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<document id="92222A988CEF3206D3F10387EA78DB1F" ID-CLB-Dataset="51311" ID-DOI="10.5281/zenodo.156493" ID-GBIF-Dataset="c3fd5e49-3b05-4a99-9bc8-455ae3056faa" ID-ISSN="1175­5326" ID-Zenodo-Dep="156493" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459617947450" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="0846107D833A6D4BFEFEFCCBFB72FDE4" docLanguage="en" docName="zt00239.pdf" docOrigin="Zootaxa 239" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Stethusa Casey 1910" docType="treatment" docVersion="13" lastPageNumber="11" masterDocId="F47F6805833E6D41FFF6FFADFFFCFFAF" masterDocTitle="A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="5" updateTime="1698204628127" updateUser="ExternalLinkService">
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<mods:title id="CA34A956ACAB765BBAC57C0C952A4085">A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="8819FA29C986A37AFBEEC3EFC1AA65A9">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="D48DBDD08C93F34A9A56B56722AF666C">2003</mods:date>
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<treatment id="0846107D833A6D4BFEFEFCCBFB72FDE4" ID-DOI="http://doi.org/10.5281/zenodo.6275921" ID-GBIF-Taxon="119331776" ID-Zenodo-Dep="6275921" LSID="urn:lsid:plazi:treatment:0846107D833A6D4BFEFEFCCBFB72FDE4" httpUri="http://treatment.plazi.org/id/0846107D833A6D4BFEFEFCCBFB72FDE4" lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5">
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<taxonomicName id="47EFDAE8833A6D45FEFEFCCBFDF5FC2F" ID-CoL="8VS4C" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[264,521,870,896]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B29B7D79833A6D45FEFEFCCBFE93FC2F" bold="true" box="[264,367,870,896]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
<bibRefCitation id="E47EDC9A833A6D45FE80FCCBFDF5FC2F" author="Casey" box="[374,521,870,896]" pageId="4" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
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<figureCitation id="18D4BDEE833A6D45FEE7FC24FE6BFC0C" box="[273,407,905,931]" captionStart-0="FIGURES 1 ­ 5" captionStart-1="FIGURE 6" captionStart-10="FIGURES 48 ­ 52" captionStart-11="FIGURES 53 ­ 57" captionStart-12="FIGURES 58 ­ 62" captionStart-13="FIGURES 63 ­ 70" captionStart-14="FIGURES 71 ­ 75" captionStart-15="FIGURES 76 ­ 83" captionStart-16="FIGURES 84 ­ 87" captionStart-17="FIGURES 88 ­ 92" captionStart-18="FIGURES 93 ­ 97" captionStart-19="FIGURES 98 ­ 100" captionStart-2="FIGURES 7 ­ 13" captionStart-20="FIGURES 101 ­ 105" captionStart-21="FIGURES 106 ­ 109" captionStart-3="FIGURES 14 ­ 17" captionStart-4="FIGURES 18 ­ 21" captionStart-5="FIGURES 22 ­ 28" captionStart-6="FIGURES 29 ­ 33" captionStart-7="FIGURES 34 ­ 39" captionStart-8="FIGURES 40 ­ 42" captionStart-9="FIGURES 43 ­ 47" captionStartId-0="5.[264,382,1536,1560]" captionStartId-1="6.[264,368,1296,1320]" captionStartId-10="22.[264,382,1402,1426]" captionStartId-11="23.[264,382,1144,1168]" captionStartId-12="25.[264,382,1456,1480]" captionStartId-13="27.[264,382,1254,1278]" captionStartId-14="30.[264,382,1719,1743]" captionStartId-15="31.[264,382,1309,1333]" captionStartId-16="33.[264,382,1203,1227]" captionStartId-17="34.[264,382,1775,1799]" captionStartId-18="35.[264,382,1460,1484]" captionStartId-19="36.[264,382,1092,1116]" captionStartId-2="8.[264,382,1402,1426]" captionStartId-20="37.[264,382,1751,1775]" captionStartId-21="38.[264,382,1692,1716]" captionStartId-3="9.[264,382,1243,1267]" captionStartId-4="10.[264,382,1704,1728]" captionStartId-5="16.[264,382,1471,1495]" captionStartId-6="17.[264,382,1348,1372]" captionStartId-7="18.[264,382,1757,1781]" captionStartId-8="19.[264,382,1234,1258]" captionStartId-9="21.[264,382,1284,1308]" captionTargetBox-0="[291,1293,316,1520]" captionTargetBox-1="[379,1208,514,1282]" captionTargetBox-10="[288,1293,492,1373]" captionTargetBox-11="[284,1298,315,1116]" captionTargetBox-12="[301,1277,317,1437]" captionTargetBox-13="[306,1276,315,1222]" captionTargetBox-14="[290,1298,484,1683]" captionTargetBox-15="[300,1285,321,1270]" captionTargetBox-16="[322,1264,569,1172]" captionTargetBox-17="[312,1273,794,1764]" captionTargetBox-18="[291,1293,318,1425]" captionTargetBox-19="[345,1241,404,1068]" captionTargetBox-2="[309,1277,513,1386]" captionTargetBox-20="[272,1313,825,1750]" captionTargetBox-21="[347,1242,1083,1660]" captionTargetBox-3="[347,1240,317,1218]" captionTargetBox-4="[348,1242,678,1683]" captionTargetBox-5="[294,1289,322,1437]" captionTargetBox-6="[291,1305,315,1327]" captionTargetBox-7="[284,1296,654,1714]" captionTargetBox-8="[344,1251,575,1191]" captionTargetBox-9="[327,1258,319,1272]" captionTargetId-0="figure@5.[281,1305,304,1529]" captionTargetId-1="figure@6.[379,1209,511,1287]" captionTargetId-10="figure@22.[278,1309,478,1384]" captionTargetId-11="figure@23.[281,1306,308,1121]" captionTargetId-12="figure@25.[296,1290,308,1444]" captionTargetId-13="figure@27.[298,1289,306,1231]" captionTargetId-14="figure@30.[282,1305,470,1692]" captionTargetId-15="figure@31.[293,1295,314,1282]" captionTargetId-16="figure@33.[318,1269,566,1177]" captionTargetId-17="figure@34.[302,1284,788,1767]" captionTargetId-18="figure@35.[281,1305,308,1436]" captionTargetId-19="figure@36.[340,1247,395,1076]" captionTargetId-2="figure@8.[302,1286,504,1395]" captionTargetId-20="figure@37.[264,1322,815,1750]" captionTargetId-21="figure@38.[337,1249,1078,1665]" captionTargetId-3="figure@9.[337,1249,307,1227]" captionTargetId-4="figure@10.[338,1248,670,1691]" captionTargetId-5="figure@16.[287,1300,315,1445]" captionTargetId-6="figure@17.[270,1317,306,1339]" captionTargetId-7="figure@18.[274,1313,646,1730]" captionTargetId-8="figure@19.[335,1251,566,1207]" captionTargetId-9="figure@21.[319,1267,312,1281]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="22" captionTargetPageId-11="23" captionTargetPageId-12="25" captionTargetPageId-13="27" captionTargetPageId-14="30" captionTargetPageId-15="31" captionTargetPageId-16="33" captionTargetPageId-17="34" captionTargetPageId-18="35" captionTargetPageId-19="36" captionTargetPageId-2="8" captionTargetPageId-20="37" captionTargetPageId-21="38" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="16" captionTargetPageId-6="17" captionTargetPageId-7="18" captionTargetPageId-8="19" captionTargetPageId-9="21" captionText-0="FIGURES 1 ­ 5. Mouthparts of Stethusa dichroa (Gravenhorst) (female (1, 5) and male (3 ­ 4), Eddyville, Illinois; female, 18 km SSE Lawrence, Kansas (2 )). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURE 6. Nomenclature of setae and pores in epipharynx of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). Scale bar 0.1 mm." captionText-10="FIGURES 48 ­ 52. Aedeagus of Stethusa klimschi (Bernhauer) (lectotype (48 ­ 51); and male from Vicksburg, Mississippi (52 )). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view; 52 – apex of left paramere. Scale bar 0.1 mm (49, 51 ­ 52), 0.2 mm (48, 50)." captionText-11="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." captionText-12="FIGURES 58 ­ 62. Abdominal segment 8 of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (58 ­ 60); and female 9 km W Buffalo Gap, Virginia (61 ­ 62 )). 58 – male tergum 8; 59 – male sternum 8; 60 – apex of male tergum 8; 61 – female tergum 8; 62 – female sternum 8. Scale bar 0.2 mm (58 ­ 59, 61 ­ 62), 0.1 mm (60)." captionText-13="FIGURES 63 ­ 70. Aedeagus and spermatheca of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (63 ­ 69); and female 9 km W Buffalo Gap, Virginia (70 )). 63 – median lobe, parameral view; 64 – apex of median lobe, parameral view; 65 – median lobe, lateral view; 66 – apex of median lobe, lateral view; 67 – details of retracted internal sac, abparameral view; 68 – details of retracted internal sac, lateral view; 69 – apex of left paramere; 70 – spermatheca. Scale bar 0.1 mm (64, 66, 69 ­ 70), 0.2 mm (63, 65, 67 ­ 68)." captionText-14="FIGURES 71 ­ 75. Abdominal segment 8 of Stethusa luederwaldti (Bernhauer) (lectotype, male (71 ­ 73); and female paralectotype (74 ­ 75 )). 71 – male tergum 8; 72 – male sternum 8; 73 – apex of male tergum 8; 74 – female tergum 8; 75 – female sternum 8. Scale bar 0.2 mm (71 ­ 72, 74 ­ 75), 0.1 mm (73)." captionText-15="FIGURES 76 ­ 83. Aedeagus and spermatheca of Stethusa luederwaldti (Bernhauer) (lectotype, male (76 ­ 82); and female paralectotype (83 )). 76 – median lobe, parameral view; 77 – apex of median lobe, parameral view; 78 – median lobe, lateral view; 79 – apex of median lobe, lateral view; 80 – details of retracted internal sac, abparameral view; 81 – details of retracted internal sac, lateral view; 82 – apex of left paramere; 83 – spermatheca. Scale bar 0.1 mm (77, 79, 82 ­ 83), 0.2 mm (76, 78, 80 ­ 81)." captionText-16="FIGURES 84 ­ 87. Abdominal segment 8 and spermatheca of Atheta texana (Casey) (holotype of At. mendosa Casey (84 ­ 86), lectotype of At. texana (87 )). 84 – female tergum 8; 85 – female sternum 8; 86 ­ 87 – spermatheca. Scale bar 0.2 mm (86 ­ 87), 0.4 mm (84 ­ 85)." captionText-17="FIGURES 88 ­ 92. Abdominal segment 8 of Atheta crenuliventris Bernhauer (lectotype of At. crenuliventris, male (88 ­ 90); and female paratype of Dimetrota bradorensis Lohse (91 ­ 92 )). 88 – male tergum 8; 89 – male sternum 8; 90 – apex of male tergum 8; 91 – female tergum 8; 92 – female sternum 8. Scale bar 0.4 mm (88 ­ 89, 91 ­ 92), 0.2 mm (90)." captionText-18="FIGURES 93 ­ 97. Aedeagus and spermatheca of Atheta crenuliventris Bernhauer (lectotype of At. crenuliventris, male (93 ­ 96); and female paratype of Dimetrota bradorensis Lohse (97 )). 93 – median lobe, parameral view; 94 – apex of median lobe, parameral view; 95 – median lobe, lateral view; 96 – apex of median lobe, lateral view; 97 – spermatheca. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.1 mm (94, 96), 0.2 mm (93, 95, 97)." captionText-19="FIGURES 98 ­ 100. Details of retracted internal sac of Atheta crenuliventris Bernhauer (lectotype). 98 ­ 99 – abparameral view; 100 – lateral view. Scale bar 0.2 mm." captionText-2="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." captionText-20="FIGURES 101 ­ 105. Abdominal segment 8 of Atheta iheringi Bernhauer (lectotype, male (101 ­ 103); and female paralectotype (104 ­ 105 )). 101 – male tergum 8; 102 – male sternum 8; 103 – apex of male tergum 8; 104 – female tergum 8; 105 – female sternum 8. Scale bar 0.4 mm (101 ­ 102, 104 ­ 105), 0.2 mm (103)." captionText-21="FIGURES 106 ­ 109. Aedeagus and spermatheca of Atheta iheringi Bernhauer (lectotype, male (106 ­ 108); and female paralectotype (109 )). 106 – median lobe, parameral view; 107 – median lobe, lateral view; 108 – apex of median lobe, lateral view; 109 – spermatheca. Scale bar 0.1 mm (108), 0.2 mm (106 ­ 107, 109)." captionText-3="FIGURES 14 ­ 17. Details of Stethusa dichroa (Gravenhorst) (male (14 ­ 16) and female (17), Eddyville, Illinois). 14 – right mesotibia; 15 – antennal article 11; 16 – right antenna; 17 – right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16)." captionText-4="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." captionText-5="FIGURES 22 ­ 28. Abdominal segment 8 of Stethusa dichroa (Gravenhorst) (18 km SSE Lawrence, Kansas (22 ­ 23, 25); 12 km NNE Lawrence (24, 27); 2 km N Eddyville, Illinois (26, 28 )). 22 – male tergum 8; 23 – male sternum 8; 24 ­ 26 – apex of male tergum 8; 27 – female tergum 8; 28 – female sternum 8. Scale bar 0.2 mm (22 ­ 23, 27 ­ 28), 0.1 mm (24 ­ 26)." captionText-6="FIGURES 29 ­ 33. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (29 ­ 32); 18 km SSE Lawrence (33 )). 29 – median lobe, parameral view; 30 – apex of median lobe, parameral view; 31 – median lobe, lateral view; 32 – apex of median lobe, lateral view; 33 – apex of left paramere. Scale bar 0.1 mm (30, 32 ­ 33), 0.2 mm (29, 31)." captionText-7="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." captionText-8="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 – female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 – spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." captionText-9="FIGURES 43 ­ 47. Abdominal segment 8 of Stethusa klimschi (Bernhauer) (male (43 ­ 45) and female (46 ­ 47) from Vicksburg, Mississippi). 43 – male tergum 8; 44 – male sternum 8; 45 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (43 ­ 44, 46 ­ 47), 0.2 mm (45)." httpUri-0="https://zenodo.org/record/156494/files/figure.png" httpUri-1="https://zenodo.org/record/156495/files/figure.png" httpUri-10="https://zenodo.org/record/156504/files/figure.png" httpUri-11="https://zenodo.org/record/156505/files/figure.png" httpUri-12="https://zenodo.org/record/156506/files/figure.png" httpUri-13="https://zenodo.org/record/156507/files/figure.png" httpUri-14="https://zenodo.org/record/156508/files/figure.png" httpUri-15="https://zenodo.org/record/156509/files/figure.png" httpUri-16="https://zenodo.org/record/156510/files/figure.png" httpUri-17="https://zenodo.org/record/156511/files/figure.png" httpUri-18="https://zenodo.org/record/156512/files/figure.png" httpUri-19="https://zenodo.org/record/156513/files/figure.png" httpUri-2="https://zenodo.org/record/156496/files/figure.png" httpUri-20="https://zenodo.org/record/156514/files/figure.png" httpUri-21="https://zenodo.org/record/156515/files/figure.png" httpUri-3="https://zenodo.org/record/156497/files/figure.png" httpUri-4="https://zenodo.org/record/156498/files/figure.png" httpUri-5="https://zenodo.org/record/156499/files/figure.png" httpUri-6="https://zenodo.org/record/156500/files/figure.png" httpUri-7="https://zenodo.org/record/156501/files/figure.png" httpUri-8="https://zenodo.org/record/156502/files/figure.png" httpUri-9="https://zenodo.org/record/156503/files/figure.png" pageId="4" pageNumber="5">Figs. 1­109</figureCitation>
)
</paragraph>
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<taxonomicName id="47EFDAE8833A6D45FEFEFC6AFEB1FC71" ID-CoL="8VRBL" box="[264,333,967,990]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFC6AFEB1FC71" box="[264,333,967,990]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
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(
<taxonomicName id="47EFDAE8833A6D45FEA8FC6AFDBAFC71" ID-CoL="8VS4C" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[350,582,967,990]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEA8FC6AFE4BFC71" box="[350,439,967,990]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
<bibRefCitation id="E47EDC9A833A6D45FE49FC6AFDBAFC71" author="Casey" box="[447,582,967,990]" pageId="4" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
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): 4 (
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species:
<taxonomicName id="47EFDAE8833A6D45FCE1FC6AFBC4FC71" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[791,1080,967,990]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="irvingi">
<emphasis id="B29B7D79833A6D45FCE1FC6AFC57FC71" box="[791,939,967,990]" italics="true" pageId="4" pageNumber="5">Atheta irvingi</emphasis>
<bibRefCitation id="E47EDC9A833A6D45FC45FC6AFBC4FC71" author="Casey" box="[947,1080,967,990]" pageId="4" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
, designated by
<bibRefCitation id="E47EDC9A833A6D45FB29FC6AFE77FC51" author="Fenyes" pageId="4" pageNumber="42" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes (1918))</bibRefCitation>
.
</paragraph>
<paragraph id="8050A16B833A6D45FEFEFBAAFC91FBB1" blockId="4.[264,1322,967,1278]" box="[264,877,1031,1054]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFBAAFC91FBB1" box="[264,877,1031,1054]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFBAAFE3FFBB1" ID-CoL="8VRBL" authority="(Stethusa)" baseAuthorityName="Stethusa" box="[264,451,1031,1054]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFBAAFEB1FBB1" box="[264,333,1031,1054]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FEABFBAAFE49FBB1" box="[349,437,1031,1054]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="014E877A833A6D45FE23FBAAFD6DFBB1" author="Casey" box="[469,657,1031,1054]" page="77" pageId="4" pageNumber="5" year="1911">
<bibRefCitation id="E47EDC9A833A6D45FE23FBAAFD9AFBB1" author="Casey" box="[469,614,1031,1054]" pageId="4" pageNumber="42" refString="Casey, T. L. (1911 a) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, 245 pp." type="book" year="1911" yearSuffix="a">Casey, 1911a</bibRefCitation>
: 77
</treatmentCitation>
(as valid subgenus).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E0833A6D45FEFEFB8AFD65FB91" box="[264,665,1063,1086]" pageId="4" pageNumber="5" type="description">
<paragraph id="8050A16B833A6D45FEFEFB8AFD65FB91" blockId="4.[264,1322,967,1278]" box="[264,665,1063,1086]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFB8AFD65FB91" box="[264,665,1063,1086]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFB8AFE3EFB91" ID-CoL="8VRBL" authority="(Stethusa)" baseAuthorityName="Stethusa" box="[264,450,1063,1086]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFB8AFEB1FB91" box="[264,333,1063,1086]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FEABFB8AFE49FB91" box="[349,437,1063,1086]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="014E877A833A6D45FE23FB8AFD68FB91" author="Fenyes" box="[469,660,1063,1086]" page="25" pageId="4" pageNumber="5" year="1918">
<bibRefCitation id="E47EDC9A833A6D45FE23FB8AFD9AFB91" author="Fenyes" box="[469,614,1063,1086]" pageId="4" pageNumber="42" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
: 25
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E0833A6D45FEFEFBEAFB13FBD1" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8050A16B833A6D45FEFEFBEAFB3EFBF1" blockId="4.[264,1322,967,1278]" box="[264,1218,1095,1118]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFBEAFB3EFBF1" box="[264,1218,1095,1118]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFBEAFE3FFBF1" ID-CoL="8VRBL" authority="(Stethusa)" baseAuthorityName="Stethusa" box="[264,451,1095,1118]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFBEAFEB1FBF1" box="[264,333,1095,1118]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FEABFBEAFE49FBF1" box="[349,437,1095,1118]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="014E877A833A6D45FE23FBEAFD63FBF1" author="Fenyes" box="[469,671,1095,1118]" page="206" pageId="4" pageNumber="5" year="1920">
<bibRefCitation id="E47EDC9A833A6D45FE23FBEAFD9BFBF1" author="Fenyes" box="[469,615,1095,1118]" pageId="4" pageNumber="42" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 206
</treatmentCitation>
(as synonym of
<taxonomicName id="47EFDAE8833A6D45FCA7FBEAFB42FBF1" ID-CoL="8VRBL" authority="Hypatheta Fenyes, 1918" authorityName="Hypatheta Fenyes" authorityYear="1918" box="[849,1214,1095,1118]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FCA7FBEAFC6BFBF1" box="[849,919,1095,1118]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FC50FBEAFBE9FBF1" box="[934,1045,1095,1118]" italics="true" pageId="4" pageNumber="5">Hypatheta</emphasis>
<bibRefCitation id="E47EDC9A833A6D45FBEAFBEAFB50FBF1" author="Fenyes" box="[1052,1196,1095,1118]" pageId="4" pageNumber="42" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8050A16B833A6D45FEFEFBCAFB13FBD1" blockId="4.[264,1322,967,1278]" box="[264,1263,1127,1150]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFBCAFB13FBD1" box="[264,1263,1127,1150]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFBCAFE3FFBD1" ID-CoL="8VRBL" authority="(Stethusa)" baseAuthorityName="Stethusa" box="[264,451,1127,1150]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFBCAFEB1FBD1" box="[264,333,1127,1150]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FEABFBCAFE49FBD1" box="[349,437,1127,1150]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="014E877A833A6D45FE23FBCAFC9DFBD1" author="Bernhauer" box="[469,865,1127,1150]" page="652" pageId="4" pageNumber="5" year="1926">
<bibRefCitation id="E47EDC9A833A6D45FE23FBCAFCD4FBD1" author="Bernhauer" box="[469,808,1127,1150]" pageId="4" pageNumber="42" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 652
</treatmentCitation>
(as synonym of
<taxonomicName id="47EFDAE8833A6D45FBE5FBCAFB17FBD1" ID-CoL="8VRBL" authority="Hypatheta" authorityName="Hypatheta" box="[1043,1259,1127,1150]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FBE5FBCAFBA4FBD1" box="[1043,1112,1127,1150]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FB9EFBCAFB2BFBD1" box="[1128,1239,1127,1150]" italics="true" pageId="4" pageNumber="5">Hypatheta</emphasis>
))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E0833A6D45FEFEFB2AFC1FFB31" box="[264,995,1159,1182]" pageId="4" pageNumber="5" type="description">
<paragraph id="8050A16B833A6D45FEFEFB2AFC1FFB31" blockId="4.[264,1322,967,1278]" box="[264,995,1159,1182]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFB2AFC1FFB31" box="[264,995,1159,1182]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFB2AFE3EFB31" ID-CoL="8VRBL" authority="(Stethusa)" baseAuthorityName="Stethusa" box="[264,450,1159,1182]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFB2AFEB1FB31" box="[264,333,1159,1182]" italics="true" pageId="4" pageNumber="5">Atheta</emphasis>
(
<emphasis id="B29B7D79833A6D45FEABFB2AFE49FB31" box="[349,437,1159,1182]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="014E877A833A6D45FE23FB2AFCFBFB31" author="Moore" box="[469,775,1159,1182]" page="353" pageId="4" pageNumber="5" year="1975">
<bibRefCitation id="E47EDC9A833A6D45FE23FB2AFD32FB31" author="Moore" box="[469,718,1159,1182]" pageId="4" pageNumber="43" refString="Moore, I. &amp; Legner, E. F. (1975) A Catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp." type="book" year="1975">Moore &amp; Legner, 1975</bibRefCitation>
: 353
</treatmentCitation>
(as valid subgenus).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E0833A6D45FEFEFB0AFE0DFB51" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8050A16B833A6D45FEFEFB0AFADDFB11" blockId="4.[264,1322,967,1278]" box="[264,1313,1191,1214]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFB0AFADDFB11" box="[264,1313,1191,1214]" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFB0AFDB4FB11" ID-CoL="8VS4C" authority="Seevers, 1978: 125" authorityName="Seevers" authorityPageNumber="125" authorityYear="1978" box="[264,584,1191,1214]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFB0AFE9CFB11" box="[264,352,1191,1214]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
:
<treatmentCitation id="014E877A833A6D45FE80FB0AFDB4FB11" author="Seevers" box="[374,584,1191,1214]" page="125" pageId="4" pageNumber="5" year="1978">
<bibRefCitation id="E47EDC9A833A6D45FE80FB0AFDF3FB11" author="Seevers" box="[374,527,1191,1214]" pageId="4" pageNumber="43" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
: 125
</treatmentCitation>
</taxonomicName>
(as valid genus in subtribe Xenotae
<bibRefCitation id="E47EDC9A833A6D45FC3BFB0AFB9FFB11" author="Seevers" box="[973,1123,1191,1214]" pageId="4" pageNumber="43" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
(
<emphasis id="B29B7D79833A6D45FB84FB0AFAF5FB11" box="[1138,1289,1191,1214]" italics="true" pageId="4" pageNumber="5">nomen nudum</emphasis>
)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8050A16B833A6D45FEFEFB6AFE0DFB51" blockId="4.[264,1322,967,1278]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="A0FF8645833A6D45FEFEFB6AFE0DFB51" pageId="4" pageNumber="5">
<taxonomicName id="47EFDAE8833A6D45FEFEFB6AFE51FB71" ID-CoL="8VS4C" authority="Ashe" authorityName="Ashe" box="[264,429,1223,1246]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FEFEFB6AFE9CFB71" box="[264,352,1223,1246]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
: Ashe
</taxonomicName>
<emphasis id="B29B7D79833A6D45FE42FB6AFE35FB71" box="[436,457,1223,1246]" italics="true" pageId="4" pageNumber="5">in</emphasis>
<treatmentCitation id="014E877A833A6D45FE39FB6AFC3DFB71" author="Newton" box="[463,961,1223,1246]" page="369" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="E47EDC9A833A6D45FE39FB6AFC74FB71" author="Newton" box="[463,904,1223,1246]" pageId="4" pageNumber="43" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter" year="2000">Newton, Thayer, Ashe &amp; Chandler, 2000</bibRefCitation>
: 369
</treatmentCitation>
(as valid genus in subtribe
<taxonomicName id="47EFDAE8833A6D45FB10FB6AFE19FB51" ID-CoL="8VRC4" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">
Athetina
<bibRefCitation id="E47EDC9A833A6D45FEA9FB4AFE19FB51" author="Casey" box="[351,485,1255,1278]" pageId="4" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E0833A6D44FEFEFA8CFE76F8C2" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8050A16B833A6D45FEFEFA8CFE76F8C4" blockId="4.[264,1323,1313,1899]" pageId="4" pageNumber="5">
<emphasis id="B29B7D79833A6D45FEFEFA8CFE74FA94" bold="true" box="[264,392,1313,1339]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
<taxonomicName id="47EFDAE8833A6D45FE61FA8FFE06FA94" box="[407,506,1314,1339]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833A6D45FE61FA8FFE06FA94" box="[407,506,1314,1339]" italics="true" pageId="4" pageNumber="5">Stethusa</emphasis>
</taxonomicName>
is distinguished from other athetine genera by the combination of the following characters: body parallel­sided; anterior margin of labrum concave; sensilla
<emphasis id="B29B7D79833A6D45FB0BFAE7FAF0FACC" box="[1277,1292,1354,1379]" italics="true" pageId="4" pageNumber="5">a</emphasis>
of epipharynx long; antennal article 2 slightly longer than article 3, articles 4­10 slightly elongate or subquadrate; ligula with narrow base and split apically (
<figureCitation id="18D4BDEE833A6D45FBC4FA34FB86FA1C" box="[1074,1146,1433,1459]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
); labial palpus with setae, and present (
<figureCitation id="18D4BDEE833A6D45FD82FA6CFD40FA74" box="[628,700,1473,1499]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
); pronotum slightly transverse, 1.2 times as wide as long, with microsetae directed anteriorly along the midline; in lateral portions of the disc microsetae directed laterally (
<typeStatus id="5F541FC9833A6D45FD91F9BCFD5FF984" box="[615,675,1553,1579]" pageId="4" pageNumber="5">Type</typeStatus>
I,
<bibRefCitation id="E47EDC9A833A6D45FD33F9BCFC30F984" author="Benick" box="[709,972,1553,1579]" pageId="4" pageNumber="42" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="18D4BDEE833A6D45FC13F9BCFBC2F984" box="[997,1086,1553,1579]" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
); pronotal macrosetae long; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia thick and long, twice as long as tibial width (
<figureCitation id="18D4BDEE833A6D45FCDAF9CCFC79F9D4" box="[812,901,1633,1659]" captionStart="FIGURES 14 ­ 17" captionStartId="9.[264,382,1243,1267]" captionTargetBox="[347,1240,317,1218]" captionTargetId="figure@9.[337,1249,307,1227]" captionTargetPageId="9" captionText="FIGURES 14 ­ 17. Details of Stethusa dichroa (Gravenhorst) (male (14 ­ 16) and female (17), Eddyville, Illinois). 14 – right mesotibia; 15 – antennal article 11; 16 – right antenna; 17 – right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16)." httpUri="https://zenodo.org/record/156497/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
); mesothoracic process broad (
<figureCitation id="18D4BDEE833A6D45FB0DF9CCFEACF90C" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="4" pageNumber="5">Fig. 19­21</figureCitation>
); tarsal formula 4­5­5; metatarsal segment 1 as long as segment 2; one long empodial seta; abdominal terga 3­5 with transverse basal impression; medial lamellae of internal sac absent; copulatory piece with pointed apex (
<figureCitation id="18D4BDEE833A6D45FCCFF974FC3AF95C" box="[825,966,1753,1779]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="4" pageNumber="5">Figs. 34, 36</figureCitation>
,
<figureCitation id="18D4BDEE833A6D45FC24F974FC0DF95C" box="[978,1009,1753,1779]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="4" pageNumber="5">56</figureCitation>
) and sclerotized suspensoria (
<figureCitation id="18D4BDEE833A6D45FECFF8ACFE62F8B4" box="[313,414,1793,1819]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="4" pageNumber="5">Figs. 36</figureCitation>
,
<figureCitation id="18D4BDEE833A6D45FE5BF8ACFE32F8B4" box="[429,462,1793,1819]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="4" pageNumber="5">56</figureCitation>
); medial lamellae absent; internal sac with two sclerotized bands (
<figureCitation id="18D4BDEE833A6D45FB06F8ACFEAAF8EC" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="4" pageNumber="5">Figs. 34, 38</figureCitation>
,
<figureCitation id="18D4BDEE833A6D45FE92F884FE7FF8EC" box="[356,387,1833,1859]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="4" pageNumber="5">54</figureCitation>
) which change their orientation when the sac everts; spermatheca short (
<figureCitation id="18D4BDEE833A6D45FB19F884FEB3F8C4" captionStart="FIGURES 40 ­ 42" captionStartId="19.[264,382,1234,1258]" captionTargetBox="[344,1251,575,1191]" captionTargetId="figure@19.[335,1251,566,1207]" captionTargetPageId="19" captionText="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 – female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 – spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." httpUri="https://zenodo.org/record/156502/files/figure.png" pageId="4" pageNumber="5">Figs. 41­42</figureCitation>
,
<figureCitation id="18D4BDEE833A6D45FEADF8FCFE80F8C4" box="[347,380,1873,1899]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="4" pageNumber="5">57</figureCitation>
).
</paragraph>
<caption id="D490F1E3833B6D44FEFEF9ADFE9DF9D0" httpUri="https://zenodo.org/record/156494/files/figure.png" pageId="5" pageNumber="6" targetBox="[291,1293,316,1520]" targetPageId="5">
<paragraph id="8050A16B833B6D44FEFEF9ADFE9DF9D0" blockId="5.[264,1323,1536,1663]" pageId="5" pageNumber="6">
<emphasis id="B29B7D79833B6D44FEFEF9ADFE48F9B7" bold="true" box="[264,436,1536,1560]" pageId="5" pageNumber="6">FIGURES 1­5.</emphasis>
Mouthparts of
<taxonomicName id="47EFDAE8833B6D44FD85F9ADFC37F9B8" authority="Gravenhorst" authorityName="Gravenhorst" box="[627,971,1536,1559]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D79833B6D44FD85F9ADFCD6F9B8" box="[627,810,1536,1559]" italics="true" pageId="5" pageNumber="6">Stethusa dichroa</emphasis>
(Gravenhorst)
</taxonomicName>
(female (1, 5) and male (3­4), Eddyville, Illinois; female, 18 km SSE Lawrence, Kansas (2)). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8050A16B833B6D44FECEF976FE76F8C2" blockId="5.[264,1322,1755,1901]" pageId="5" pageNumber="6">
<taxonomicName id="47EFDAE8833B6D44FECEF976FE66F95B" box="[312,410,1755,1780]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833B6D44FECEF976FE66F95B" box="[312,410,1755,1780]" italics="true" pageId="5" pageNumber="6">Stethusa</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="47EFDAE8833B6D44FD32F976FCECF95B" box="[708,784,1755,1780]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D79833B6D44FD32F976FCECF95B" box="[708,784,1755,1780]" italics="true" pageId="5" pageNumber="6">Atheta</emphasis>
</taxonomicName>
(including
<emphasis id="B29B7D79833B6D44FC5CF976FB90F95B" box="[938,1132,1755,1780]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="47EFDAE8833B6D44FC5CF976FB94F95B" box="[938,1128,1755,1780]" class="Insecta" family="Staphylinidae" genus="Dimetrota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="auct">Dimetrota auct</taxonomicName>
.
</emphasis>
) by a broader mesothoracic process (
<figureCitation id="18D4BDEE833B6D44FDE4F8AEFD60F8B2" box="[530,668,1795,1821]" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="5" pageNumber="6">Figs. 19­21</figureCitation>
); the lack of the median lamellae of the internal sac; the distinct shape of the copulatory piece (
<figureCitation id="18D4BDEE833B6D44FD3AF886FCA6F8EA" box="[716,858,1835,1861]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="5" pageNumber="6">Figs. 36, 39</figureCitation>
,
<figureCitation id="18D4BDEE833B6D44FC90F886FC52F8EA" box="[870,942,1835,1861]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="5" pageNumber="6">55­56</figureCitation>
); and a short spermatheca (
<figureCitation id="18D4BDEE833B6D44FB1BF886FEB3F8C2" captionStart="FIGURES 40 ­ 42" captionStartId="19.[264,382,1234,1258]" captionTargetBox="[344,1251,575,1191]" captionTargetId="figure@19.[335,1251,566,1207]" captionTargetPageId="19" captionText="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 – female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 – spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." httpUri="https://zenodo.org/record/156502/files/figure.png" pageId="5" pageNumber="6">Figs. 41­42</figureCitation>
,
<figureCitation id="18D4BDEE833B6D44FEADF8FEFE80F8C2" box="[347,380,1875,1901]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="5" pageNumber="6">57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E083386D47FECEFEB4FDFCFE04" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8050A16B83386D47FECEFEB4FDFCFE04" blockId="6.[264,1323,281,427]" pageId="6" pageNumber="7">
<taxonomicName id="47EFDAE883386D47FECEFEB4FE66FE9D" ID-CoL="8VS4C" box="[312,410,281,306]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983386D47FECEFEB4FE66FE9D" box="[312,410,281,306]" italics="true" pageId="6" pageNumber="7">Stethusa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="47EFDAE883386D47FDCEFEB4FC6CFE9C" ID-CoL="8VRN4" authority="Mulsant &amp; Rey, 1874" authorityName="Mulsant &amp; Rey" authorityYear="1874" box="[568,912,281,307]" class="Insecta" family="Staphylinidae" genus="Earota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983386D47FDCEFEB4FD74FE9D" box="[568,648,281,306]" italics="true" pageId="6" pageNumber="7">Earota</emphasis>
Mulsant &amp;
<bibRefCitation id="E47EDC9A83386D47FCEEFEB4FC6CFE9C" author="Rey" box="[792,912,281,307]" pageId="6" pageNumber="43" refString="Muslant, E. &amp; Rey, C. (1874) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts utiles de Lyon, serie 4. 6 [1873], 33 - 738." type="journal article" year="1874">
Rey,
<quantity id="47170C8E83386D47FCA3FEB4FC6CFE9C" box="[853,912,281,307]" metricMagnitude="1" metricUnit="m" metricValue="4.7599599999999995" pageId="6" pageNumber="7" unit="in" value="1874.0">1874</quantity>
</bibRefCitation>
</taxonomicName>
in having sensilla
<emphasis id="B29B7D7983386D47FB99FEB4FB82FE9D" box="[1135,1150,281,306]" italics="true" pageId="6" pageNumber="7">a</emphasis>
of the epipharynx long; pronotal setation of
<typeStatus id="5F541FC983386D47FD81FEECFD56FEF4" box="[631,682,321,347]" pageId="6" pageNumber="7">type</typeStatus>
I; slightly narrower mesothoracic process; by the lack of the median lamellae of the internal sac; and the distinct shape of the copulatory piece (
<figureCitation id="18D4BDEE83386D47FEE7FE3CFE63FE04" box="[273,415,401,427]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="6" pageNumber="7">Figs. 36, 39</figureCitation>
,
<figureCitation id="18D4BDEE83386D47FE5DFE3CFE08FE04" box="[427,500,401,427]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="6" pageNumber="7">55­56</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D490F1E383386D47FEFEFABDFC7BFAE6" httpUri="https://zenodo.org/record/156495/files/figure.png" pageId="6" pageNumber="7" targetBox="[379,1208,514,1282]" targetPageId="6">
<paragraph id="8050A16B83386D47FEFEFABDFC7BFAE6" blockId="6.[264,1322,1296,1353]" pageId="6" pageNumber="7">
<emphasis id="B29B7D7983386D47FEFEFABDFE72FA87" bold="true" box="[264,398,1296,1320]" pageId="6" pageNumber="7">FIGURE 6.</emphasis>
Nomenclature of setae and pores in epipharynx of
<taxonomicName id="47EFDAE883386D47FC20FABDFAD6FA88" authority="Gravenhorst" authorityName="Gravenhorst" box="[982,1322,1296,1319]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983386D47FC20FABDFB77FA88" box="[982,1163,1296,1319]" italics="true" pageId="6" pageNumber="7">Stethusa dichroa</emphasis>
(Gravenhorst)
</taxonomicName>
(female, 18 km SSE Lawrence, Kansas). Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C8F5F2E083386D46FECEFA08FB9FF9A4" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="8050A16B83386D47FECEFA08FB9FFA48" blockId="6.[264,1323,1445,1871]" pageId="6" pageNumber="7">
<emphasis id="B29B7D7983386D47FECEFA08FE2CFA10" bold="true" box="[312,464,1445,1471]" pageId="6" pageNumber="7">Description.</emphasis>
Length 2.0­
<quantity id="47170C8E83386D47FD91FA08FD38FA10" box="[615,708,1445,1471]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="3.5">3.5 mm</quantity>
, pronotal width
<quantity id="47170C8E83386D47FC74FA08FBD7FA10" box="[898,1067,1445,1471]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="7.7" metricValueMin="4.6" pageId="6" pageNumber="7" unit="mm" value="0.615" valueMax="0.77" valueMin="0.46">0.46­0.77 mm</quantity>
. Body parallel­sided. Body brown to dark brown, with brownish yellow elytra and yellow legs.
</paragraph>
<paragraph id="8050A16B83386D46FECEFA58FB92FCBC" blockId="6.[264,1323,1445,1871]" lastBlockId="7.[264,1323,281,1867]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Head transverse; eyes very large, eye length to temple length ratio 2.5­5; infraorbital carina complete. Antennal article 2 slightly longer than article 3, articles 4­10 slightly elongate, subquadrate or transverse, apical article without coeloconic sensilla (not to be mixed with subbasal impression densely covered with microsetae, present in
<taxonomicName id="47EFDAE883386D47FB46F9C0FAD6F929" box="[1200,1322,1645,1670]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983386D47FB46F9C0FAD6F929" box="[1200,1322,1645,1670]" italics="true" pageId="6" pageNumber="7">S. dichroa</emphasis>
</taxonomicName>
and
<taxonomicName id="47EFDAE883386D47FECDF938FE11F901" box="[315,493,1685,1710]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B29B7D7983386D47FECDF938FE11F901" box="[315,493,1685,1710]" italics="true" pageId="6" pageNumber="7">S. luederwaldti</emphasis>
</taxonomicName>
), as long as or longer than articles 9 and 10 combined. Labrum (
<figureCitation id="18D4BDEE83386D47FB0DF938FEE9F978" captionStart="FIGURES 1 ­ 5" captionStartId="5.[264,382,1536,1560]" captionTargetBox="[291,1293,316,1520]" captionTargetId="figure@5.[281,1305,304,1529]" captionTargetPageId="5" captionText="FIGURES 1 ­ 5. Mouthparts of Stethusa dichroa (Gravenhorst) (female (1, 5) and male (3 ­ 4), Eddyville, Illinois; female, 18 km SSE Lawrence, Kansas (2 )). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156494/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
) transverse, with concave anterior margin. Epipharynx (
<figureCitation id="18D4BDEE83386D47FC4CF910FBF1F978" box="[954,1037,1725,1751]" captionStart="FIGURES 1 ­ 5" captionStartId="5.[264,382,1536,1560]" captionTargetBox="[291,1293,316,1520]" captionTargetId="figure@5.[281,1305,304,1529]" captionTargetPageId="5" captionText="FIGURES 1 ­ 5. Mouthparts of Stethusa dichroa (Gravenhorst) (female (1, 5) and male (3 ­ 4), Eddyville, Illinois; female, 18 km SSE Lawrence, Kansas (2 )). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156494/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
,
<figureCitation id="18D4BDEE83386D47FBEDF910FBD6F978" box="[1051,1066,1725,1751]" captionStart="FIGURE 6" captionStartId="6.[264,368,1296,1320]" captionTargetBox="[379,1208,514,1282]" captionTargetId="figure@6.[379,1209,511,1287]" captionTargetPageId="6" captionText="FIGURE 6. Nomenclature of setae and pores in epipharynx of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156495/files/figure.png" pageId="6" pageNumber="7">6</figureCitation>
) with long sensilla
<emphasis id="B29B7D7983386D47FAE2F910FADFF979" box="[1300,1315,1725,1750]" italics="true" pageId="6" pageNumber="7">a</emphasis>
, with three pairs of small marginal setae, medial field with 36­38 pores, lateral rows with three pores each, anterolateral groups with three to four pores each, transverse row with six pores, posterolateral groups with four pores each, with one medial proximal pore on each side and no lateral proximal pores. Mandibles (
<figureCitation id="18D4BDEE83396D46FC81FEB4FC1CFE9C" box="[887,992,281,307]" captionStart="FIGURES 1 ­ 5" captionStartId="5.[264,382,1536,1560]" captionTargetBox="[291,1293,316,1520]" captionTargetId="figure@5.[281,1305,304,1529]" captionTargetPageId="5" captionText="FIGURES 1 ­ 5. Mouthparts of Stethusa dichroa (Gravenhorst) (female (1, 5) and male (3 ­ 4), Eddyville, Illinois; female, 18 km SSE Lawrence, Kansas (2 )). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156494/files/figure.png" pageId="7" pageNumber="8">Figs. 3­5</figureCitation>
) broad, right mandible with a small medial tooth; dorsal molar area with velvety patch consisting of very small denticles (poorly visible at 400x). Maxilla (
<figureCitation id="18D4BDEE83396D46FD1EFEC4FC9DFE2C" box="[744,865,361,387]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Figs. 9­12</figureCitation>
) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae (
<figureCitation id="18D4BDEE83396D46FCF1FE14FCA3FE7C" box="[775,863,441,467]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
); distal comb of lacinia is divided into isolated groups of 6 and 2 closely placed spines (
<figureCitation id="18D4BDEE83396D46FC96FE4CFC16FE54" box="[864,1002,481,507]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Figs. 10­11</figureCitation>
), middle portion produced medially and covered with numerous fine setae (
<figureCitation id="18D4BDEE83396D46FCAFFDA4FC19FD8C" box="[857,997,521,547]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Figs. 10­11</figureCitation>
), ventral surface of lacinia with a medial group of 4 strong setae and marginal group of 5 strong setae (
<figureCitation id="18D4BDEE83396D46FB81FD9CFB32FDE4" box="[1143,1230,561,587]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 10</figureCitation>
), dorsal surface of lacinia with a row of 13 strong setae (
<figureCitation id="18D4BDEE83396D46FCC1FDF4FC70FDDC" box="[823,908,601,627]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
). Labium as in
<figureCitation id="18D4BDEE83396D46FBB0FDF4FB27FDDC" box="[1094,1243,601,627]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Figs. 7­8, 13</figureCitation>
; ligula with narrow base and split apically (
<figureCitation id="18D4BDEE83396D46FD4BFD2CFCF9FD34" box="[701,773,641,667]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
); medial area of prementum with 2 pores and with 8­20 pseudopores, lateral areas each with two asetose pores, single setose pore and 8­ 12 pseudopores (
<figureCitation id="18D4BDEE83396D46FE27FD7CFDE5FD44" box="[465,537,721,747]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
). Hypopharyngeal lobes as in
<figureCitation id="18D4BDEE83396D46FC7DFD7CFC28FD44" box="[907,980,721,747]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
. Labial palpus with setae,, and present (
<figureCitation id="18D4BDEE83396D46FE27FD54FDEBFCBC" box="[465,535,761,787]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
). Mentum (
<figureCitation id="18D4BDEE83396D46FD5FFD54FD02FCBC" box="[681,766,761,787]" captionStart="FIGURES 7 ­ 13" captionStartId="8.[264,382,1402,1426]" captionTargetBox="[309,1277,513,1386]" captionTargetId="figure@8.[302,1286,504,1395]" captionTargetPageId="8" captionText="FIGURES 7 ­ 13. Mouthparts of Stethusa dichroa (Gravenhorst) (female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7 ­ 8, 10 ­ 13), 0.2 mm (9)." httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
) with concave anterior margin.
</paragraph>
<paragraph id="8050A16B83396D46FECEFC8CFB54FB64" blockId="7.[264,1323,281,1867]" pageId="7" pageNumber="8">
Pronotum (
<figureCitation id="18D4BDEE83396D46FE36FC8CFDE5FC94" box="[448,537,801,827]" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="7" pageNumber="8">Fig. 18</figureCitation>
) slightly transverse, 1.2 times as wide as long, with microsetae directed anteriorly in midline; in lateral portions of disc microsetae directed laterally (
<typeStatus id="5F541FC983396D46FB19FCE4FAD7FCCC" box="[1263,1323,841,867]" pageId="7" pageNumber="8">Type</typeStatus>
I,
<bibRefCitation id="E47EDC9A83396D46FED6FCDCFDDBFC24" author="Benick" box="[288,551,881,907]" pageId="7" pageNumber="42" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); macrosetae short; hypomera fully visible in lateral view. Meso­ and metasternum as in
<figureCitation id="18D4BDEE83396D46FDE0FC34FD61FC1C" box="[534,669,921,947]" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="7" pageNumber="8">Figs. 19­21</figureCitation>
, mesosternal process moderately wide, extending about ½ length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially or (in
<taxonomicName id="47EFDAE883396D46FD43FC47FCD2FBAC" box="[693,814,1002,1027]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983396D46FD43FC47FCD2FBAC" box="[693,814,1002,1027]" italics="true" pageId="7" pageNumber="8">S. dichroa</emphasis>
</taxonomicName>
) mesosternum with very short carina (
<figureCitation id="18D4BDEE83396D46FB0DFC44FED4FB84" captionStart="FIGURES 18 ­ 21" captionStartId="10.[264,382,1704,1728]" captionTargetBox="[348,1242,678,1683]" captionTargetId="figure@10.[338,1248,670,1691]" captionTargetPageId="10" captionText="FIGURES 18 ­ 21. Details of Stethusa dichroa (Gravenhorst) (male (18) and female (19), Eddyville, Illinois), S. spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (20 )), and S. klimschi (Bernhauer) (male, Opelousas, Louisiana (21 )). 18 – pronotum; 19 ­ 21 – meso­ and metathorax, ventral view. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156498/files/figure.png" pageId="7" pageNumber="8">Fig. 19</figureCitation>
); relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 3:1:2 or 4:1:2; mesocoxal cavities margined posteriorly; mesocoxae moderately widely separated. Medial macroseta of mesotibia long and thick, twice as long as tibial width. Tarsal segmentation 4­5­5, metatarsal segment 1 as long as segment 2. One empodial seta, shorter than claws. Posterior margin of elytra straight. Wings fully developed.
</paragraph>
<paragraph id="8050A16B83396D46FECEFB74FE41FAEC" blockId="7.[264,1323,281,1867]" pageId="7" pageNumber="8">Abdominal terga 3­5 with moderate basal impressions. Tergum 7 is 1.1 times as long as tergum 6. Punctation on terga 6­7 sparser than on terga 3­5. Tergum 7 with wide white palisade fringe.</paragraph>
<paragraph id="8050A16B83396D46FECEFAFCFB9FF9A4" blockId="7.[264,1323,281,1867]" pageId="7" pageNumber="8">
Internal sac of aedeagus without medial lamellae; with two sclerotized bands (which may be homologous to medial lamellae; however, unlike the medial lamellae the distal end of the bands is oriented towards the base of median lobe in retracted internal sac); copulatory piece with pointed apex (
<figureCitation id="18D4BDEE83396D46FD9DFA64FD07FA4C" box="[619,763,1481,1507]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="7" pageNumber="8">Figs. 34, 36</figureCitation>
,
<figureCitation id="18D4BDEE83396D46FCFEFA64FCDBFA4C" box="[776,807,1481,1507]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="7" pageNumber="8">56</figureCitation>
) and sclerotized suspensoria (
<figureCitation id="18D4BDEE83396D46FB7DFA64FB13FA4C" box="[1163,1263,1481,1507]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="7" pageNumber="8">Figs. 36</figureCitation>
,
<figureCitation id="18D4BDEE83396D46FB0AFA64FAE1FA4C" box="[1276,1309,1481,1507]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="7" pageNumber="8">56</figureCitation>
), in lateral view curved (
<figureCitation id="18D4BDEE83396D46FDE1FA5CFD5AF9A4" box="[535,678,1521,1547]" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 – everted internal sac, parameral view; 35 ­ 36 – details of retracted internal sac, abparameral view; 37 – everted internal sac, lateral view; 38 ­ 39 – details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="7" pageNumber="8">Figs. 37, 39</figureCitation>
); spermatheca short (
<figureCitation id="18D4BDEE83396D46FC69FA5CFBD5F9A4" box="[927,1065,1521,1547]" captionStart="FIGURES 40 ­ 42" captionStartId="19.[264,382,1234,1258]" captionTargetBox="[344,1251,575,1191]" captionTargetId="figure@19.[335,1251,566,1207]" captionTargetPageId="19" captionText="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 – female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 – spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." httpUri="https://zenodo.org/record/156502/files/figure.png" pageId="7" pageNumber="8">Figs. 41­42</figureCitation>
,
<figureCitation id="18D4BDEE83396D46FBC3FA5CFBAAF9A4" box="[1077,1110,1521,1547]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 – details of retracted internal sac, abparameral view; 54 – details of retracted internal sac, lateral view; 55 – copulatory piece, lateral view; 56 – copulatory piece, abparameral view; 57 – spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="7" pageNumber="8">57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E083396D46FECEF9B4FADFF99C" box="[312,1315,1561,1587]" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8050A16B83396D46FECEF9B4FADFF99C" blockId="7.[264,1323,281,1867]" box="[312,1315,1561,1587]" pageId="7" pageNumber="8">
<emphasis id="B29B7D7983396D46FECEF9B4FE2CF99C" bold="true" box="[312,464,1561,1587]" pageId="7" pageNumber="8">
<typeStatus id="5F541FC983396D46FECEF9B4FE8FF99C" box="[312,371,1561,1587]" pageId="7" pageNumber="8">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="47EFDAE883396D46FE1EF9B7FCDFF99C" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[488,803,1561,1587]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="irvingi">
<emphasis id="B29B7D7983396D46FE1EF9B7FD76F99C" box="[488,650,1562,1587]" italics="true" pageId="7" pageNumber="8">Atheta irvingi</emphasis>
<bibRefCitation id="E47EDC9A83396D46FD64F9B4FCDFF99C" author="Casey" box="[658,803,1561,1587]" pageId="7" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation id="E47EDC9A83396D46FB82F9B4FAEAF99C" author="Fenyes" box="[1140,1302,1561,1587]" pageId="7" pageNumber="42" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8F5F2E083396D4BFECEF9ECFB72FDE4" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8050A16B83396D49FECEF9ECFE11FE04" blockId="7.[264,1323,281,1867]" lastBlockId="8.[264,1323,281,427]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B29B7D7983396D46FECEF9ECFE3FF9F4" bold="true" box="[312,451,1601,1627]" pageId="7" pageNumber="8">Discussion.</emphasis>
When describing
<taxonomicName id="47EFDAE883396D46FD5DF9EFFC42F9F4" authority="Casey (1910)" authorityName="Casey" authorityYear="1910" box="[683,958,1601,1627]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983396D46FD5DF9EFFCF1F9F4" box="[683,781,1602,1627]" italics="true" pageId="7" pageNumber="8">Stethusa</emphasis>
,
<bibRefCitation id="E47EDC9A83396D46FCE8F9ECFC42F9F4" author="Casey" box="[798,958,1601,1627]" pageId="7" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey (1910)</bibRefCitation>
</taxonomicName>
did not explicitly fix the
<typeStatus id="5F541FC983396D46FB0EF9ECFAD7F9F4" box="[1272,1323,1601,1627]" pageId="7" pageNumber="8">type</typeStatus>
species of the subgenus. While designating the
<typeStatus id="5F541FC983396D46FCBFF9C4FC80F92C" box="[841,892,1641,1667]" pageId="7" pageNumber="8">type</typeStatus>
species of
<taxonomicName id="47EFDAE883396D46FBF2F9C7FAF7F92C" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[1028,1291,1641,1667]" class="Insecta" family="Staphylinidae" genus="Noverota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983396D46FBF2F9C7FB8CF92C" box="[1028,1136,1642,1667]" italics="true" pageId="7" pageNumber="8">Noverota</emphasis>
<bibRefCitation id="E47EDC9A83396D46FB8CF9C4FAF7F92C" author="Casey" box="[1146,1291,1641,1667]" pageId="7" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">
Casey,
<quantity id="47170C8E83396D46FB26F9C4FAF7F92C" box="[1232,1291,1641,1667]" metricMagnitude="1" metricUnit="m" metricValue="4.8514" pageId="7" pageNumber="8" unit="in" value="1910.0">1910</quantity>
</bibRefCitation>
</taxonomicName>
in the same paper, Casey stated (1910, p. 90): “The first species may be regarded as the
<typeStatus id="5F541FC983396D46FB06F93CFAD9F904" box="[1264,1317,1681,1707]" pageId="7" pageNumber="8">type</typeStatus>
, as in all other cases where the
<typeStatus id="5F541FC983396D46FD9CF914FD60F97C" box="[618,668,1721,1747]" pageId="7" pageNumber="8">type</typeStatus>
is not specifically named”. Unfortunately it is not clear how to apply this recommendation to
<taxonomicName id="47EFDAE883396D46FD22F94FFCC9F954" box="[724,821,1762,1787]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983396D46FD22F94FFCC9F954" box="[724,821,1762,1787]" italics="true" pageId="7" pageNumber="8">Stethusa</emphasis>
</taxonomicName>
. The first species described by Casey in
<taxonomicName id="47EFDAE883396D46FEFEF8A7FE95F88C" box="[264,361,1802,1827]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983396D46FEFEF8A7FE95F88C" box="[264,361,1802,1827]" italics="true" pageId="7" pageNumber="8">Stethusa</emphasis>
</taxonomicName>
is
<taxonomicName id="47EFDAE883396D46FE64F8A7FD44F88C" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[402,696,1801,1827]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="affluens">
<emphasis id="B29B7D7983396D46FE64F8A7FDE6F88C" box="[402,538,1802,1827]" italics="true" pageId="7" pageNumber="8">At. affluens</emphasis>
<bibRefCitation id="E47EDC9A83396D46FDD5F8A4FD44F88C" author="Casey" box="[547,696,1801,1827]" pageId="7" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
. However, before Casey proceeds to describing the new species he mentions that
<taxonomicName id="47EFDAE883396D46FD8DF89FFD21F8E4" box="[635,733,1842,1867]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983396D46FD8DF89FFD21F8E4" box="[635,733,1842,1867]" italics="true" pageId="7" pageNumber="8">Stethusa</emphasis>
</taxonomicName>
includes many allies of
<taxonomicName id="47EFDAE883396D49FBE6F89FFEBAFE9C" authority="Bernhauer, 1909" authorityName="Bernhauer" authorityYear="1909" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" lastPageId="8" lastPageNumber="9" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983396D46FBE6F89FFB63F8E4" box="[1040,1183,1842,1867]" italics="true" pageId="7" pageNumber="8">At. klimschi</emphasis>
<bibRefCitation id="E47EDC9A83396D49FB5CF89CFEBAFE9C" author="Bernhauer" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="42" refString="Bernhauer, M. (1909) Neue Aleocharini aus Nordamerika. (Col.) (4. Stuck.). Deutsche Entomologische Zeitschrift, 1909 (4), 515 - 528." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
</taxonomicName>
; therefore,
<taxonomicName id="47EFDAE883366D49FE3CFEB4FDA9FE9D" box="[458,597,281,306]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983366D49FE3CFEB4FDA9FE9D" box="[458,597,281,306]" italics="true" pageId="8" pageNumber="9">At. klimschi</emphasis>
</taxonomicName>
was also a species originally included in
<taxonomicName id="47EFDAE883366D49FBCCFEB4FB60FE9D" box="[1082,1180,281,306]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983366D49FBCCFEB4FB60FE9D" box="[1082,1180,281,306]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
</taxonomicName>
. Because it is not clear which of the two species,
<taxonomicName id="47EFDAE883366D49FD33FEECFCACFEF5" box="[709,848,321,346]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983366D49FD33FEECFCACFEF5" box="[709,848,321,346]" italics="true" pageId="8" pageNumber="9">At. klimschi</emphasis>
</taxonomicName>
or
<taxonomicName id="47EFDAE883366D49FC8EFEECFC03FEF5" box="[888,1023,321,346]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="affluens">
<emphasis id="B29B7D7983366D49FC8EFEECFC03FEF5" box="[888,1023,321,346]" italics="true" pageId="8" pageNumber="9">At. affluens</emphasis>
</taxonomicName>
qualifies as the
<typeStatus id="5F541FC983366D49FB49FEECFB0DFEF4" box="[1215,1265,321,347]" pageId="8" pageNumber="9">type</typeStatus>
species under Casey's first species rule, I maintain that Casey has not validly fixed the
<typeStatus id="5F541FC983366D49FB0EFEC4FAD7FE2C" box="[1272,1323,361,387]" pageId="8" pageNumber="9">type</typeStatus>
species of
<taxonomicName id="47EFDAE883366D49FE72FE3CFE1AFE05" box="[388,486,401,426]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983366D49FE72FE3CFE1AFE05" box="[388,486,401,426]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="D490F1E383366D49FEFEFAD7FDE8FA55" httpUri="https://zenodo.org/record/156496/files/figure.png" pageId="8" pageNumber="9" targetBox="[309,1277,513,1386]" targetPageId="8">
<paragraph id="8050A16B83366D49FEFEFAD7FDE8FA55" blockId="8.[264,1323,1402,1530]" pageId="8" pageNumber="9">
<emphasis id="B29B7D7983366D49FEFEFAD7FE43FA3D" bold="true" box="[264,447,1402,1426]" pageId="8" pageNumber="9">FIGURES 7­13.</emphasis>
Mouthparts of
<taxonomicName id="47EFDAE883366D49FD80FAD6FC35FA3D" authority="Gravenhorst" authorityName="Gravenhorst" box="[630,969,1403,1426]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983366D49FD80FAD6FD2CFA3D" box="[630,720,1403,1426]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
<emphasis id="B29B7D7983366D49FD2FFAD6FCD7FA3D" box="[729,811,1403,1426]" italics="true" pageId="8" pageNumber="9">dichroa</emphasis>
(Gravenhorst)
</taxonomicName>
(female, 18 km SSE Lawrence, Kansas). 7 – hypopharynx; 8 – prementum; 9 – right maxilla, ventral view; 10 – right galea, ventral view; 11 – right galea, dorsal view; 12 – right lacinia, dorsal view; 13 – mentum. Scale bar 0.1 mm (7­8, 10­13), 0.2 mm (9).
</paragraph>
</caption>
<paragraph id="8050A16B83366D49FECEF9FBFDF6F8CF" blockId="8.[264,1323,1622,1888]" pageId="8" pageNumber="9">
<bibRefCitation id="E47EDC9A83366D49FECEF9FBFE1EF9DF" author="Fenyes" box="[312,482,1622,1648]" pageId="8" pageNumber="42" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes (1920)</bibRefCitation>
listed the subgenus
<taxonomicName id="47EFDAE883366D49FD23F9FBFCCAF9C0" box="[725,822,1622,1647]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983366D49FD23F9FBFCCAF9C0" box="[725,822,1622,1647]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
</taxonomicName>
as a synonym of the subgenus
<emphasis id="B29B7D7983366D49FB46F9FBFAD6F9C0" box="[1200,1322,1622,1647]" italics="true" pageId="8" pageNumber="9">Hypatheta</emphasis>
<bibRefCitation id="E47EDC9A83366D49FEFEF9D3FE58F937" author="Fenyes" box="[264,420,1662,1688]" pageId="8" pageNumber="42" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
and included the
<typeStatus id="5F541FC983366D49FD80F9D3FD54F937" box="[630,680,1662,1688]" pageId="8" pageNumber="9">type</typeStatus>
species of
<taxonomicName id="47EFDAE883366D49FCDBF9D3FC73F938" box="[813,911,1662,1687]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983366D49FCDBF9D3FC73F938" box="[813,911,1662,1687]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
</taxonomicName>
in
<emphasis id="B29B7D7983366D49FC43F9D3FBD3F938" box="[949,1071,1662,1687]" italics="true" pageId="8" pageNumber="9">Hypatheta</emphasis>
. The
<typeStatus id="5F541FC983366D49FB8AF9D3FB53F937" box="[1148,1199,1662,1688]" pageId="8" pageNumber="9">type</typeStatus>
species of
<emphasis id="B29B7D7983366D49FEFEF90BFE7EF910" box="[264,386,1702,1727]" italics="true" pageId="8" pageNumber="9">Hypatheta</emphasis>
is
<taxonomicName id="47EFDAE883366D49FE47F90BFC5DF96F" authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" box="[433,929,1702,1728]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="castanoptera">
<emphasis id="B29B7D7983366D49FE47F90BFD63F910" box="[433,671,1702,1727]" italics="true" pageId="8" pageNumber="9">Atheta castanoptera</emphasis>
(
<bibRefCitation id="E47EDC9A83366D49FD47F90BFC6BF96F" author="Mannerheim" box="[689,919,1702,1728]" pageId="8" pageNumber="43" refString="Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des Brachelytres, de l'ordre des Insectes Coleopteres. St. Petersbourg. 87 pp." type="book" year="1830">Mannerheim, 1830</bibRefCitation>
)
</taxonomicName>
, currently placed in
<emphasis id="B29B7D7983366D49FB60F90BFE04F948" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="47EFDAE883366D49FB60F90BFB1FF910" box="[1174,1251,1702,1727]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">Atheta</taxonomicName>
<taxonomicNameLabel id="A9A8C00283366D49FB1AF90BFAD7F910" box="[1260,1323,1702,1727]" pageId="8" pageNumber="9" sensu="stricto">s. str.</taxonomicNameLabel>
<taxonomicName id="47EFDAE883366D49FEFEF963FE04F948" box="[264,504,1742,1767]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="castanoptera">Atheta castanoptera</taxonomicName>
</emphasis>
and other related species of the subgenus
<emphasis id="B29B7D7983366D49FBF5F963FB67F948" box="[1027,1179,1742,1767]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="47EFDAE883366D49FBF5F963FBACF948" box="[1027,1104,1742,1767]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">Atheta</taxonomicName>
<taxonomicNameLabel id="A9A8C00283366D49FBACF963FB67F948" box="[1114,1179,1742,1767]" pageId="8" pageNumber="9" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
differ from
<taxonomicName id="47EFDAE883366D49FEFEF95BFE95F8A0" box="[264,361,1782,1807]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983366D49FEFEF95BFE95F8A0" box="[264,361,1782,1807]" italics="true" pageId="8" pageNumber="9">Stethusa</emphasis>
</taxonomicName>
in having a narrow mesosternal process, a long spermatheca of distinct shape with a thick distal portion and a thin proximal portion and by the presence of medial lamellae of the internal sac.
</paragraph>
<caption id="D490F1E383376D48FEFEFB76FC32FA97" httpUri="https://zenodo.org/record/156497/files/figure.png" pageId="9" pageNumber="10" targetBox="[347,1240,317,1218]" targetPageId="9">
<paragraph id="8050A16B83376D48FEFEFB76FC32FA97" blockId="9.[264,1323,1243,1336]" pageId="9" pageNumber="10">
<emphasis id="B29B7D7983376D48FEFEFB76FE2CFB5C" bold="true" box="[264,464,1243,1267]" pageId="9" pageNumber="10">FIGURES 14­17.</emphasis>
Details of
<taxonomicName id="47EFDAE883376D48FD92FB71FC41FB5D" authority="Gravenhorst" authorityName="Gravenhorst" box="[612,957,1243,1267]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983376D48FD92FB71FD42FB5C" box="[612,702,1244,1267]" italics="true" pageId="9" pageNumber="10">Stethusa</emphasis>
<emphasis id="B29B7D7983376D48FD3CFB71FCE0FB5C" box="[714,796,1244,1267]" italics="true" pageId="9" pageNumber="10">dichroa</emphasis>
(Gravenhorst)
</taxonomicName>
(male (14­16) and female (17), Eddyville, Illinois). 14 – right mesotibia; 15 – antennal article 11; 16 – right antenna; 17 – right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16).
</paragraph>
</caption>
<paragraph id="8050A16B83376D48FECEFA39FD34F9E1" blockId="9.[264,1324,1428,1894]" pageId="9" pageNumber="10">
<bibRefCitation id="E47EDC9A83376D48FECEFA39FE16FA01" author="Seevers" box="[312,490,1428,1454]" pageId="9" pageNumber="43" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers (1978)</bibRefCitation>
raised
<taxonomicName id="47EFDAE883376D48FDB6FA39FD5EFA02" box="[576,674,1428,1453]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983376D48FDB6FA39FD5EFA02" box="[576,674,1428,1453]" italics="true" pageId="9" pageNumber="10">Stethusa</emphasis>
</taxonomicName>
to generic rank. The only character listed in the diagnosis of the genus (p. 125) is the oval pale areas near the base of the terminal antennal segment. Seevers noted that
<taxonomicName id="47EFDAE883376D48FDC1FA49FC6FFA51" authority="Bernhauer, 1909" authorityName="Bernhauer" authorityYear="1909" box="[567,915,1508,1534]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983376D48FDC1FA49FD4BFA52" box="[567,695,1508,1533]" italics="true" pageId="9" pageNumber="10">S. klimschi</emphasis>
(
<bibRefCitation id="E47EDC9A83376D48FD30FA49FC77FA51" author="Bernhauer" box="[710,907,1508,1534]" pageId="9" pageNumber="42" refString="Bernhauer, M. (1909) Neue Aleocharini aus Nordamerika. (Col.) (4. Stuck.). Deutsche Entomologische Zeitschrift, 1909 (4), 515 - 528." type="journal article" year="1909">Bernhauer, 1909</bibRefCitation>
)
</taxonomicName>
lacks this feature but retained it in
<taxonomicName id="47EFDAE883376D48FEFEF9A1FE95F98A" box="[264,361,1548,1573]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983376D48FEFEF9A1FE95F98A" box="[264,361,1548,1573]" italics="true" pageId="9" pageNumber="10">Stethusa</emphasis>
</taxonomicName>
. In the key to genera (p. 51) another diagnostic character of
<taxonomicName id="47EFDAE883376D48FB95F9A1FB38F98A" box="[1123,1220,1548,1573]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983376D48FB95F9A1FB38F98A" box="[1123,1220,1548,1573]" italics="true" pageId="9" pageNumber="10">Stethusa</emphasis>
</taxonomicName>
is mentioned, the broad mesosternal process.
</paragraph>
<paragraph id="8050A16B83376D48FECEF9F1FDEDF941" blockId="9.[264,1324,1428,1894]" pageId="9" pageNumber="10">
<bibRefCitation id="E47EDC9A83376D48FECEF9F1FD8AF9D9" box="[312,630,1628,1654]" pageId="9" pageNumber="43" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter">
Ashe (Newton
<emphasis id="B29B7D7983376D48FE1AF9F1FDD5F9DA" box="[492,553,1628,1653]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2000)
</bibRefCitation>
followed
<bibRefCitation id="E47EDC9A83376D48FD06F9F1FC5EF9D9" author="Seevers" box="[752,930,1628,1654]" pageId="9" pageNumber="43" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers (1978)</bibRefCitation>
and mentioned that thirteen species belong to
<taxonomicName id="47EFDAE883376D48FE42F929FDEAF932" box="[436,534,1668,1693]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983376D48FE42F929FDEAF932" box="[436,534,1668,1693]" italics="true" pageId="9" pageNumber="10">Stethusa</emphasis>
</taxonomicName>
. In the key to genera (pp. 317, 319) &quot;a subbasal &quot;spongy&quot; sensory patch on each side&quot; of the terminal antennal segment is the only mentioned diagnostic character of the genus.
</paragraph>
<paragraph id="8050A16B83376D4BFECEF951FE8EFE54" blockId="9.[264,1324,1428,1894]" lastBlockId="10.[264,1323,281,587]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
My examination of the antennae of
<taxonomicName id="47EFDAE883376D48FD1CF951FC99F8BA" box="[746,869,1788,1813]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983376D48FD1CF951FC99F8BA" box="[746,869,1788,1813]" italics="true" pageId="9" pageNumber="10">S. dichroa</emphasis>
</taxonomicName>
at high magnification (400x) demonstrated that in this species the terminal antennal article has two subbasal impressions (
<figureCitation id="18D4BDEE83376D48FB0DF889FED5F8C9" captionStart="FIGURES 14 ­ 17" captionStartId="9.[264,382,1243,1267]" captionTargetBox="[347,1240,317,1218]" captionTargetId="figure@9.[337,1249,307,1227]" captionTargetPageId="9" captionText="FIGURES 14 ­ 17. Details of Stethusa dichroa (Gravenhorst) (male (14 ­ 16) and female (17), Eddyville, Illinois). 14 – right mesotibia; 15 – antennal article 11; 16 – right antenna; 17 – right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16)." httpUri="https://zenodo.org/record/156497/files/figure.png" pageId="9" pageNumber="10">Fig. 16</figureCitation>
). The microsetae in these impressions are denser and wider, somewhat scale­like, compared to microsetae covering the rest of the article (
<figureCitation id="18D4BDEE83346D4BFC79FEB4FC17FE9C" box="[911,1003,281,307]" captionStart="FIGURES 14 ­ 17" captionStartId="9.[264,382,1243,1267]" captionTargetBox="[347,1240,317,1218]" captionTargetId="figure@9.[337,1249,307,1227]" captionTargetPageId="9" captionText="FIGURES 14 ­ 17. Details of Stethusa dichroa (Gravenhorst) (male (14 ­ 16) and female (17), Eddyville, Illinois). 14 – right mesotibia; 15 – antennal article 11; 16 – right antenna; 17 – right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16)." httpUri="https://zenodo.org/record/156497/files/figure.png" pageId="10" pageNumber="11">Fig. 15</figureCitation>
). All species of
<taxonomicName id="47EFDAE883346D4BFB3EFEB4FAD5FE9D" box="[1224,1321,281,306]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983346D4BFB3EFEB4FAD5FE9D" box="[1224,1321,281,306]" italics="true" pageId="10" pageNumber="11">Stethusa</emphasis>
</taxonomicName>
described by
<bibRefCitation id="E47EDC9A83346D4BFE55FEECFDC4FEF4" author="Casey" box="[419,568,321,347]" pageId="10" pageNumber="42" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, 183 pp." type="book" year="1910">Casey (1910</bibRefCitation>
,
<bibRefCitation id="E47EDC9A83346D4BFDB2FEECFD77FEF4" author="Casey" box="[580,651,321,347]" pageId="10" pageNumber="42" refString="Casey, T. L. (1911 a) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, 245 pp." type="book" year="1911" yearSuffix="a">1911a</bibRefCitation>
) that possessed this modification of the terminal antennal article turned out to be synonyms of
<taxonomicName id="47EFDAE883346D4BFD43FEC4FCD0FE2D" box="[693,812,361,386]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983346D4BFD43FEC4FCD0FE2D" box="[693,812,361,386]" italics="true" pageId="10" pageNumber="11">S. dichroa</emphasis>
</taxonomicName>
(see below). Two other valid Nearctic species of
<taxonomicName id="47EFDAE883346D4BFEAAFE3CFE42FE05" box="[348,446,401,426]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983346D4BFEAAFE3CFE42FE05" box="[348,446,401,426]" italics="true" pageId="10" pageNumber="11">Stethusa</emphasis>
</taxonomicName>
,
<taxonomicName id="47EFDAE883346D4BFE3AFE3CFDB7FE05" box="[460,587,401,426]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983346D4BFE3AFE3CFDB7FE05" box="[460,587,401,426]" italics="true" pageId="10" pageNumber="11">S. klimschi</emphasis>
</taxonomicName>
and
<taxonomicName id="47EFDAE883346D4BFD75FE3CFCFBFE05" box="[643,775,401,426]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spuriella">
<emphasis id="B29B7D7983346D4BFD75FE3CFCFBFE05" box="[643,775,401,426]" italics="true" pageId="10" pageNumber="11">S. spuriella</emphasis>
</taxonomicName>
, lack the basal impressions of the last antennal segment. Therefore, this feature is not considered here a diagnostic character of the genus
<taxonomicName id="47EFDAE883346D4BFEFEFE4CFE95FE55" box="[264,361,481,506]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983346D4BFEFEFE4CFE95FE55" box="[264,361,481,506]" italics="true" pageId="10" pageNumber="11">Stethusa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8050A16B83346D4BFECEFDA4FB72FDE4" blockId="10.[264,1323,281,587]" pageId="10" pageNumber="11">
Based on the differences between
<taxonomicName id="47EFDAE883346D4BFD3EFDA4FCD6FD8D" box="[712,810,521,546]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983346D4BFD3EFDA4FCD6FD8D" box="[712,810,521,546]" italics="true" pageId="10" pageNumber="11">Stethusa</emphasis>
</taxonomicName>
and
<taxonomicName id="47EFDAE883346D4BFC92FDA4FC4CFD8D" box="[868,944,521,546]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983346D4BFC92FDA4FC4CFD8D" box="[868,944,521,546]" italics="true" pageId="10" pageNumber="11">Atheta</emphasis>
</taxonomicName>
listed in the Diagnosis, both are considered here as valid genera, pending a revision of the genera of
<taxonomicName id="47EFDAE883346D4BFBD0FD9CFB76FDE4" box="[1062,1162,561,587]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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