174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="C65B72E76BEAD1AC8A48EDECE9C95E00" ID-CLB-Dataset="28328" ID-DOI="10.11646/zootaxa.4568.1.8" ID-GBIF-Dataset="5513b97d-9796-43fd-94b3-557aada1afe4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2599310" ID-ZooBank="7F3892E0-31FF-43DC-BBB0-94DD82119B61" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553063194481" checkinUser="plazi" docAuthor="Kakizoe, Showtaro, Maruyama, Munetoshi & Masumoto, Kimio" docDate="2019" docId="0825441A9260FFEEEDCA3B3DFE4F988E" docLanguage="en" docName="zootaxa.4568.1.8.pdf" docOrigin="Zootaxa 4568 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megasybacodes brevitarsis Kakizoe & Maruyama & Masumoto 2019, new species" docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="F41C3C629262FFECED5D3A71FF8E9B64" masterDocTitle="Megasybacodes brevitarsis, a new genus and species of Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae) from Borneo" masterLastPageNumber="148" masterPageNumber="139" pageNumber="141" updateTime="1698702630802" updateUser="ExternalLinkService">
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<mods:mods id="A57BEB33A04B1B8FF63462FE9F3A3D79" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8BCB0F12A98F23F106E53F8E7CDD391C">
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<mods:title id="8EBD0945AC4C26363606A23407DB7F0D">Megasybacodes brevitarsis, a new genus and species of Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae) from Borneo</mods:title>
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</mods:titleInfo>
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<mods:name id="89D10C94E9DBA5043F453844C7BF09C5" type="personal">
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<mods:role id="3CE157300D2CE11A3907FA11B205F935">
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<mods:roleTerm id="B4B281D8938ADD5385DCC5665E60A01A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="74705D1BA104FE1595E19CA354E8C4F9">Kakizoe, Showtaro</mods:namePart>
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</mods:name>
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<mods:name id="4202C9640D7A268F4EAEDF47D8FA6E37" type="personal">
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<mods:roleTerm id="0E5742C59FF00F88B72411E0288C643F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6B60B855F8A7E4FB86816B285294C46A">Maruyama, Munetoshi</mods:namePart>
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</mods:name>
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<mods:name id="1F7768E38986AE0BFDA7CD28442C169F" type="personal">
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<mods:role id="DA20851B31CD259683D2937F97DE0DF2">
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<mods:roleTerm id="47F85EB1549A67C93DDAAE050C2B479F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="098B6A7746CDF9B28BD72D81786B7E27">Masumoto, Kimio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3150534ABA61FB3B069B30E7800D2D30">text</mods:typeOfResource>
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<mods:title id="30312C00E460D374FEB74C2001664BA7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="906CB662FE49E2930D02474EB53200DC">
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<mods:date id="D5BDDB43A0CFDD58C9881A23A8CC2D43">2019</mods:date>
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<mods:number id="20B64212A79E735E4664F3ECE98D35A5">2019-03-19</mods:number>
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<mods:number id="2EDF045C9C4C16DC99264D30E1001126">4568</mods:number>
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<mods:detail id="AC6189F1CCE6AEC933955B64F1798465" type="issue">
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<mods:number id="4E911B0E210FD6E19F1A9B04C088FBB1">1</mods:number>
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<mods:identifier id="EF983589C93EE846F922F0A27D182315" type="DOI">10.11646/zootaxa.4568.1.8</mods:identifier>
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<mods:identifier id="40F998622CF8A1C41FAE893B1E71B0BF" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0825441A9260FFEEEDCA3B3DFE4F988E" ID-DOI="http://doi.org/10.5281/zenodo.5935559" ID-GBIF-Taxon="154435733" ID-Zenodo-Dep="5935559" LSID="urn:lsid:plazi:treatment:0825441A9260FFEEEDCA3B3DFE4F988E" httpUri="http://treatment.plazi.org/id/0825441A9260FFEEEDCA3B3DFE4F988E" lastPageNumber="141" pageId="2" pageNumber="141">
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<subSubSection id="C896A6879260FFEEEDCA3B3DFBA59A02" box="[151,1067,332,358]" pageId="2" pageNumber="141" type="nomenclature">
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<paragraph id="8033F50C9260FFEEEDCA3B3DFBA59A02" blockId="2.[151,1067,332,391]" box="[151,1067,332,358]" pageId="2" pageNumber="141">
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<heading id="DB7B42609260FFEEEDCA3B3DFBA59A02" bold="true" box="[151,1067,332,358]" fontSize="11" level="1" pageId="2" pageNumber="141" reason="1">
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<emphasis id="B2F8291E9260FFEEEDCA3B3DFBA59A02" bold="true" box="[151,1067,332,358]" pageId="2" pageNumber="141">
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<taxonomicName id="478C8E8F9260FFEEEDCA3B3DFC009A02" ID-CoL="3YZNR" authority="Kakizoe, Maruyama & Masumoto" authorityName="Kakizoe & Maruyama & Masumoto" authorityYear="2019" box="[151,910,332,358]" class="Insecta" family="Scarabaeidae" genus="Megasybacodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="brevitarsis" status="sp. nov.">
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<emphasis id="B2F8291E9260FFEEEDCA3B3DFE569A02" bold="true" box="[151,472,332,358]" italics="true" pageId="2" pageNumber="141">Megasybacodes brevitarsis</emphasis>
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Kakizoe, Maruyama & Masumoto
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</taxonomicName>
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,
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<taxonomicNameLabel id="A9CB94659260FFEEEEC13B3DFBA59A02" box="[924,1067,332,358]" pageId="2" pageNumber="141" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C896A6879260FFEEEDCA3B1FFE9C9AE3" box="[151,274,366,391]" pageId="2" pageNumber="141" type="description">
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<paragraph id="8033F50C9260FFEEEDCA3B1FFE9C9AE3" blockId="2.[151,1067,332,391]" box="[151,274,366,391]" pageId="2" pageNumber="141">
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(
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<figureCitation id="18B7E9899260FFEEEDC23B1FFE869AE3" box="[159,264,366,391]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Figs. 1–5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C896A6879260FFEEEDCA3BC7FCB39A97" pageId="2" pageNumber="141" type="materials_examined">
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<paragraph id="8033F50C9260FFEEEDCA3BC7FCB39A97" blockId="2.[151,1437,438,1003]" pageId="2" pageNumber="141">
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<emphasis id="B2F8291E9260FFEEEDCA3BC7FEC69AAB" bold="true" box="[151,328,438,463]" pageId="2" pageNumber="141">Type material.</emphasis>
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<materialsCitation id="30E4FF519260FFEEEC0F3BC7FCB39A97" ID-GBIF-Occurrence="2013824516" collectingDate="2003-04" country="Malaysia" county="Kyushu University Museum" location="Japan" municipality="Fukuoka" pageId="2" pageNumber="141" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="5F374BAE9260FFEEEC0F3BC7FE339AAB" box="[338,445,438,463]" pageId="2" pageNumber="141" type="holotype">Holotype</typeStatus>
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, female, deposited in the
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<collectingCounty id="69528D809260FFEEEFB73BC7FBBF9AAA" box="[746,1073,438,463]" pageId="2" pageNumber="141">Kyushu University Museum</collectingCounty>
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||
,
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<collectingMunicipality id="60576F769260FFEEE9623BC7FB2A9AAB" box="[1087,1188,438,463]" pageId="2" pageNumber="141">Fukuoka</collectingMunicipality>
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,
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<location id="8553A3D79260FFEEE9EF3BC6FB789AAA" LSID="urn:lsid:plazi:treatment:0825441A9260FFEEEDCA3B3DFE4F988E:8553A3D79260FFEEE9EF3BC6FB789AAA" box="[1202,1270,439,462]" country="Malaysia" county="Kyushu University Museum" municipality="Fukuoka" name="Japan" pageId="2" pageNumber="141" stateProvince="Sabah">Japan</location>
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:
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<collectingCountry id="F89BB59C9260FFEEE8593BC6FA169AAB" box="[1284,1432,439,463]" name="Malaysia" pageId="2" pageNumber="141">MALAYSIA</collectingCountry>
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,
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<location id="8553A3D79260FFEEEDCA3BAAFF629A96" LSID="urn:lsid:plazi:treatment:0825441A9260FFEEEDCA3B3DFE4F988E:8553A3D79260FFEEEDCA3BAAFF629A96" box="[151,236,475,498]" country="Malaysia" county="Kyushu University Museum" municipality="Fukuoka" name="Borneo" pageId="2" pageNumber="141" stateProvince="Sabah">Borneo</location>
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,
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<collectingRegion id="42483BEE9260FFEEEDAA3BABFEB09A97" box="[247,318,474,499]" country="Malaysia" name="Sabah" pageId="2" pageNumber="141">Sabah</collectingRegion>
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,
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<location id="8553A3D79260FFEEEC173BAAFE799A97" LSID="urn:lsid:plazi:treatment:0825441A9260FFEEEDCA3B3DFE4F988E:8553A3D79260FFEEEC173BAAFE799A97" box="[330,503,474,499]" country="Malaysia" county="Kyushu University Museum" municipality="Fukuoka" name="Near Keningau" pageId="2" pageNumber="141" stateProvince="Sabah">Near Keningau</location>
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,
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<date id="F432D3CC9260FFEEEF5F3BAAFDEF9A97" box="[514,609,475,499]" pageId="2" pageNumber="141" value="2003-04">
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<collectingDate id="E4762A249260FFEEEF5F3BAAFDEF9A97" box="[514,609,475,499]" pageId="2" pageNumber="141" value="2003-04">IV 2003</collectingDate>
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</date>
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, local collector leg.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C896A6879260FFEEED9A3B8FFE2B983E" pageId="2" pageNumber="141" type="description">
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<paragraph id="8033F50C9260FFEEED9A3B8FFB9999AF" blockId="2.[151,1437,438,1003]" pageId="2" pageNumber="141">
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||
<emphasis id="B2F8291E9260FFEEED9A3B8FFDCE9973" bold="true" box="[199,576,510,535]" pageId="2" pageNumber="141">
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Description of
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<typeStatus id="5F374BAE9260FFEEEC273B8FFE6F9973" box="[378,481,510,535]" pageId="2" pageNumber="141" type="holotype">holotype</typeStatus>
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female.
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</emphasis>
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Head (
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<figureCitation id="18B7E9899260FFEEEFC93B8FFD719973" box="[660,767,510,535]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Figs. 1–3</figureCitation>
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) narrower than pronotum with a ratio of pronotal width to head width ca. 1.06; clypeus densely punctate-microsetose. Pronotum (
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<figureCitation id="18B7E9899260FFEEEEEC3853FB97995F" box="[945,1049,546,571]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Figs. 1, 3</figureCitation>
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) distinctly shorter than elytra with a ratio of elytral length to pronotal length ca. 1.98, with surface strongly wrinkled except carinae. Elytra (
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<figureCitation id="18B7E9899260FFEEE8773837FA1D993B" box="[1322,1427,582,607]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Figs. 1, 3</figureCitation>
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) with a ratio of length to width ca. 1.39; strongly wrinkled and almost glabrous except for microsetae at top of carinae. Protibiae (
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<figureCitation id="18B7E9899260FFEEEC2F38FFFE3999C3" box="[370,439,654,679]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Fig. 1</figureCitation>
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) densely setiferous near apex. Mesotibiae (
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<figureCitation id="18B7E9899260FFEEEEEA38FFFC7299C3" box="[951,1020,654,679]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Fig. 1</figureCitation>
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) densely setiferous on anterior and lateral margins. Metatibiae (
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<figureCitation id="18B7E9899260FFEEEC8E38C3FD9D99AF" box="[467,531,690,715]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[281,1326,226,1900]" captionTargetId="figure@5.[204,1378,195,1934]" captionTargetPageId="5" captionText="FIGURES 1–5. Megasybacodes brevitarsis Kakizoe, Maruyama & Masumoto, new genus and new species (holotype, female). 1, dorsal view; 2, ventral view; 3, dorso-lateral view; 4, right mesotarsus (dorsal view); 5, epipharynx. Scale bar = 5.0 mm for Figs. 1–3, 1.0 mm for Fig. 4 and 0.5 mm for Fig. 5." httpUri="https://zenodo.org/record/2599312/files/figure.png" pageId="2" pageNumber="141">Fig. 1</figureCitation>
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) with setae of similar densities to mesotibiae.
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</paragraph>
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<paragraph id="8033F50C9260FFEEED9A38A7FE09998B" blockId="2.[151,1437,438,1003]" box="[199,391,726,751]" pageId="2" pageNumber="141">
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<emphasis id="B2F8291E9260FFEEED9A38A7FE85998B" bold="true" box="[199,267,726,751]" pageId="2" pageNumber="141">Male.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="8033F50C9260FFEEED9A388BFE2B983E" blockId="2.[151,1437,438,1003]" pageId="2" pageNumber="141">
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||
<emphasis id="B2F8291E9260FFEEED9A388BFEF39877" bold="true" box="[199,381,762,787]" pageId="2" pageNumber="141">Measurements.</emphasis>
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Body length
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||
<quantity id="477458E99260FFEEEF4A388AFDF19876" box="[535,639,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="8.91" pageId="2" pageNumber="141" unit="mm" value="8.91">8.91 mm</quantity>
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||
; maximum width of head
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<quantity id="477458E99260FFEEEEF5388BFB9E9876" box="[936,1040,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="2.51" pageId="2" pageNumber="141" unit="mm" value="2.51">2.51 mm</quantity>
|
||
; median dorsal length of pronotum
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<quantity id="477458E99260FFEEEDCA396FFF739852" box="[151,253,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.66" pageId="2" pageNumber="141" unit="mm" value="2.66">2.66 mm</quantity>
|
||
, maximum width
|
||
<quantity id="477458E99260FFEEEC95396EFDA19852" box="[456,559,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="3.71" pageId="2" pageNumber="141" unit="mm" value="3.71">3.71 mm</quantity>
|
||
; sutural length of elytra
|
||
<quantity id="477458E99260FFEEEE1D396FFC289852" box="[832,934,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="5.27" pageId="2" pageNumber="141" unit="mm" value="5.27">5.27 mm</quantity>
|
||
, maximum width
|
||
<quantity id="477458E99260FFEEE92C396EFB569852" box="[1137,1240,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="3.79" pageId="2" pageNumber="141" unit="mm" value="3.79">3.79 mm</quantity>
|
||
; maximum width of epipharynx
|
||
<quantity id="477458E99260FFEEEC673933FE2F983E" box="[314,417,834,859]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="141" unit="mm" value="0.52">0.52 mm</quantity>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C896A6879260FFEEED9A3917FCE198C7" pageId="2" pageNumber="141" type="diagnosis">
|
||
<paragraph id="8033F50C9260FFEEED9A3917FCE198C7" blockId="2.[151,1437,438,1003]" pageId="2" pageNumber="141">
|
||
<emphasis id="B2F8291E9260FFEEED9A3917FECF981B" bold="true" box="[199,321,870,895]" pageId="2" pageNumber="141">Diagnosis.</emphasis>
|
||
This species is similar to the members of the genus
|
||
<taxonomicName id="478C8E8F9260FFEEEECA3917FB83981B" authorityName="Fairmaire" authorityYear="1896" box="[919,1037,870,895]" class="Insecta" family="Scarabaeidae" genus="Sybacodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2F8291E9260FFEEEECA3917FB83981B" box="[919,1037,870,895]" italics="true" pageId="2" pageNumber="141">Sybacodes</emphasis>
|
||
</taxonomicName>
|
||
in general appearance but is easily distinguished by the generic diagnostic features described above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C896A6879260FFEEED9A39DFFA1E98A3" box="[199,1424,942,967]" pageId="2" pageNumber="141" type="etymology">
|
||
<paragraph id="8033F50C9260FFEEED9A39DFFA1E98A3" blockId="2.[151,1437,438,1003]" box="[199,1424,942,967]" pageId="2" pageNumber="141">
|
||
<emphasis id="B2F8291E9260FFEEED9A39DFFEC098A3" bold="true" box="[199,334,942,967]" pageId="2" pageNumber="141">Etymology.</emphasis>
|
||
A combination of the Latin word
|
||
<emphasis id="B2F8291E9260FFEEEF9139DFFC8198A3" box="[716,783,942,967]" italics="true" pageId="2" pageNumber="141">brevis</emphasis>
|
||
meaning “short” and
|
||
<emphasis id="B2F8291E9260FFEEE95B39DEFBB798A2" box="[1030,1081,943,966]" italics="true" pageId="2" pageNumber="141">tarsi</emphasis>
|
||
, in reference to the short tarsi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C896A6879260FFEEED9A39A3FE4F988E" box="[199,449,978,1003]" pageId="2" pageNumber="141" type="distribution">
|
||
<paragraph id="8033F50C9260FFEEED9A39A3FE4F988E" blockId="2.[151,1437,438,1003]" box="[199,449,978,1003]" pageId="2" pageNumber="141">
|
||
<emphasis id="B2F8291E9260FFEEED9A39A3FEEE988F" bold="true" box="[199,352,978,1003]" pageId="2" pageNumber="141">Distribution.</emphasis>
|
||
Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |