191 lines
21 KiB
XML
191 lines
21 KiB
XML
<document id="1B1B3B324C7EAA87D48D864A24E69AD5" ID-DOI="10.11646/zootaxa.3683.2.3" ID-GBIF-Dataset="bb1cc4ed-5a7d-4c04-a042-2deb834b1d21" ID-ISSN="1175-5326" ID-Zenodo-Dep="219725" ID-ZooBank="1CDD7727-E1BF-4C46-B8FD-A13522FFC698" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460644285648" checkinUser="plazi" docAuthor="Dominiak, Patrycja & Alwin, Alicja" docDate="2013" docId="081B3D0DFFFFFFC442AB332503348E58" docLanguage="en" docName="zt03683p144.pdf" docOrigin="Zootaxa 3683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea antonii Dominiak, sp. nov." docType="treatment" docUuid="3BABE4D9-78F6-481A-B9BD-C3137E990676" docUuidSource="ZooBank" docVersion="10" lastPageNumber="134" masterDocId="F4224575FFFEFFC5423C302F02178965" masterDocTitle="Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)" masterLastPageNumber="144" masterPageNumber="133" pageNumber="134" updateTime="1698336543250" updateUser="plazi">
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<mods:title id="B2EA9EEB6CD8B460FA5025CAFB9540FA">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<mods:name id="868DFC2A38BF37AE00D1F397B64EF98B" type="personal">
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<mods:namePart id="339C523D3DA3022F581B310EFCBDF4B3">Dominiak, Patrycja</mods:namePart>
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<mods:namePart id="08B9B219CD6C1CA662261BE43163D4E6">Alwin, Alicja</mods:namePart>
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<mods:typeOfResource id="EE66130C5A3A38A5B17B6634BA1B6F07">text</mods:typeOfResource>
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<mods:title id="F0AC1A2BA2AFB61BAFD63DCE3B65F013">Zootaxa</mods:title>
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<mods:date id="37CC8ECFAA1C7C0991DBE442E654F965">2013</mods:date>
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<mods:number id="632055283AC4F0262DB2B57499040210">3683</mods:number>
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<mods:identifier id="8236403AB8A98CAE4F1AEE1C9968B1C3" type="DOI">10.11646/zootaxa.3683.2.3</mods:identifier>
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<treatment id="081B3D0DFFFFFFC442AB332503348E58" ID-DOI="http://doi.org/10.5281/zenodo.4499930" ID-GBIF-Taxon="119571703" ID-Zenodo-Dep="4499930" LSID="urn:lsid:zoobank.org:act:3BABE4D9-78F6-481A-B9BD-C3137E990676" httpUri="http://treatment.plazi.org/id/081B3D0DFFFFFFC442AB332503348E58" lastPageNumber="134" pageId="1" pageNumber="134">
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<subSubSection id="C8A8DF90FFFFFFC442AB3325031D8A20" pageId="1" pageNumber="134" type="nomenclature">
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<paragraph id="800D8C1BFFFFFFC442AB3325004D8A41" blockId="1.[151,602,778,837]" box="[151,602,778,804]" pageId="1" pageNumber="134">
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<heading id="DB453B77FFFFFFC442AB3325004D8A41" bold="true" box="[151,602,778,804]" fontSize="11" level="1" pageId="1" pageNumber="134" reason="1">
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<emphasis id="B2C65009FFFFFFC442AB3325004D8A41" bold="true" box="[151,602,778,804]" pageId="1" pageNumber="134">
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<taxonomicName id="47B2F798FFFFFFC442AB332503E28A41" ID-CoL="9HN78" authority="Dominiak" authorityName="Dominiak" box="[151,501,778,804]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii" status="sp. nov.">
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<emphasis id="B2C65009FFFFFFC442AB332503658A41" bold="true" box="[151,370,778,804]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
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Dominiak
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</taxonomicName>
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<taxonomicNameLabel id="A9F5ED72FFFFFFC443C03325004D8A41" box="[508,602,778,804]" pageId="1" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="800D8C1BFFFFFFC442AB3302031D8A20" blockId="1.[151,602,778,837]" box="[151,266,813,837]" pageId="1" pageNumber="134">
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(
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<figureCitation id="1889909EFFFFFFC442A3330203158A20" box="[159,258,813,837]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFFFFC442AB335A00B08D78" pageId="1" pageNumber="134" type="materials_examined">
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<paragraph id="800D8C1BFFFFFFC442AB335A00B08D78" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC442AB335A03518AEB" bold="true" box="[151,326,885,910]" pageId="1" pageNumber="134">
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<typeStatus id="5F0932B9FFFFFFC442AB335A02C58AE8" box="[151,210,885,909]" pageId="1" pageNumber="134">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="5F0932B9FFFFFFC44372335A03AC8AE8" box="[334,443,885,909]" pageId="1" pageNumber="134" type="holotype">Holotype</typeStatus>
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: adult male.
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<collectingCountry id="F8A5CC8BFFFFFFC44070335A014E8AEB" box="[588,857,885,910]" name="United Arab Emirates" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC44070335A014E8AEB" bold="true" box="[588,857,885,910]" pageId="1" pageNumber="134">United Arab Emirates</emphasis>
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</collectingCountry>
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. Sharjah Desert Park, N 25˚17' E 55˚42',
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<quantity id="474A21FEFFFFFFC4470F335A07798AE8" box="[1331,1390,885,909]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="134" unit="m" value="70.0">70 m</quantity>
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asl,
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<date id="F40CAADBFFFFFFC442AB33B603378AD4" box="[151,288,921,945]" pageId="1" pageNumber="134" value="2008-12-15">15.XII.2008</date>
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–
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<date id="F40CAADBFFFFFFC4430D33B6038C8AD4" box="[305,411,921,945]" pageId="1" pageNumber="134" value="2009-01-12">12.I.2009</date>
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, light trap, leg. A. van Harten (UG).
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<typeStatus id="5F0932B9FFFFFFC4415E33B501C48AD7" box="[866,979,922,946]" pageId="1" pageNumber="134" type="paratype">Paratypes</typeStatus>
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:
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<emphasis id="B2C65009FFFFFFC441DF33B606E88AD7" bold="true" box="[995,1279,921,946]" pageId="1" pageNumber="134">
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<collectingCountry id="F8A5CC8BFFFFFFC441DF33B606ED8AD7" box="[995,1274,921,946]" name="United Arab Emirates" pageId="1" pageNumber="134">United Arab Emirates</collectingCountry>
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.
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</emphasis>
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Al-Ajban, N 24˚36' E 55˚01',
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<quantity id="474A21FEFFFFFFC4436D3392039D8AB0" box="[337,394,957,981]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="134" unit="m" value="60.0">60 m</quantity>
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asl,
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<date id="F40CAADBFFFFFFC44383339200678AB0" box="[447,624,957,981]" pageId="1" pageNumber="134" value="2006-01-30" valueMax="2006-02-26" valueMin="2006-01-30">
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30.I–26.
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<bibRefCitation id="E423F1EAFFFFFFC44027339200678AB0" author="II" box="[539,624,957,981]" pageId="1" pageNumber="134" refString="Dominiak, P. & Szadziewski, R. (2006) Biting midges of the genus Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae) collected from Polish peat-bogs. Dipteron, 22, 4 - 7." type="journal article" year="2006">II.2006</bibRefCitation>
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</date>
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, Malaise trap,
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<specimenCount id="96B44792FFFFFFC44129339201798AB0" box="[789,878,957,981]" pageId="1" pageNumber="134" type="male">2 males</specimenCount>
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, leg. A. van Harten (UG). Sharjah Desert Park, N 25˚17' E 55˚42',
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<quantity id="474A21FEFFFFFFC4436533CE03828A9C" box="[345,405,993,1017]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="134" unit="m" value="70.0">70 m</quantity>
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asl,
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<date id="F40CAADBFFFFFFC443F233CE00418A9C" box="[462,598,993,1017]" pageId="1" pageNumber="134" value="2008-12-11">11.XII.2008</date>
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||
–
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<date id="F40CAADBFFFFFFC4405A33CE00C58A9C" box="[614,722,993,1017]" pageId="1" pageNumber="134" value="2009-01-06">06.I.2009</date>
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, light trap,
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<specimenCount id="96B44792FFFFFFC4416633CE01BD8A9C" box="[858,938,993,1017]" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
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||
, leg. A. van Harten (UG);
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<date id="F40CAADBFFFFFFC446DA33CE07808A9C" box="[1254,1431,993,1017]" pageId="1" pageNumber="134" value="2009-01-12" valueMax="2009-02-02" valueMin="2009-01-12">12.I–02.II.2009</date>
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, light trap,
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<specimenCount id="96B44792FFFFFFC44336342A034F8D78" box="[266,344,1029,1053]" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
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, leg. A. van Harten (
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<collectionCode id="E6A314DEFFFFFFC4407D342A008B8D78" box="[577,668,1029,1053]" pageId="1" pageNumber="134">BMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFFFFC442FB340607078D00" pageId="1" pageNumber="134" type="diagnosis">
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<paragraph id="800D8C1BFFFFFFC442FB340607078D00" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC442FB340603568D27" bold="true" box="[199,321,1065,1090]" pageId="1" pageNumber="134">Diagnosis.</emphasis>
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<taxonomicName id="47B2F798FFFFFFC443703406000B8D24" box="[332,540,1065,1089]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
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<emphasis id="B2C65009FFFFFFC443703406000B8D24" box="[332,540,1065,1089]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
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</taxonomicName>
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is distinct in having a fifth palpal segment nearly two times longer than the fourth one, as well as a concave and well sclerotized posterior margin of 9th sternite of male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFFFFC442FB345E06DC8F00" pageId="1" pageNumber="134" type="description">
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<paragraph id="800D8C1BFFFFFFC442FB345E00E68C9C" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC442FB345E03B48DEF" bold="true" box="[199,419,1137,1162]" pageId="1" pageNumber="134">Description. Male.</emphasis>
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Head. Distal flagellomeres with two rows of setae (
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<figureCitation id="1889909EFFFFFFC441DB345E063F8DEC" box="[999,1064,1137,1161]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 1</figureCitation>
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). Total length of flagellum
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<quantity id="474A21FEFFFFFFC44761345E02E88DC8" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="5.8" metricValueMin="5.6" pageId="1" pageNumber="134" unit="mm" value="0.57" valueMax="0.58" valueMin="0.56">0.56– 0.58 mm</quantity>
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, AR 0.89–0.94 (n=2). Clypeus entire with lateral lobes, bearing from 8 to 10 strong setae. Third palpal segment slender, length 60–72 μm, PR III 4.28–5.64 (n=4). Fifth palpal segment elongate, 1.7–2 times longer than fourth segment, with long setae located also in middle and at base (
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<figureCitation id="1889909EFFFFFFC4419334CF01E08D9D" box="[943,1015,1248,1272]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
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). Thorax. Transverse suture present, number of supraalar setae varing from 6 to 8. Scutellum yellow, with 7–8 strong and few smaller setae. Wing length
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<quantity id="474A21FEFFFFFFC442DB350703878C25" box="[231,400,1320,1344]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.9" metricValueMin="7.8" pageId="1" pageNumber="134" unit="mm" value="0.885" valueMax="0.99" valueMin="0.78">0.78–0.99 mm</quantity>
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, CR 0.44–0.45 (n=4), two radial cells present. Legs unicolor,
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<collectingCountry id="F8A5CC8BFFFFFFC4465D350606938C25" box="[1121,1156,1321,1344]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
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I 1.9–2.1 (n=4),
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<collectingCountry id="F8A5CC8BFFFFFFC4477D350607728C25" box="[1345,1381,1321,1344]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
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II 2.0 (n=2),
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<collectingCountry id="F8A5CC8BFFFFFFC442DF357F03108C02" box="[227,263,1360,1383]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
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III 1.8–2.0 (n=4). Genitalia (
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<figureCitation id="1889909EFFFFFFC44000356000978C02" box="[572,640,1359,1383]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 3</figureCitation>
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). Tergite 9 rounded distally, with a pair of short apicolateral processes. Posterior margin of sternite 9 concave and well sclerotized, with very small median projections. Gonostylus long, slightly arched. Parameres asymmetrical. Posterior process broadly fused to right basal arm, long and slender, gradually tapering distally, markedly curved on apical portion. Arch of aedeagus low. Submedian projections stout, triangular, highly sclerotized only on ventral margins.
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</paragraph>
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<paragraph id="800D8C1BFFFFFFC442FB362900498F78" blockId="1.[151,1437,885,1853]" box="[199,606,1541,1566]" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC442FB362903F68F7B" bold="true" box="[199,481,1541,1566]" pageId="1" pageNumber="134">Female and immatures.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="800D8C1BFFFFFFC442FB360606DC8F00" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
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<emphasis id="B2C65009FFFFFFC442FB360603FF8F27" bold="true" box="[199,488,1577,1602]" pageId="1" pageNumber="134">Derivation of the name.</emphasis>
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This species is named after Antonius van Harten, the leader of the entomological projects in the Middle East, who collected and kindly sent us a lot of materials of biting midges.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFFFFC442FB365E00808FEC" box="[199,663,1649,1674]" pageId="1" pageNumber="134" type="distribution">
|
||
<paragraph id="800D8C1BFFFFFFC442FB365E00808FEC" blockId="1.[151,1437,885,1853]" box="[199,663,1649,1674]" pageId="1" pageNumber="134">
|
||
<emphasis id="B2C65009FFFFFFC442FB365E03778FEF" bold="true" box="[199,352,1649,1674]" pageId="1" pageNumber="134">Distribution.</emphasis>
|
||
The
|
||
<collectingCountry id="F8A5CC8BFFFFFFC443A6365E00848FEC" box="[410,659,1649,1673]" name="United Arab Emirates" pageId="1" pageNumber="134">United Arab Emirates</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8A8DF90FFFFFFC442FB36BA03348E58" pageId="1" pageNumber="134" type="discussion">
|
||
<paragraph id="800D8C1BFFFFFFC442FB36BA03348E58" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
|
||
<emphasis id="B2C65009FFFFFFC442FB36BA03588FCB" bold="true" box="[199,335,1685,1710]" pageId="1" pageNumber="134">Comments.</emphasis>
|
||
Among all other species of this subgenus,
|
||
<taxonomicName id="47B2F798FFFFFFC4416236B906278FCB" box="[862,1072,1686,1710]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
|
||
<emphasis id="B2C65009FFFFFFC4416236B906278FCB" box="[862,1072,1686,1710]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
|
||
</taxonomicName>
|
||
is characterized in having the posterior margin of sternite 9 concave and strongly chitinized, and the aedeagus with prominent submedian projections. The species resembles
|
||
<taxonomicName id="47B2F798FFFFFFC4401236F001248F90" authority="Winnertz" authorityName="Winnertz" box="[558,819,1757,1782]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="modesta">
|
||
<emphasis id="B2C65009FFFFFFC4401236F000558F93" box="[558,578,1759,1782]" italics="true" pageId="1" pageNumber="134">D</emphasis>
|
||
.
|
||
<emphasis id="B2C65009FFFFFFC4406F36F100A78F93" box="[595,688,1758,1782]" italics="true" pageId="1" pageNumber="134">modesta</emphasis>
|
||
(Winnertz)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47B2F798FFFFFFC4414C36F006558F90" authority="Clastrier" authorityName="Clastrier" box="[880,1090,1757,1782]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="wirthi">
|
||
<emphasis id="B2C65009FFFFFFC4414C36F001938F93" box="[880,900,1759,1782]" italics="true" pageId="1" pageNumber="134">D</emphasis>
|
||
.
|
||
<emphasis id="B2C65009FFFFFFC441A836F101C08F93" box="[916,983,1758,1782]" italics="true" pageId="1" pageNumber="134">wirthi</emphasis>
|
||
Clastrier
|
||
</taxonomicName>
|
||
(known only from female) in having a similar shape of the fifth palpal segment that is elongate, but in
|
||
<taxonomicName id="47B2F798FFFFFFC441EA372C065E8E7F" box="[982,1097,1794,1818]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
|
||
<emphasis id="B2C65009FFFFFFC441EA372C01FD8E7F" box="[982,1002,1795,1818]" italics="true" pageId="1" pageNumber="134">D</emphasis>
|
||
.
|
||
<emphasis id="B2C65009FFFFFFC441C5372C065E8E7F" box="[1017,1097,1795,1818]" italics="true" pageId="1" pageNumber="134">antonii</emphasis>
|
||
</taxonomicName>
|
||
bears additional long setae in median part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |