77DC" ID-Table-UUID="D5FAB0EAFFF42931FF61FB0FFB8B77DC" box="[151,1117,1160,1185]" httpUri="http://table.plazi.org/id/D5FAB0EAFFF42931FF61FB0FFB8B77DC" pageId="2" pageNumber="597" startId="2.[151,239,1160,1184]" targetBox="[163,1386,1201,1417]" targetIsTable="true" targetPageId="2" targetType="table">
|
||
<paragraph id="813AE062FFF42931FF61FB0FFB8B77DC" blockId="2.[151,1117,1160,1185]" box="[151,1117,1160,1185]" pageId="2" pageNumber="597">
|
||
<emphasis id="B3F13C70FFF42931FF61FB0FFEDC77DD" bold="true" box="[151,266,1160,1184]" pageId="2" pageNumber="597">TABLE 1.</emphasis>
|
||
Comparison of
|
||
<taxonomicName id="46859BE1FFF42931FE41FB0FFD1F77DD" authority="Fan & Gu & Cao, 2023" authorityName="Fan & Gu & Cao" authorityYear="2023" box="[439,713,1160,1184]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="tenuicerci" status="sp. nov.">
|
||
<emphasis id="B3F13C70FFF42931FE41FB0FFD1F77DD" box="[439,713,1160,1184]" italics="true" pageId="2" pageNumber="597">Burmadactylus tenuicerci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3F13C70FFF42931FD39FB0EFCF377DC" bold="true" box="[719,805,1161,1185]" pageId="2" pageNumber="597">
|
||
<taxonomicNameLabel id="A8C2810BFFF42931FD39FB0EFCF377DC" box="[719,805,1161,1185]" pageId="2" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="46859BE1FFF42931FCAEFB0EFB8B77DC" authority="Heads, 2009" authorityName="Heads" authorityYear="2009" box="[856,1117,1160,1185]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="grimaldi">
|
||
<emphasis id="B3F13C70FFF42931FCAEFB0EFC0477DD" box="[856,978,1160,1184]" italics="true" pageId="2" pageNumber="597">B. grimaldi</emphasis>
|
||
<bibRefCitation id="E5149D93FFF42931FC2EFB0FFB8B77DC" author="Heads, S. W." box="[984,1117,1160,1185]" pageId="2" pageNumber="597" pagination="75 - 82" refId="ref1726" refString="Heads, S. W. (2009) A new pygmy mole cricket in cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia, 26, 75 - 82." type="journal article" year="2009">Heads, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="813AE062FFF42931FF5FFB36FB3E7618" pageId="2" pageNumber="597">
|
||
<table id="F38512C2FFF4D6CCFF55FB36FABC76F4" box="[163,1386,1201,1417]" gridcols="3" gridrows="4" pageId="2" pageNumber="597">
|
||
<tr id="3FB5E220FFF4D6CCFF55FB36FABC77B4" box="[163,1386,1201,1225]" gridrow="0" pageId="2" pageNumber="597">
|
||
<th id="7C648B5CFFF4D6CCFF55FB36FE9E77B4" box="[163,328,1201,1225]" gridcol="0" gridrow="0" pageId="2" pageNumber="597">Characters</th>
|
||
<th id="7C648B5CFFF4D6CCFE68FB36FC8777B4" box="[414,849,1201,1225]" gridcol="1" gridrow="0" pageId="2" pageNumber="597">
|
||
<taxonomicName id="46859BE1FFF42931FE68FB36FD7477B5" authority="Fan & Gu & Cao, 2023" authorityName="Fan & Gu & Cao" authorityYear="2023" box="[414,674,1201,1225]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="tenuicerci" status="sp. nov.">
|
||
<emphasis id="B3F13C70FFF42931FE68FB36FD7477B5" box="[414,674,1201,1225]" italics="true" pageId="2" pageNumber="597">Burmadactylus tenuicerci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3F13C70FFF42931FD5EFB36FD2F77B4" bold="true" box="[680,761,1201,1225]" pageId="2" pageNumber="597">
|
||
<taxonomicNameLabel id="A8C2810BFFF42931FD5EFB36FD2F77B4" box="[680,761,1201,1225]" pageId="2" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="7C648B5CFFF4D6CCFC77FB36FABC77B4" box="[897,1386,1201,1225]" gridcol="2" gridrow="0" pageId="2" pageNumber="597">
|
||
<emphasis id="B3F13C70FFF42931FC77FB36FBAE77B4" box="[897,1144,1201,1225]" italics="true" pageId="2" pageNumber="597">Burmadactylus grimaldi</emphasis>
|
||
<bibRefCitation id="E5149D93FFF42931FB89FB36FB2877B4" author="Heads, S. W." box="[1151,1278,1201,1225]" pageId="2" pageNumber="597" pagination="75 - 82" refId="ref1726" refString="Heads, S. W. (2009) A new pygmy mole cricket in cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia, 26, 75 - 82." type="journal article" year="2009">Heads, 2009</bibRefCitation>
|
||
</th>
|
||
</tr>
|
||
<tr id="3FB5E220FFF4D6CCFF55FB5EFABC7668" box="[163,1386,1241,1301]" gridrow="1" pageId="2" pageNumber="597">
|
||
<th id="7C648B5CFFF4D6CCFF55FB5EFE9E7668" box="[163,328,1241,1301]" gridcol="0" gridrow="1" pageId="2" pageNumber="597">Mesotibia</th>
|
||
<td id="7C648B5CFFF4D6CCFE68FB5EFC877668" box="[414,849,1241,1301]" gridcol="1" gridrow="1" pageId="2" pageNumber="597">Basally inflated and almost as long as mesofemur</td>
|
||
<td id="7C648B5CFFF4D6CCFC77FB5EFABC7668" box="[897,1386,1241,1301]" gridcol="2" gridrow="1" pageId="2" pageNumber="597">Basally not inflated and shorter than mesofemur</td>
|
||
</tr>
|
||
<tr id="3FB5E220FFF4D6CCFF55FAA2FABC7643" box="[163,1386,1317,1342]" gridrow="2" pageId="2" pageNumber="597">
|
||
<th id="7C648B5CFFF4D6CCFF55FAA2FE9E7643" box="[163,328,1317,1342]" gridcol="0" gridrow="2" pageId="2" pageNumber="597">Cercus</th>
|
||
<td id="7C648B5CFFF4D6CCFE68FAA2FC877643" box="[414,849,1317,1342]" gridcol="1" gridrow="2" pageId="2" pageNumber="597">Second segment distinctly slender</td>
|
||
<td id="7C648B5CFFF4D6CCFC77FAA2FABC7643" box="[897,1386,1317,1342]" gridcol="2" gridrow="2" pageId="2" pageNumber="597">Second segment not slender</td>
|
||
</tr>
|
||
<tr id="3FB5E220FFF4D6CCFF55FACAFABC76F4" box="[163,1386,1357,1417]" gridrow="3" pageId="2" pageNumber="597">
|
||
<th id="7C648B5CFFF4D6CCFF55FACAFE9E76F4" box="[163,328,1357,1417]" gridcol="0" gridrow="3" pageId="2" pageNumber="597">Paraproctal lobe</th>
|
||
<td id="7C648B5CFFF4D6CCFE68FACAFC8776F4" box="[414,849,1357,1417]" gridcol="1" gridrow="3" pageId="2" pageNumber="597">Covered with sparse setae, one of the setae located the apex thick and long</td>
|
||
<td id="7C648B5CFFF4D6CCFC77FACAFABC76F4" box="[897,1386,1357,1417]" gridcol="2" gridrow="3" pageId="2" pageNumber="597">Covered with dense and short setae</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C99FB3E9FFF42931FF31FA5AFD5D7566" pageId="2" pageNumber="597" type="etymology">
|
||
<paragraph id="813AE062FFF42931FF31FA5AFD5D7566" blockId="2.[151,1436,1501,1816]" pageId="2" pageNumber="597">
|
||
<emphasis id="B3F13C70FFF42931FF31FA5AFE98768A" bold="true" box="[199,334,1501,1527]" pageId="2" pageNumber="597">Etymology.</emphasis>
|
||
The specific epithet is from the Latin ‘tenui’ and ‘cercus’, used to describe the slender and elongated second segment of cercus in
|
||
<emphasis id="B3F13C70FFF42931FE2CF986FD5D7566" box="[474,651,1537,1563]" italics="true" pageId="2" pageNumber="597">
|
||
<taxonomicName id="46859BE1FFF42931FE2CF986FD517566" authorityName="Heads" authorityYear="2009" box="[474,647,1537,1563]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="genus">Burmadactylus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C99FB3E9FFF42931FF31F9A2FC01746A" pageId="2" pageNumber="597" type="discussion">
|
||
<paragraph id="813AE062FFF42931FF31F9A2FC01746A" blockId="2.[151,1436,1501,1816]" pageId="2" pageNumber="597">
|
||
<emphasis id="B3F13C70FFF42931FF31F9A2FEED7542" bold="true" box="[199,315,1573,1599]" pageId="2" pageNumber="597">Remarks.</emphasis>
|
||
Owing to the metatarsus of this specimen with a subapical denticular process, it should be assigned in
|
||
<taxonomicName id="46859BE1FFF42931FF45F9CEFDCE751E" authority="Gunther, 1979" authorityName="Gunther" authorityYear="1979" box="[179,536,1609,1635]" class="Insecta" family="Tridactylidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="subFamily" subFamily="Dentridactylinae">
|
||
Dentridactylinae
|
||
<bibRefCitation id="E5149D93FFF42931FE82F9CEFDCE751E" author="Gunther, K. K." box="[372,536,1609,1635]" pageId="2" pageNumber="597" pagination="255 - 264" refId="ref1670" refString="Gunther, K. K. (1979) Einige Bemerkungen uber die Gattungen der Familie Tridactylidae Brunner und zur Klassifikation der Tridactylodea (Orthopteroida, Caelifera). Deutsche Entomologische Zeitschrift, 26 (4 - 5), 255 - 264. https: // doi. org / 10.1002 / mmnd. 4800260408" type="journal article" year="1979">Günther, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="46859BE1FFF42931FDD4F9CDFD66751E" authority="Fan & Gu & Cao, 2023" authorityName="Fan & Gu & Cao" authorityYear="2023" box="[546,688,1610,1635]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="tenuicerci" status="sp. nov.">
|
||
<emphasis id="B3F13C70FFF42931FDD4F9CDFD66751E" box="[546,688,1610,1635]" italics="true" pageId="2" pageNumber="597">B. tenuicerci</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B3F13C70FFF42931FD40F9CEFCD9751E" bold="true" box="[694,783,1609,1635]" pageId="2" pageNumber="597">
|
||
<taxonomicNameLabel id="A8C2810BFFF42931FD40F9CEFCD9751E" box="[694,783,1609,1635]" pageId="2" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from other fossil
|
||
<taxonomicName id="46859BE1FFF42931FB7CF9CEFA93751E" authorityName="Gunther" authorityYear="1979" box="[1162,1349,1609,1635]" class="Insecta" family="Tridactylidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="subFamily" subFamily="Dentridactylinae">Dentridactylinae</taxonomicName>
|
||
species by presence of small paraproctal lobe which are much longer (as long as the cercus) in other members of the subfamily (
|
||
<bibRefCitation id="E5149D93FFF42931FEE1F916FE3175D6" author="Azar, D. & Nel, A." box="[279,487,1681,1707]" pageId="2" pageNumber="597" pagination="169 - 175" refId="ref1637" refString="Azar, D. & Nel, A. (2008) First Tridactylidae from the Eocene French amber (Insecta: Orthoptera). Alavesia, 2, 169 - 175." type="journal article" year="2008">Azar & Nel, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E5149D93FFF42931FE05F916FD5175D6" author="Poinar, G." box="[499,647,1681,1707]" pageId="2" pageNumber="597" pagination="238 - 243" refId="ref1759" refString="Poinar, G. (2020) Pygmy mole crickets (Orthoptera: Tridactylidae) in Dominican and Burmese amber. Historical Biology, 32 (2), 238 - 243. https: // doi. org / 10.1080 / 08912963.2018.1483362" type="journal article" year="2020">Poinar, 2020</bibRefCitation>
|
||
). This new species can be assigned to
|
||
<taxonomicName id="46859BE1FFF42931FBCCF916FAAE75D6" authority="Heads, 2009" authorityName="Heads" authorityYear="2009" box="[1082,1400,1681,1707]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3F13C70FFF42931FBCCF916FB3275D6" box="[1082,1252,1681,1707]" italics="true" pageId="2" pageNumber="597">Burmadactylus</emphasis>
|
||
<bibRefCitation id="E5149D93FFF42931FB1DF916FAAE75D6" author="Heads, S. W." box="[1259,1400,1681,1707]" pageId="2" pageNumber="597" pagination="75 - 82" refId="ref1726" refString="Heads, S. W. (2009) A new pygmy mole cricket in cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia, 26, 75 - 82." type="journal article" year="2009">Heads, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
by the following characters: paraproctal lobe small, conical shape; swimming plates absent; margin of subgenital plate with long setae; metatarsus slightly over twice as long as apical spurs. The differences between the new species and the
|
||
<typeStatus id="5E3E5EC0FFF42931FF36F979FF267465" box="[192,240,1790,1816]" pageId="2" pageNumber="597">type</typeStatus>
|
||
species
|
||
<taxonomicName id="46859BE1FFF42931FEB9F979FDBC746A" authority="Heads, 2009" authorityName="Heads" authorityYear="2009" box="[335,618,1789,1815]" class="Insecta" family="Tridactylidae" genus="Burmadactylus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis id="B3F13C70FFF42931FEB9F979FE01746A" box="[335,471,1789,1815]" italics="true" pageId="2" pageNumber="597">B. grimaldii</emphasis>
|
||
<bibRefCitation id="E5149D93FFF42931FE28F97AFDBC746A" author="Heads, S. W." box="[478,618,1789,1815]" pageId="2" pageNumber="597" pagination="75 - 82" refId="ref1726" refString="Heads, S. W. (2009) A new pygmy mole cricket in cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia, 26, 75 - 82." type="journal article" year="2009">Heads, 2009</bibRefCitation>
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</taxonomicName>
|
||
are demonstrated in the
|
||
<tableCitation id="CC07D5D9FFF42931FC8BF97AFC07746A" box="[893,977,1789,1815]" captionStart="TABLE 1" captionStartId="2.[151,239,1160,1184]" captionTargetPageId="2" captionText="TABLE 1. Comparison of Burmadactylus tenuicerci sp. nov. and B. grimaldi Heads, 2009" httpUri="http://table.plazi.org/id/D5FAB0EAFFF42931FF61FB0FFB8B77DC" pageId="2" pageNumber="597" tableUuid="D5FAB0EAFFF42931FF61FB0FFB8B77DC">Table 1</tableCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |